Gorilla Characters In Comics
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Gorillas are herbivorous, predominantly ground-dwelling great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the
eastern gorilla The eastern gorilla (''Gorilla beringei'') is a critically endangered species of the genus ''Gorilla'' and the largest living primate. At present, the species is subdivided into two subspecies. There are 3,800 eastern lowland gorillas or Graue ...
and the
western gorilla The western gorilla (''Gorilla gorilla'') is a great ape found in Africa, one of two species of the hominid genus ''Gorilla''. Large and robust with males weighing around , the hair is significantly lighter in color than the eastern gorilla, ''G ...
, and either four or five
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
. The DNA of gorillas is highly similar to that of humans, from 95 to 99% depending on what is included, and they are the next closest living relatives to humans after chimpanzees and
bonobos The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee and less often the dwarf chimpanzee or gracile chimpanzee, is an endangered great ape and one of the two species making up the genus '' Pan,'' the other being the comm ...
. Gorillas are the largest living primates, reaching heights between 1.25 and 1.8 metres, weights between 100 and 270 kg, and arm spans up to 2.6 metres, depending on species and sex. They tend to live in troops, with the leader being called a silverback. The Eastern gorilla is distinguished from the Western by darker fur colour and some other minor morphological differences. Gorillas tend to live 35–40 years in the wild. The oldest gorilla known is Fatou (b. 1957), who is still alive at the advanced age of 65 years. Gorillas' natural habitats cover tropical or subtropical forest in
Sub-Saharan Africa Sub-Saharan Africa is, geographically, the area and regions of the continent of Africa that lies south of the Sahara. These include West Africa, East Africa, Central Africa, and Southern Africa. Geopolitically, in addition to the List of sov ...
. Although their range covers a small percentage of Sub-Saharan Africa, gorillas cover a wide range of elevations. The mountain gorilla inhabits the Albertine Rift montane
cloud forest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest (TMCF), is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud c ...
s of the Virunga Volcanoes, ranging in altitude from . Lowland gorillas live in dense forests and lowland swamps and marshes as low as sea level, with western lowland gorillas living in Central West African countries and
eastern lowland gorilla The eastern lowland gorilla (''Gorilla beringei graueri'') or Grauer's gorilla is a Critically Endangered subspecies of eastern gorilla endemic to the mountainous forests of eastern Democratic Republic of the Congo. Important populations of t ...
s living in the Democratic Republic of the Congo near its border with
Rwanda Rwanda (; rw, u Rwanda ), officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of Central Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator ...
. There are thought to be around 316,000 western gorillas in the wild, and 5,000 eastern gorillas. Both species are classified as Critically Endangered by the IUCN; all subspecies are classified as Critically Endangered with the exception of the mountain gorilla, which is classified as Endangered. There are many threats to their survival, such as poaching, habitat destruction, and disease, which threaten the survival of the species. However, conservation efforts have been successful in some areas where they live.


Etymology

The word "gorilla" comes from the history of
Hanno the Navigator Hanno the Navigator (sometimes "Hannon"; xpu, 𐤇𐤍𐤀 , ; ) was a Carthaginian explorer of the fifth century BC, best known for his naval exploration of the western coast of Africa. The only source of his voyage is a ''periplus'' transla ...
( 500 BC), a
Carthaginian The term Carthaginian ( la, Carthaginiensis ) usually refers to a citizen of Ancient Carthage. It can also refer to: * Carthaginian (ship), a three-masted schooner built in 1921 * Insurgent privateers; nineteenth-century South American privateers, ...
explorer on an expedition to the west African coast to the area that later became Sierra Leone. Members of the expedition encountered "savage people, the greater part of whom were women, whose bodies were hairy, and whom our interpreters called Gorillae". It is unknown whether what the explorers encountered were what we now call gorillas, another species of ape or monkeys, or humans. Skins of gorillai women, brought back by Hanno, are reputed to have been kept at Carthage until Rome destroyed the city 350 years later at the end of the
Punic Wars The Punic Wars were a series of wars between 264 and 146BC fought between Roman Republic, Rome and Ancient Carthage, Carthage. Three conflicts between these states took place on both land and sea across the western Mediterranean region and i ...
, 146 BC. The American physician and missionary Thomas Staughton Savage and naturalist
Jeffries Wyman Jeffries Wyman (August 11, 1814 – September 4, 1874) was an American naturalist and anatomist, born in Chelmsford, Massachusetts. Wyman died in Bethlehem, New Hampshire of a pulmonary hemorrhage. Career He graduated Harvard College in 183 ...
first described the
western gorilla The western gorilla (''Gorilla gorilla'') is a great ape found in Africa, one of two species of the hominid genus ''Gorilla''. Large and robust with males weighing around , the hair is significantly lighter in color than the eastern gorilla, ''G ...
in 1847 from specimens obtained in
Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...
. They called it ''Troglodytes gorilla'', using the then-current name of the chimpanzee genus. The species name was derived , as described by Hanno.


Evolution and classification

The closest relatives of gorillas are the other two Homininae genera, chimpanzees and humans, all of them having diverged from a common ancestor about 7 million years ago. Human gene sequences differ only 1.6% on average from the sequences of corresponding gorilla genes, but there is further difference in how many
copies Copy may refer to: * Copying or the product of copying (including the plural "copies"); the duplication of information or an artifact **Cut, copy and paste, a method of reproducing text or other data in computing **File copying **Photocopying, a p ...
each gene has. Until recently, gorillas were considered to be a single species, with three subspecies: the western lowland gorilla, the eastern lowland gorilla and the mountain gorilla. There is now agreement that there are two species, each with two subspecies. More recently, a third subspecies has been claimed to exist in one of the species. The separate species and subspecies developed from a single type of gorilla during the Ice Age, when their forest habitats shrank and became isolated from each other. Primatologists continue to explore the relationships between various gorilla populations. The species and subspecies listed here are the ones upon which most scientists agree. The proposed third subspecies of ''Gorilla beringei'', which has not yet received a trinomen, is the Bwindi population of the mountain gorilla, sometimes called the
Bwindi gorilla The Bwindi Impenetrable National Park (BINP) is in southwestern Uganda. The park is part of the Bwindi Impenetrable Forest and is situated along the Democratic Republic of the Congo (DRC) border next to the Virunga National Park and on the edge ...
. Some variations that distinguish the classifications of gorilla include varying density, size, hair colour, length, culture, and facial widths. Population genetics of the lowland gorillas suggest that the western and eastern lowland populations diverged around 261 thousand years ago.


Characteristics

Wild male gorillas weigh , while adult females weigh . Adult males are tall, with an arm span that stretches from . Female gorillas are shorter at , with smaller arm spans. Colin Groves (1970) calculated the average weight of 42 wild adult male gorillas at 144 kg, while Smith and Jungers (1997) found the average weight of 19 wild adult male gorillas to be 169 kg. Adult male gorillas are known as silverbacks due to the characteristic silver hair on their backs reaching to the hips. The tallest gorilla recorded was a silverback with an arm span of , a chest of , and a weight of , shot in Alimbongo, northern Kivu in May 1938. The heaviest gorilla recorded was a silverback shot in Ambam, Cameroon, which weighed . Males in captivity can be
overweight Being overweight or fat is having more body fat than is optimally healthy. Being overweight is especially common where food supplies are plentiful and lifestyles are sedentary. , excess weight reached epidemic proportions globally, with mo ...
and reach weights up to . The eastern gorilla is more darkly coloured than the western gorilla, with the mountain gorilla being the darkest of all. The mountain gorilla also has the thickest hair. The western lowland gorilla can be brown or greyish with a reddish forehead. In addition, gorillas that live in lowland forest are more slender and agile than the more bulky mountain gorillas. The eastern gorilla also has a longer face and broader chest than the western gorilla. Like humans, gorillas have individual fingerprints. Their eye colour is dark brown, framed by a black ring around the iris. Gorilla facial structure is described as
mandibular prognathism Prognathism, also called Habsburg jaw or Habsburgs' jaw primarily in the context of its prevalence amongst members of the House of Habsburg, is a positional relationship of the mandible or maxilla to the skeletal base where either of the jaws pr ...
, that is, the mandible protrudes farther out than the maxilla. Adult males also have a prominent
sagittal crest A sagittal crest is a ridge of bone running lengthwise along the midline of the top of the skull (at the sagittal suture) of many mammalian and reptilian skulls, among others. The presence of this ridge of bone indicates that there are exceptiona ...
. Gorillas move around by knuckle-walking, although they sometimes walk upright for short distances, typically while carrying food or in defensive situations. A 2018 study investigating the hand posture of 77 mountain gorillas at
Bwindi Impenetrable National Park The Bwindi Impenetrable National Park (BINP) is in southwestern Uganda. The park is part of the Bwindi Impenetrable Forest and is situated along the Democratic Republic of the Congo (DRC) border next to the Virunga National Park and on the edge o ...
(8% of the population) found that knuckle walking was done only 60% of the time, and they also supported their weight on their fists, the backs of their hands/feet, and on their palms/soles (with the digits flexed). Such a range of hand postures was previously thought to have been used by only orangutans. Studies of gorilla
handedness In human biology, handedness is an individual's preferential use of one hand, known as the dominant hand, due to it being stronger, faster or more Fine motor skill, dextrous. The other hand, comparatively often the weaker, less dextrous or sim ...
have yielded varying results, with some arguing for no preference for either hand, and others right-hand dominance for the general population. Studies have shown gorilla blood is not reactive to anti-A and anti-B
monoclonal antibodies A monoclonal antibody (mAb, more rarely called moAb) is an antibody produced from a cell Lineage made by cloning a unique white blood cell. All subsequent antibodies derived this way trace back to a unique parent cell. Monoclonal antibodies ca ...
, which would, in humans, indicate
type O blood The ABO blood group system is used to denote the presence of one, both, or neither of the A and B antigens on erythrocytes. For human blood transfusions, it is the most important of the 43 different blood type (or group) classification system ...
. Due to novel sequences, though, it is different enough to not conform with the human ABO blood group system, into which the other great apes fit. A gorilla's lifespan is normally between 35 and 40 years, although zoo gorillas may live for 50 years or more. Colo, a female western gorilla at the Columbus Zoo and Aquarium, was the oldest known gorilla at 60 years of age when she died on 17 January 2017.
Ozzie Ozzie or Ozzy is a masculine given name, sometimes a short form (hypocorism) of Oswald, Oscar, Osborne, Osman and Ozymandias and other names, surname and nickname which may refer to: Animals * Ozzie (gorilla) (born c. 1961) People * Chris Osgood ...
was 61 years old at the time of his death in January 2022.


Distribution and habitat

Gorillas have a patchy distribution. The range of the two species is separated by the Congo River and its tributaries. The western gorilla lives in west central Africa, while the eastern gorilla lives in east central Africa. Between the species, and even within the species, gorillas live in a variety of habitats and elevations. Gorilla habitat ranges from montane forest to swampland. Eastern gorillas inhabit montane and submontane forests between above sea level. Mountain gorillas live in montane forests at the higher end of the elevation range, while eastern lowland gorillas live in submontane forests at the lower end. In addition, eastern lowland gorillas live in montane bamboo forests, as well as lowland forests ranging from in elevation. Western gorillas live in both lowland swamp forests and montane forests, at elevations ranging from sea level to . Western lowland gorillas live in swamp and lowland forests ranging up to , and Cross River gorillas live in low-lying and submontane forests ranging from .


Ecology


Diet and foraging

A gorilla's day is divided between rest periods and travel or feeding periods. Diets differ between and within species. Mountain gorillas mostly eat foliage, such as leaves, stems, pith, and shoots, while fruit makes up a very small part of their diets. Mountain gorilla food is widely distributed and neither individuals nor groups have to compete with one another. Their home ranges vary from , and their movements range around or less on an average day. Despite eating a few species in each habitat, mountain gorillas have flexible diets and can live in a variety of habitats. Eastern lowland gorillas have more diverse diets, which vary seasonally. Leaves and pith are commonly eaten, but fruits can make up as much as 25% of their diets. Since fruit is less available, lowland gorillas must travel farther each day, and their home ranges vary from , with day ranges . Eastern lowland gorillas will also eat insects, preferably ants. Western lowland gorillas depend on fruits more than the others and they are more dispersed across their range. They travel even farther than the other gorilla subspecies, at per day on average, and have larger home ranges of . Western lowland gorillas have less access to terrestrial herbs, although they can access aquatic herbs in some areas. Termites and ants are also eaten. Gorillas rarely drink water "because they consume succulent vegetation that is almost half water as well as morning dew", although both mountain and lowland gorillas have been observed drinking.


Nesting

Gorillas construct nests for daytime and night use. Nests tend to be simple aggregations of branches and leaves about in diameter and are constructed by individuals. Gorillas, unlike chimpanzees or orangutans, tend to sleep in nests on the ground. The young nest with their mothers, but construct nests after three years of age, initially close to those of their mothers. Gorilla nests are distributed arbitrarily and use of tree species for site and construction appears to be opportunistic. Nest-building by great apes is now considered to be not just
animal architecture Structures built by animals, often called animal architecture, abound in nature. Examples of animal structures include termite mounds, wasp and beehives, burrow complexes of rodents, beaver dams, elaborate nests of birds, and webs of spiders. ...
, but as an important instance of
tool use Tool use by animals is a phenomenon in which an animal uses any kind of tool in order to achieve a goal such as acquiring food and water, grooming, defence, communication, recreation or construction. Originally thought to be a skill possessed o ...
. Gorillas make a new nest to sleep on daily and do not use the previous one. This even if remaining in the same place. Usually, they are made an hour before dusk, to be ready to sleep when night falls. Gorillas sleep longer than humans, an average of 12 hours per day.


Threats and competition

One possible predator of gorillas is the
leopard The leopard (''Panthera pardus'') is one of the five extant species in the genus '' Panthera'', a member of the cat family, Felidae. It occurs in a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, a ...
. Gorilla remains have been found in leopard scat, but this may be the result of scavenging. When the group is attacked by humans, leopards, or other gorillas, an individual silverback will protect the group, even at the cost of his own life. Gorillas do not appear to directly compete with
chimpanzee The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative th ...
s in areas where they overlap. When fruit is abundant gorilla and chimpanzee diets converge, but when fruit is scarce gorillas resort to vegetation. The two apes may also feed on different species, whether fruit or insects. Gorillas and chimpanzees usually ignore or avoid each other when feeding on the same tree, but coalitions of chimpanzees have been observed attacking families of gorillas including silverbacks and killing infants.


Behaviour


Social structure

Gorillas live in groups called troops. Troops tend to be made of one adult male or silverback, with a harem of multiple adult females and their offspring. However, multiple-male troops also exist. A silverback is typically more than 12 years of age, and is named for the distinctive patch of silver hair on his back, which comes with maturity. Silverbacks have large canine teeth that also come with maturity. Both males and females tend to emigrate from their natal groups. For mountain gorillas, females disperse from their natal troops more than males. Mountain gorillas and western lowland gorillas also commonly transfer to second new groups. Mature males also tend to leave their groups and establish their own troops by attracting emigrating females. However, male mountain gorillas sometimes stay in their natal troops and become subordinate to the silverback. If the silverback dies, these males may be able to become dominant or mate with the females. This behaviour has not been observed in eastern lowland gorillas. In a single male group, when the silverback dies, the females and their offspring disperse and find a new troop. Without a silverback to protect them, the infants will likely fall victim to
infanticide Infanticide (or infant homicide) is the intentional killing of infants or offspring. Infanticide was a widespread practice throughout human history that was mainly used to dispose of unwanted children, its main purpose is the prevention of reso ...
. Joining a new group is likely to be a tactic against this. However, while gorilla troops usually disband after the silverback dies, female eastern lowlands gorillas and their offspring have been recorded staying together until a new silverback transfers into the group. This likely serves as protection from leopards. The silverback is the centre of the troop's attention, making all the decisions, mediating conflicts, determining the movements of the group, leading the others to feeding sites, and taking responsibility for the safety and well-being of the troop. Younger males subordinate to the silverback, known as blackbacks, may serve as backup protection. Blackbacks are aged between 8 and 12 years and lack the silver back hair. The bond that a silverback has with his females forms the core of gorilla social life. Bonds between them are maintained by grooming and staying close together. Females form strong relationships with males to gain mating opportunities and protection from predators and infanticidal outside males. However, aggressive behaviours between males and females do occur, but rarely lead to serious injury. Relationships between females may vary. Maternally related females in a troop tend to be friendly towards each other and associate closely. Otherwise, females have few friendly encounters and commonly act aggressively towards each other. Females may fight for social access to males and a male may intervene. Male gorillas have weak social bonds, particularly in multiple-male groups with apparent dominance hierarchies and strong competition for mates. Males in all-male groups, though, tend to have friendly interactions and socialise through play, grooming, and staying together, and occasionally they even engage in homosexual interactions. Severe aggression is rare in stable groups, but when two mountain gorilla groups meet the two silverbacks can sometimes engage in a fight to the death, using their canines to cause deep, gaping injuries.


Reproduction and parenting

Females mature at 10–12 years (earlier in captivity), and males at 11–13 years. A female's first ovulatory cycle occurs when she is six years of age, and is followed by a two-year period of adolescent infertility. The estrous cycle lasts 30–33 days, with outward ovulation signs subtle compared to those of chimpanzees. The gestation period lasts 8.5 months. Female mountain gorillas first give birth at 10 years of age and have four-year interbirth intervals. Males can be fertile before reaching adulthood. Gorillas mate year round. Females will purse their lips and slowly approach a male while making eye contact. This serves to urge the male to mount her. If the male does not respond, then she will try to attract his attention by reaching towards him or slapping the ground. In multiple-male groups, solicitation indicates female preference, but females can be forced to mate with multiple males. Males incite copulation by approaching a female and displaying at her or touching her and giving a "train grunt". Recently, gorillas have been observed engaging in face-to-face sex, a trait once considered unique to humans and
bonobo The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee and less often the dwarf chimpanzee or gracile chimpanzee, is an endangered great ape and one of the two species making up the genus '' Pan,'' the other being the comm ...
s. Gorilla infants are vulnerable and dependent, thus mothers, their primary caregivers, are important to their survival. Male gorillas are not active in caring for the young, but they do play a role in socialising them to other youngsters. The silverback has a largely supportive relationship with the infants in his troop and shields them from aggression within the group. Infants remain in contact with their mothers for the first five months and mothers stay near the silverback for protection. Infants suckle at least once per hour and sleep with their mothers in the same nest. Infants begin to break contact with their mothers after five months, but only for a brief period each time. By 12 months old, infants move up to from their mothers. At around 18–21 months, the distance between mother and offspring increases and they regularly spend time away from each other. In addition, nursing decreases to once every two hours. Infants spend only half of their time with their mothers by 30 months. They enter their juvenile period at their third year, and this lasts until their sixth year. At this time, gorillas are weaned and they sleep in a separate nest from their mothers. After their offspring are weaned, females begin to ovulate and soon become pregnant again. The presence of play partners, including the silverback, minimizes conflicts in weaning between mother and offspring.


Communication

Twenty-five distinct vocalisations are recognised, many of which are used primarily for group communication within dense vegetation. Sounds classified as grunts and barks are heard most frequently while traveling, and indicate the whereabouts of individual group members. They may also be used during social interactions when discipline is required. Screams and roars signal alarm or warning, and are produced most often by silverbacks. Deep, rumbling belches suggest contentment and are heard frequently during feeding and resting periods. They are the most common form of intragroup communication. For this reason, conflicts are most often resolved by displays and other threat behaviours that are intended to intimidate without becoming physical. As a result, fights do not occur very frequently. The ritualized charge display is unique to gorillas. The entire sequence has nine steps: (1) progressively quickening hooting, (2) symbolic feeding, (3) rising bipedally, (4) throwing vegetation, (5) chest-beating with cupped hands, (6) one leg kick, (7) sideways running, two-legged to four-legged, (8) slapping and tearing vegetation, and (9) thumping the ground with palms to end display. A gorilla's chest-beat may vary in frequency depending on its size. Smaller ones tend to have higher frequencies, while larger ones tend to be lower. They also do it the most when females are ready to mate.


Intelligence

Gorillas are considered highly intelligent. A few individuals in captivity, such as
Koko Koko or KOKO may refer to: Animals *Koko (gorilla) (1971–2018), a gorilla trained to communicate in American Sign Language *Koko (dog) (2005–2012), the Australian kelpie in the 2011 film ''Red Dog'' *Koko (horse), an Irish racehorse that won ...
, have been taught a subset of sign language. Like the other great apes, gorillas can laugh, grieve, have "rich emotional lives", develop strong family bonds, make and use tools, and think about the past and future. Some researchers believe gorillas have spiritual feelings or religious sentiments. They have been shown to have cultures in different areas revolving around different methods of food preparation, and will show individual
colour preferences In the psychology of color, color preferences are the tendency for an individual or a group to prefer some colors over others, such as having a favorite color or a traditional color. Introduction An 'ecological valence theory' (EVT) has bee ...
.


Tool use

The following observations were made by a team led by Thomas Breuer of the Wildlife Conservation Society in September 2005. Gorillas are now known to use tools in the wild. A female gorilla in the Nouabalé-Ndoki National Park in the
Republic of Congo The Republic of the Congo (french: République du Congo, ln, Republíki ya Kongó), also known as Congo-Brazzaville, the Congo Republic or simply either Congo or the Congo, is a country located in the western coast of Central Africa to the w ...
was recorded using a stick as if to gauge the depth of water whilst crossing a swamp. A second female was seen using a tree stump as a bridge and also as a support whilst fishing in the swamp. This means all of the great apes are now known to use tools. In September 2005, a two-and-a-half-year-old gorilla in the
Republic of Congo The Republic of the Congo (french: République du Congo, ln, Republíki ya Kongó), also known as Congo-Brazzaville, the Congo Republic or simply either Congo or the Congo, is a country located in the western coast of Central Africa to the w ...
was discovered using rocks to smash open palm nuts inside a game sanctuary. While this was the first such observation for a gorilla, over 40 years previously, chimpanzees had been seen using tools in the wild 'fishing' for termites. Nonhuman great apes are endowed with semiprecision grips, and have been able to use both simple tools and even weapons, such as improvising a club from a convenient fallen branch.


Scientific study

American physician and missionary Thomas Staughton Savage obtained the first specimens (the skull and other bones) during his time in
Liberia Liberia (), officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to its north, Ivory Coast to its east, and the Atlantic Ocean ...
. The first scientific description of gorillas dates back to an article by Savage and the naturalist
Jeffries Wyman Jeffries Wyman (August 11, 1814 – September 4, 1874) was an American naturalist and anatomist, born in Chelmsford, Massachusetts. Wyman died in Bethlehem, New Hampshire of a pulmonary hemorrhage. Career He graduated Harvard College in 183 ...
in 1847 in ''Proceedings of the Boston Society of Natural History'', where ''Troglodytes gorilla'' is described, now known as the
western gorilla The western gorilla (''Gorilla gorilla'') is a great ape found in Africa, one of two species of the hominid genus ''Gorilla''. Large and robust with males weighing around , the hair is significantly lighter in color than the eastern gorilla, ''G ...
. Other species of gorilla were described in the next few years. The explorer Paul Du Chaillu was the first westerner to see a live gorilla during his travel through western equatorial Africa from 1856 to 1859. He brought dead specimens to the UK in 1861. The first systematic study was not conducted until the 1920s, when Carl Akeley of the
American Museum of Natural History The American Museum of Natural History (abbreviated as AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. In Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 26 inter ...
traveled to Africa to hunt for an animal to be shot and stuffed. On his first trip, he was accompanied by his friends
Mary Bradley Mary Bradley (born 10 May 1942) is a Social Democratic and Labour Party (SDLP) politician who was a Member of the Northern Ireland Assembly (MLA) for Foyle from 2003 to 2011. In 1985, she was elected to Derry City Council, representing the Soc ...
, a mystery writer, her husband, and their young daughter Alice, who would later write science fiction under the pseudonym
James Tiptree Jr. Alice Bradley Sheldon (born Alice Hastings Bradley; August 24, 1915 – May 19, 1987) was an American science fiction and fantasy author better known as James Tiptree, Jr., a pen name she used from 1967 to her death. It was not publicly known un ...
After their trip, Mary Bradley wrote ''On the Gorilla Trail''. She later became an advocate for the conservation of gorillas, and wrote several more books (mainly for children). In the late 1920s and early 1930s,
Robert Yerkes Robert Mearns Yerkes (; May 26, 1876 – February 3, 1956) was an American psychologist, ethologist, eugenicist and primatologist best known for his work in intelligence testing and in the field of comparative psychology. Yerkes was a pioneer ...
and his wife Ava helped further the study of gorillas when they sent Harold Bigham to Africa. Yerkes also wrote a book in 1929 about the great apes. After World War II,
George Schaller George Beals Schaller (born 1933) is a German-born American mammalogist, biologist, conservationist and author. Schaller is recognized by many as the world's preeminent field biologist, studying wildlife throughout Africa, Asia and South Americ ...
was one of the first researchers to go into the field and study primates. In 1959, he conducted a systematic study of the mountain gorilla in the wild and published his work. Years later, at the behest of
Louis Leakey Louis Seymour Bazett Leakey (7 August 1903 – 1 October 1972) was a Kenyan-British palaeoanthropologist and archaeologist whose work was important in demonstrating that humans evolved in Africa, particularly through discoveries made at Olduvai ...
and the ''
National Geographic ''National Geographic'' (formerly the ''National Geographic Magazine'', sometimes branded as NAT GEO) is a popular American monthly magazine published by National Geographic Partners. Known for its photojournalism, it is one of the most widely ...
'', Dian Fossey conducted a much longer and more comprehensive study of the mountain gorilla. When she published her work, many misconceptions and myths about gorillas were finally disproved, including the myth that gorillas are violent. Western lowland gorillas (''G. g. gorilla'') are believed to be one of the zoonotic origins of HIV/AIDS. The SIVgor Simian immunodeficiency virus that infects them is similar to a certain strain of HIV-1.


Genome sequencing

The gorilla became the next-to-last great ape genus to have its genome sequenced. The first gorilla genome was generated with short read and Sanger sequencing using DNA from a female western lowland gorilla named Kamilah. This gave scientists further insight into the evolution and origin of humans. Despite the chimpanzees being the closest extant relatives of humans, 15% of the human genome was found to be more like that of the gorilla. In addition, 30% of the gorilla genome "is closer to human or chimpanzee than the latter are to each other; this is rarer around coding genes, indicating pervasive selection throughout great ape evolution, and has functional consequences in gene expression." Analysis of the gorilla genome has cast doubt on the idea that the rapid evolution of hearing genes gave rise to language in humans, as it also occurred in gorillas.


Captivity

Gorillas became highly prized by western zoos since the 19th century, though the earliest attempts to keep them in captive facilities ended in their early death. In the late 1920s the care of captive gorillas significantly improved. Colo (December 22, 1956 – January 17, 2017) of the Columbus Zoo and Aquarium was the first gorilla to be born in captivity. Captive gorillas exhibit stereotypic behaviors, including eating disorders — such as regurgitation, reingestion and coprophagy — self-injurious or conspecific aggression, pacing, rocking, sucking of fingers or lip smacking, and overgrooming. Negative vigilance of visitor behaviors have been identified as starting, posturing and charging at visitors. Groups of bachelor gorillas containing young silverbacks have significantly higher levels of aggression and wounding rates than mixed age and sex groups. The use of both internal and external privacy screens on exhibit windows has been shown to alleviate stresses from visual effects of high crowd densities, leading to decreased stereotypic behaviors in the gorillas. Playing naturalistic auditory stimuli as opposed to classical music, rock music, or no auditory enrichment (which allows for crowd noise, machinery, etc. to be heard) has been noted to reduce stress behavior as well. Enrichment modifications to feed and foraging, where clover-hay is added to an exhibit floor, decrease stereotypic activities while simultaneously increasing positive food-related behaviors. Recent research on captive gorilla welfare emphasizes a need to shift to individual assessments instead of a one-size-fits-all group approach to understanding how welfare increases or decreases based on a variety of factors. Individual characteristics such as age, sex, personality and individual histories are essential in understanding that stressors will affect each individual gorilla and their welfare differently.


Conservation status

All species (and subspecies) of gorilla are listed as endangered or critically endangered on the IUCN Red List. All gorillas are listed in Appendix I of the Convention on International Trade in Endangered Species (CITES), meaning that international export/import of the species, including in parts and derivatives, is regulated. Around 316,000 western lowland gorillas are thought to exist in the wild, 4,000 in zoos, thanks to conservation; eastern lowland gorillas have a population of under 5,000 in the wild and 24 in zoos. Mountain gorillas are the most severely endangered, with an estimated population of about 880 left in the wild and none in zoos. Threats to gorilla survival include habitat destruction and poaching for the
bushmeat Bushmeat is meat from wildlife species that are hunted for human consumption, most often referring to the meat of game in Africa. Bushmeat represents a primary source of animal protein and a cash-earning commodity for inhabitants of humid tropi ...
trade. Gorillas are closely related to humans, and are susceptible to diseases that humans also get infected by. In 2004, a population of several hundred gorillas in the
Odzala National Park Odzala-Kokoua National Park (or Odzala National Park) is a national park in the Republic of the Congo. The park was first protected in 1935, declared a biosphere reserve in 1977, and granted official designation by presidential decree in 2001. Odz ...
,
Republic of Congo The Republic of the Congo (french: République du Congo, ln, Republíki ya Kongó), also known as Congo-Brazzaville, the Congo Republic or simply either Congo or the Congo, is a country located in the western coast of Central Africa to the w ...
was essentially wiped out by the Ebola virus. A 2006 study published in '' Science'' concluded more than 5,000 gorillas may have died in recent outbreaks of the Ebola virus in central Africa. The researchers indicated in conjunction with commercial hunting of these apes, the virus creates "a recipe for rapid
ecological extinction Ecological extinction is "the reduction of a species to such low abundance that, although it is still present in the community, it no longer interacts significantly with other species". Ecological extinction stands out because it is the interacti ...
". In captivity, its also been observed that gorillas can also be infected with COVID-19. Conservation efforts include the
Great Apes Survival Project The Great Apes Survival Partnership (GRASP) is a UNEP and UNESCO-led World Summit on Sustainable Development (WSSD) Type II Partnership, established in 2001, that aims to conserve the non-human great apes (chimpanzees, bonobos, gorillas and oranguta ...
, a partnership between the United Nations Environment Programme and the UNESCO, and also an international treaty, the Agreement on the Conservation of Gorillas and Their Habitats, concluded under UNEP-administered Convention on Migratory Species. The
Gorilla Agreement The Agreement on the Conservation of Gorillas and Their Habitats, also known as the Gorilla Agreement, is a multilateral environmental agreement that binds the parties to conserve gorillas in their territories. It was concluded 2007 under the aus ...
is the first legally binding instrument exclusively targeting gorilla conservation; it came into effect on 1 June 2008. Governments of countries where gorillas live placed a ban on their killing and trading, but weak law enforcement still poses a threat to them, since the governments rarely apprehend poachers, traders and consumers that rely on gorillas for profit.


Cultural significance

In Cameroon's Lebialem highlands, folk stories connect people and gorillas via totems; a gorilla's death means the connected person will die also. This creates a local conservation ethic. Many different indigenous peoples interact with wild gorillas. Some have detailed knowledge; the
Baka Baka, baká or BAKA may refer to: Ethnicities and languages * Baka people (Cameroon and Gabon), an African ethnic group * Baka people (Congo and South Sudan), an African ethnic group * Baka language, a dialect cluster of Cameroon and Gabon * Baka ...
have words to distinguish at least ten types of gorilla individuals, by sex, age, and relationships. In 1861, alongside tales of hunting enormous gorillas, the traveller and anthropologist Paul Du Chaillu reported the Cameroonian story that a pregnant woman who sees a gorilla will give birth to one. In 1911, the anthropologist Albert Jenks noted the Bulu people's knowledge of gorilla behaviour and ecology, and their gorilla stories. In one such story, "The Gorilla and the Child", a gorilla speaks to people, seeking help and trust, and stealing a baby; a man accidentally kills the baby while attacking the gorilla. Even far from where gorillas live, savannah tribes pursue "cult-like worship" of the apes. Some beliefs are widespread among indigenous peoples. The
Fang A fang is a long, pointed tooth. In mammals, a fang is a modified maxillary tooth, used for biting and tearing flesh. In snakes, it is a specialized tooth that is associated with a venom gland (see snake venom). Spiders also have external fang ...
name for gorilla is ''ngi'' while the Bulu name is ''njamong''; the root ''ngi'' means fire, denoting a positive energy. From the Central African Republic to Cameroon and Gabon, stories of reincarnations as gorillas, totems, and transformations similar to those recorded by Du Chaillu are still told in the 21st century. Since gaining international attention, gorillas have been a recurring element of many aspects of popular culture and media. They were usually portrayed as murderous and aggressive. Inspired by Emmanuel Frémiet's ''Gorilla Carrying off a Woman'', gorillas have been depicted kidnapping human women. This theme was used in films such as ''
Ingagi ''Ingagi'' is a 1930 pre-Code mockumentary exploitation film directed by William S. Campbell. It purports to be a documentary about "Sir Hubert Winstead" of London on an expedition to the Belgian Congo, and depicts a tribe of gorilla-worshipping ...
'' (1930) and most notably '' King Kong'' (1933). The comedic play '' The Gorilla'', which debuted in 1925, featured an escaped gorilla taking a woman from her house. Several films would use the "escaped gorilla" trope including ''
The Strange Case of Doctor Rx ''The Strange Case of Doctor Rx'' is a 1942 black-and-white murder mystery/ horror B film by Universal Studios directed by William Nigh and starring Patric Knowles, Lionel Atwill, Anne Gwynne, Ray "Crash" Corrigan and Samuel S. Hinds. Although ...
'' (1942), '' The Gorilla Man'' (1943), ''
Gorilla at Large ''Gorilla at Large'' is a 1954 American horror mystery film made in 3-D. The film stars Cameron Mitchell, Anne Bancroft, Lee J. Cobb and Raymond Burr, with Lee Marvin and Warren Stevens in supporting roles. Directed by Harmon Jones, it was ma ...
'' (1954) and the Disney cartoons ''
The Gorilla Mystery ''The Gorilla Mystery'' is a 1930 Mickey Mouse animated film produced by Walt Disney for Columbia Pictures, as part of the ''Mickey Mouse'' film series. It was the twenty-second Mickey Mouse short to be produced, the seventh of that year. The sh ...
'' (1930) and ''
Donald Duck and the Gorilla ''Donald Duck and the Gorilla'' is a Donald Duck short adventure horror monster film which was originally released in 1944. It stars Donald Duck along with his three nephews; Huey, Dewey and Louie and Ajax, the killer gorilla. The 1930 Mickey Mo ...
'' (1944). Gorillas have been used as opponents to jungle-themed heroes such as Tarzan and Sheena, Queen of the Jungle, as well as superheroes. The DC comics supervillain
Gorilla Grodd Gorilla Grodd is a supervillain character appearing in American comic books and other media published by DC Comics, primarily as an enemy of The Flash. The character was created by John Broome and Carmine Infantino, and first appeared in ''Th ...
is an enemy of
the Flash The Flash (or simply Flash) is the name of several superheroes appearing in American comic books published by DC Comics. Created by writer Gardner Fox and artist Harry Lampert, the original Flash first appeared in ''Flash Comics'' #1 (cover date ...
. Gorillas also serve as antagonists in the 1968 film '' Planet of the Apes''. More positive and sympathetic portrayals of gorillas include the films '' Son of Kong'' (1933), '' Mighty Joe Young'' (1949), '' Gorillas in the Mist'' (1988) and ''
Instinct Instinct is the inherent inclination of a living organism towards a particular complex behaviour, containing both innate (inborn) and learned elements. The simplest example of an instinctive behaviour is a fixed action pattern (FAP), in which a v ...
'' (1999) and the 1992 novel ''
Ishmael Ishmael ''Ismaḗl''; Classical/Qur'anic Arabic: إِسْمَٰعِيْل; Modern Standard Arabic: إِسْمَاعِيْل ''ʾIsmāʿīl''; la, Ismael was the first son of Abraham, the common patriarch of the Abrahamic religions; and is cons ...
''. Gorillas have been featured in video games as well, notably '' Donkey Kong''.


See also

*
Bili ape The Bili apes or Bondo mystery apes were names given in 2003 in sensational reports in the popular media to a purportedly new species of highly aggressive, giant ape supposedly inhabiting the wetlands and savannah around of the village of Bili in ...
*
Gorilla (advertisement) ''Gorilla'' is a British advertising campaign launched by the advertising agency Fallon London on behalf of Cadbury Schweppes in 2007, to promote Cadbury Dairy Milk brand chocolate. The centrepiece of the campaign was a 90-second television a ...
* Gorilla suit *
Great Ape Project The Great Ape Project (GAP), founded in 1993, is an international organization of primatologists, anthropologists, ethicists, and others who advocate a United Nations Declaration of the Rights of Great Apes that would confer basic legal rights ...
* Harambe – a captive gorilla, shot while interacting with a child who fell into his zoo enclosure, who became a popular internet meme *
International Primate Day International Primate Day, September 1, is an annual educational observance event organized since 2005 largely by British-based Animal Defenders International (ADI) and supported annually by various primate-oriented advocacy organizations, speaks ...
*
List of individual apes This is a list of non-human apes of encyclopedic interest. It includes individual chimpanzees, gorillas, orangutans, bonobos, and gibbons that are in some way famous or notable. Actors * Bam Bam, an orangutan, played Precious (Passions), Preciou ...
*
List of fictional apes This list of fictional primates is a subsidiary to the list of fictional animals. The list is restricted to notable non-human primate characters from the world of fiction including chimpanzees, gorillas, orangutans, bonobos, gibbons, monkeys, le ...
* Monkey Day


References


Literature cited

* * ** ** ** ** ** ** ** **


External links


Animal Diversity Web
– includes photos, artwork, and skull specimens of ''Gorilla gorilla''
Pulitzer Center on Crisis Reporting
International Gorilla Conservation Programme The International Gorilla Conservation Programme (IGCP) was formed in 1991 to ensure that the critically endangered mountain gorillas are conserved in their habitat in the mountain forests of the Virunga Massif in Rwanda, Uganda and the Democrati ...
(Video)
Primate Info Net ''Gorilla'' Factsheet
– taxonomy, ecology, behavior and conservation

– features a video and photos
World Wildlife Fund: Gorillas
– conservation, facts and photos
Gorilla protection
– Gorilla conservation
Welcome to the Year of the Gorilla 2009

Virunga National Park
– The Official Website for Virunga National Park, the Last Refuge for Congo's Mountain Gorillas
Human Timeline (Interactive)
Smithsonian, National Museum of Natural History (August 2016).
Genome of ''Gorilla gorilla''
via Ensembl
Genome of ''Gorilla gorilla'' (version Kamilah_GGO_v0/gorGor6)
via UCSC Genome Browser
Data of the genome of ''Gorilla gorilla''
via NCBI
Data of the genome assembly of ''Gorilla gorilla'' (version Kamilah_GGO_v0/gorGor6)
via NCBI {{Authority control   Apes Herbivorous mammals   Fauna of Sub-Saharan Africa Primate genera Taxa named by Isidore Geoffroy Saint-Hilaire Taxa described in 1852