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The godwits are a group of large, long-billed, long-legged and strongly migratory
wader 245px, A flock of Dunlins and Red knots">Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflat ...
s of the
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Limosa''. Their long bills allow them to probe deeply in the sand for aquatic worms and
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
s. In their winter range, they flock together where food is plentiful. They frequent tidal shorelines, breeding in northern climates in summer and migrating south in winter. A female
bar-tailed godwit The bar-tailed godwit (''Limosa lapponica'') is a large and strongly migratory wader in the family Scolopacidae, which feeds on bristle-worms and shellfish on coastal mudflats and estuaries. It has distinctive red breeding plumage, long legs, an ...
made a flight of 29,000 km (18,000 mi), flying of it without stopping. In 2020 a male bar-tailed godwit flew about non-stop in its migration from Alaska to New Zealand, previously a record for avian non-stop flight. In October 2022, a 5 month old, male bar-tailed godwit was tracked from
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
to
Tasmania ) , nickname = , image_map = Tasmania in Australia.svg , map_caption = Location of Tasmania in AustraliaCoordinates: , subdivision_type = Country , subdi ...
, a trip that took 11 days, and recorded a non-stop flight of . The godwits can be distinguished from the
curlew The curlews () are a group of nine species of birds in the genus ''Numenius'', characterised by their long, slender, downcurved bills and mottled brown plumage. The English name is imitative of the Eurasian curlew's call, but may have been in ...
s by their straight or slightly upturned bills, and from the
dowitcher The three dowitchers are medium-sized long-billed wading birds in the genus ''Limnodromus''. The English name "dowitchers" is from Iroquois, recorded in English by the 1830s. The OED's earliest example is from 1841, but full-text searching giv ...
s by their longer legs. The winter plumages are fairly drab, but three species have reddish underparts when breeding. The females are appreciably larger than the males. Godwits were once a popular British dish. Sir
Thomas Browne Sir Thomas Browne (; 19 October 160519 October 1682) was an English polymath and author of varied works which reveal his wide learning in diverse fields including science and medicine, religion and the esoteric. His writings display a deep curi ...
writing in about 1682 noted that godwits "were accounted the daintiest dish in England".


Taxonomy

The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
''Limosa'' was introduced by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosopher. Brisson was born at Fontenay-le-Comte. The earlier part of his life was spent in the pursuit of natural history; his published works ...
in 1760 with the black-tailed godwit (''Limosa limosa'') as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
. The genus name ''Limosa'' is from
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
and means "muddy", from ''limus'', "mud". The English name "godwit" was first recorded in about 1416–17 and is believed to imitate the bird's call. The genus contains four living species: *
Bar-tailed godwit The bar-tailed godwit (''Limosa lapponica'') is a large and strongly migratory wader in the family Scolopacidae, which feeds on bristle-worms and shellfish on coastal mudflats and estuaries. It has distinctive red breeding plumage, long legs, an ...
, ''Limosa lapponica'' * Black-tailed godwit, ''Limosa limosa'' *
Hudsonian godwit The Hudsonian godwit (''Limosa haemastica'') is a large shorebird in the sandpiper family, Scolopacidae. The genus name ''Limosa'' is from Latin and means "muddy", from ''limus'', "mud". The specific ''haemastica'' is from Ancient Greek and means ...
, ''Limosa haemastica'' *
Marbled godwit The marbled godwit (''Limosa fedoa'') is a large migratory shorebird in the family Scolopacidae. On average, it is the largest of the four species of godwit. Taxonomy In 1750 the English naturalist George Edwards included an illustration and a ...
, ''Limosa fedoa''


Fossil species

In addition, there are two or three species of
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
prehistoric godwits. ''Limosa vanrossemi'' is known from the
Monterey Formation The Monterey Formation is an extensive Miocene oil-rich geological sedimentary formation in California, with outcrops of the formation in parts of the California Coast Ranges, Peninsular Ranges, and on some of California's off-shore islan ...
(Late
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
, approx. 6 mya) of
Lompoc Lompoc ( ; Chumash: ''Lum Poc'') is a city in Santa Barbara County, California. Located on the Central Coast, Lompoc has a population of 43,834 as of July 2021. Lompoc has been inhabited for thousands of years by the Chumash people, who called ...
, United States. ''Limosa lacrimosa'' is known from the Early Pliocene of Western Mongolia ( Kurochkin, 1985). ''Limosa gypsorum'' of the Late
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
( Montmartre Formation, some 35 mya) of France may have actually been a
curlew The curlews () are a group of nine species of birds in the genus ''Numenius'', characterised by their long, slender, downcurved bills and mottled brown plumage. The English name is imitative of the Eurasian curlew's call, but may have been in ...
or some bird ancestral to both curlews and godwits (and possibly other
Scolopacidae Sandpipers are a large family, Scolopacidae, of waders. They include many species called sandpipers, as well as those called by names such as curlew and snipe. The majority of these species eat small invertebrates picked out of the mud or soil. ...
), or even a
rail Rail or rails may refer to: Rail transport *Rail transport and related matters *Rail (rail transport) or railway lines, the running surface of a railway Arts and media Film * ''Rails'' (film), a 1929 Italian film by Mario Camerini * ''Rail'' ( ...
, being placed in the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispec ...
genus '' Montirallus'' by some (Olson, 1985). Certainly, curlews and godwits are rather ancient and in some respects primitive lineages of scolopacids, further complicating the assignment of such possibly basal forms. In a 2001 study comparing the ratios cerebrum to brain volumes in various dinosaur species, Hans C. E. Larsson found that more derived dinosaurs generally had proportionally more voluminous cerebrum. ''Limosa gypsorum'', then regarded as a '' Numenius'' species, was a discrepancy in this general trend. ''L. gypsorum'' was only 63% of the way between a typical reptilian ratio and that of modern birds. However, this may be explainable if the endocast was distorted, as it had been previously depicted in the past by Deschaseaux, who is described by Larsson as calling the endocast "slightly anteroposteriorly sheared and laterally compressed."


Citations


General sources

* * Larsson, H. C. E. 2001. Endocranial anatomy of ''Carcharodontosaurus saharicus'' (Theropoda: Allosauroidea) and its implications for theropod brain evolution. pp. 19–33. In: ''Mesozoic Vertebrate Life''. Tanke, D. H., Carpenter, K., Skrepnick, M. W. (eds.). Indiana University Press. * Olson, Storrs L. (1985): Section X.D.2.b. "Scolopacidae". In: Farner, D.S.; King, J.R. & Parkes, Kenneth C. (eds.): ''Avian Biology'' 8: 174–175. Academic Press, New York. {{Taxonbar, from=Q585636 * Taxa named by Mathurin Jacques Brisson