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''Enteroctopus'' is an
octopus An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttle ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
whose members are sometimes known as giant octopus.


Etymology

The generic name ''Enteroctopus'' was created by
Alphonse Tremeau de Rochebrune Alphonse may refer to: * Alphonse (given name) * Alphonse (surname) * Alphonse Atoll, one of two atolls in the Seychelles' Alphonse Group See also *Alphons *Alfonso (disambiguation) Alfonso (and variants Alphonso, Afonso, Alphons, and Alphonse) is ...
and Jules François Mabille in 1887 and published in 1889, joining
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic peri ...
'gut' and , thus 'octopus ith arms similar toguts.'


Description

''Enteroctopus'' is a genus of generally temperate
octopus An octopus ( : octopuses or octopodes, see below for variants) is a soft-bodied, eight- limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttle ...
es. Members of this genus are characterized by their large size and are often known as giant octopuses. ''Enteroctopus'' species have distinct longitudinal wrinkles or folds dorsally and laterally on their bodies. Their heads are distinctly narrower than the mantle width. The
hectocotylus A hectocotylus (plural: ''hectocotyli'') is one of the arms of male cephalopods that is specialized to store and transfer spermatophores to the female. Structurally, hectocotyli are muscular hydrostats. Depending on the species, the male may use i ...
of the males in this genus, found on the third right arm, is long and narrow in comparison with other genera in the family
Octopodidae The Octopodidae comprise the family containing the majority of known octopus species. Genera The World Register of Marine Species lists these genera: *'' Abdopus'' Norman & Finn, 2001 *'' Ameloctopus'' Norman, 1992 *'' Amphioctopus'' P. Fischer, ...
, often comprising one-fifth the length of the arm. Octopuses in this genus have large, paddle-like
papilla Papilla (Latin, 'nipple') or papillae may refer to: In animals * Papilla (fish anatomy), in the mouth of fish * Basilar papilla, a sensory organ of lizards, amphibians and fish * Dental papilla, in a developing tooth * Dermal papillae, part of ...
e instead of the more conical papillae in other octopus genera.


Species

Genus ''Enteroctopus'' at present consists of four species, tabulated below:


Type species

''E. membranaceus'' has often been regarded as
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
of the genus, not because it was designated as such by
Rochebrune Rochebrune may refer to the following places in France: * Rochebrune, Hautes-Alpes, a commune in the department of Hautes-Alpes * Rochebrune, Drôme, a commune in the department of Drôme * Pic de Rochebrune, a mountain of Cottian Alps * Pic de P ...
and Mabille when they erected the genus, but because it was the first named species in the genus. Robson in his 1929 monograph of octopods regarded ''E. membranaceus'' as a '' species dubium'' because the original description was insufficient to identify an individual species, the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
was an immature specimen, and the type specimen no longer existed. As such, the genus was considered invalid until Hochberg resurrected it in 1998. Hochberg noted that Robson had considered ''E. membranaceus'' a
junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...
of '' E. megalocyathus'', the second species assigned to the genus by Rochebrune and Mabille in their 1889 description. Additionally, since Rochebrune and Mabille did not actually assign type status to ''E. membranaceus'', Hochberg concluded that ''Enteroctopus'' was indeed a valid genus and transferred type-species status to ''E. megalocyathus'' based on his conclusion that ''E. megalocyathus'' and ''E. membranaceus'' are the same species.


Distribution

Species in the genus ''Enteroctopus'' are restricted to the temperate areas of the
Northern Northern may refer to the following: Geography * North, a point in direction * Northern Europe, the northern part or region of Europe * Northern Highland, a region of Wisconsin, United States * Northern Province, Sri Lanka * Northern Range, a ra ...
and Southern Hemispheres. ''E. dofleini'' is the only member of the genus found in the Northern Hemisphere, and also the most widely distributed: It is found from
Southern California Southern California (commonly shortened to SoCal) is a geographic and Cultural area, cultural region that generally comprises the southern portion of the U.S. state of California. It includes the Los Angeles metropolitan area, the second most po ...
, along the North Pacific Rim to
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
, including the
Okhotsk Okhotsk ( rus, Охотск, p=ɐˈxotsk) is an types of inhabited localities in Russia, urban locality (a urban-type settlement, work settlement) and the administrative center of Okhotsky District of Khabarovsk Krai, Russia, located at the mou ...
and
Bering Sea The Bering Sea (, ; rus, Бе́рингово мо́ре, r=Béringovo móre) is a marginal sea of the Northern Pacific Ocean. It forms, along with the Bering Strait, the divide between the two largest landmasses on Earth: Eurasia and The Ameri ...
s. The other three species are found in the Southern Hemisphere; ''E. megalocyathus'' occurs on the southeastern coast of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, ''E. magnificus'' on the southwestern coast of
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
from
Namibia Namibia (, ), officially the Republic of Namibia, is a country in Southern Africa. Its western border is the Atlantic Ocean. It shares land borders with Zambia and Angola to the north, Botswana to the east and South Africa to the south and ea ...
to
Port Elizabeth Gqeberha (), formerly Port Elizabeth and colloquially often referred to as P.E., is a major seaport and the most populous city in the Eastern Cape province of South Africa. It is the seat of the Nelson Mandela Bay Metropolitan Municipality, Sou ...
,
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
, and ''E. zealandicus'' in temperate
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...
.


Size

The member of this genus that best embodies the common name "giant octopus" is ''
Enteroctopus dofleini The giant Pacific octopus (''Enteroctopus dofleini''), also known as the North Pacific giant octopus, is a large marine cephalopod belonging to the genus '' Enteroctopus''. Its spatial distribution includes the coastal North Pacific, along Mexic ...
'', which holds the record of being the world's largest octopus based on direct measurements of a individual, weighed live. This octopus had a total length near to . The remaining members of the genus are substantially smaller, with ''E. megalocyathus'' having an average mass of and reaching a total length of . ''E. magnificus'' reaches a total length of around .


See also

*
Gigantic octopus The kraken () is a legendary sea monster of enormous size said to appear off the coasts of Norway. Kraken, the subject of sailors' superstitions and mythos, was first described in the modern age at the turn of the 18th century, in a travelogu ...
*
Giant squid The giant squid (''Architeuthis dux'') is a species of deep-ocean dwelling squid in the family Architeuthidae. It can grow to a tremendous size, offering an example of abyssal gigantism: recent estimates put the maximum size at around Trace ...
*
Colossal squid The colossal squid (''Mesonychoteuthis hamiltoni'') is part of the family Cranchiidae. It is sometimes called the Antarctic squid or giant cranch squid and is believed to be the largest squid species in terms of mass. It is the only recognize ...
*
Cephalopod size Cephalopods vary enormously in size. The smallest are only about long and weigh less than at maturity, while the largest—the giant and colossal squids—can exceed in length and weigh close to half a tonne (), making them the largest living ...
*
Kraken The kraken () is a legendary sea monster of enormous size said to appear off the coasts of Norway. Kraken, the subject of sailors' superstitions and mythos, was first described in the modern age at the turn of the 18th century, in a travelogu ...
*
Lusca In Caribbean folklore, the Lusca is a name given to a sea monster said to exist in the region of the blue holes nearby Andros, an island in the Bahamas. Description It is described as a giant octopus, a giant cuttlefish, or a half dragon, half oct ...


Footnotes


References

{{Authority control Enteroctopodidae Taxa named by Jules François Mabille