Georg Von Békésy
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Georg von Békésy (, ; 3 June 1899 – 13 June 1972) was a Hungarian-American
biophysicist Biophysics is an interdisciplinary science that applies approaches and methods traditionally used in physics to study biological phenomena. Biophysics covers all scales of biological organization, from molecular to organismic and populations ...
. By using
strobe A strobe light or stroboscopic lamp, commonly called a strobe, is a device used to produce regular flashes of light. It is one of a number of devices that can be used as a stroboscope. The word originated from the Ancient Greek ('), meaning ...
photography and
silver Silver is a chemical element; it has Symbol (chemistry), symbol Ag () and atomic number 47. A soft, whitish-gray, lustrous transition metal, it exhibits the highest electrical conductivity, thermal conductivity, and reflectivity of any metal. ...
flakes as a marker, he was able to observe that the
basilar membrane The basilar membrane is a stiff structural element within the cochlea of the inner ear which separates two liquid-filled tubes that run along the coil of the cochlea, the scala media and the scala tympani. The basilar membrane moves up and down ...
moves like a
surface wave In physics, a surface wave is a mechanical wave that propagates along the Interface (chemistry), interface between differing media. A common example is gravity waves along the surface of liquids, such as ocean waves. Gravity waves can also occu ...
when stimulated by
sound In physics, sound is a vibration that propagates as an acoustic wave through a transmission medium such as a gas, liquid or solid. In human physiology and psychology, sound is the ''reception'' of such waves and their ''perception'' by the br ...
. Because of the structure of the
cochlea The cochlea is the part of the inner ear involved in hearing. It is a spiral-shaped cavity in the bony labyrinth, in humans making 2.75 turns around its axis, the modiolus (cochlea), modiolus. A core component of the cochlea is the organ of Cort ...
and the basilar membrane, different
frequencies Frequency is the number of occurrences of a repeating event per unit of time. Frequency is an important parameter used in science and engineering to specify the rate of oscillatory and vibratory phenomena, such as mechanical vibrations, audio ...
of sound cause the maximum amplitudes of the waves to occur at different places on the basilar membrane along the coil of the
cochlea The cochlea is the part of the inner ear involved in hearing. It is a spiral-shaped cavity in the bony labyrinth, in humans making 2.75 turns around its axis, the modiolus (cochlea), modiolus. A core component of the cochlea is the organ of Cort ...
. High frequencies cause more vibration at the base of the cochlea while low frequencies create more vibration at the apex. He concluded that his observations showed how different sound wave frequencies are locally dispersed before exciting different
nerve fiber An axon (from Greek ἄξων ''áxōn'', axis) or nerve fiber (or nerve fibre: see spelling differences) is a long, slender projection of a nerve cell, or neuron, in vertebrates, that typically conducts electrical impulses known as action pot ...
s that lead from the cochlea to the brain. In 1961, he was awarded the
Nobel Prize in Physiology or Medicine The Nobel Prize in Physiology or Medicine () is awarded yearly by the Nobel Assembly at the Karolinska Institute for outstanding discoveries in physiology or medicine. The Nobel Prize is not a single prize, but five separate prizes that, acco ...
for his research on the function of the
cochlea The cochlea is the part of the inner ear involved in hearing. It is a spiral-shaped cavity in the bony labyrinth, in humans making 2.75 turns around its axis, the modiolus (cochlea), modiolus. A core component of the cochlea is the organ of Cort ...
in the mammalian hearing organ.


Biography

Békésy was born on 3 June 1899 in
Budapest Budapest is the Capital city, capital and List of cities and towns of Hungary, most populous city of Hungary. It is the List of cities in the European Union by population within city limits, tenth-largest city in the European Union by popul ...
,
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
, as the first of three children (György 1899, Lola 1901 and Miklós 1903) to Sándor Békésy (1860–1923), an economic diplomat, and to his mother Paula Mazaly. The Békésy family was originally
Reformed Reform is beneficial change. Reform, reformed or reforming may also refer to: Media * ''Reform'' (album), a 2011 album by Jane Zhang * Reform (band), a Swedish jazz fusion group * ''Reform'' (magazine), a Christian magazine Places * Reform, Al ...
but converted to
Catholicism The Catholic Church (), also known as the Roman Catholic Church, is the List of Christian denominations by number of members, largest Christian church, with 1.27 to 1.41 billion baptized Catholics Catholic Church by country, worldwid ...
. His mother, Paula Mazaly (1877–1974) was born in Szagolyca (now Čađavica,
Croatia Croatia, officially the Republic of Croatia, is a country in Central Europe, Central and Southeast Europe, on the coast of the Adriatic Sea. It borders Slovenia to the northwest, Hungary to the northeast, Serbia to the east, Bosnia and Herze ...
). His maternal grandfather was from
Pécs Pécs ( , ; ; Slovak language, Slovak: ''Päťkostolie''; also known by #Name, alternative names) is List of cities and towns of Hungary#Largest cities in Hungary, the fifth largest city in Hungary, on the slopes of the Mecsek mountains in the c ...
. His father was born in Kolozsvár (now Cluj-Napoca,
Romania Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...
). Békésy went to school in Budapest,
Munich Munich is the capital and most populous city of Bavaria, Germany. As of 30 November 2024, its population was 1,604,384, making it the third-largest city in Germany after Berlin and Hamburg. Munich is the largest city in Germany that is no ...
, and
Zürich Zurich (; ) is the list of cities in Switzerland, largest city in Switzerland and the capital of the canton of Zurich. It is in north-central Switzerland, at the northwestern tip of Lake Zurich. , the municipality had 448,664 inhabitants. The ...
. He studied
chemistry Chemistry is the scientific study of the properties and behavior of matter. It is a physical science within the natural sciences that studies the chemical elements that make up matter and chemical compound, compounds made of atoms, molecules a ...
in
Bern Bern (), or Berne (), ; ; ; . is the ''de facto'' Capital city, capital of Switzerland, referred to as the "federal city".; ; ; . According to the Swiss constitution, the Swiss Confederation intentionally has no "capital", but Bern has gov ...
and received his PhD in physics on the subject: "Fast way of determining molecular weight" from the
University of Budapest A university () is an institution of tertiary education and research which awards academic degrees in several academic disciplines. ''University'' is derived from the Latin phrase , which roughly means "community of teachers and scholars". Univ ...
in 1926. He then spent one year working in an engineering firm. He published his first paper on the pattern of vibrations of the inner ear in 1928. He was offered a position at
Uppsala University Uppsala University (UU) () is a public university, public research university in Uppsala, Sweden. Founded in 1477, it is the List of universities in Sweden, oldest university in Sweden and the Nordic countries still in operation. Initially fou ...
by Róbert Bárány, which he declined because of the hard Swedish winters. Before and during
World War II World War II or the Second World War (1 September 1939 – 2 September 1945) was a World war, global conflict between two coalitions: the Allies of World War II, Allies and the Axis powers. World War II by country, Nearly all of the wo ...
, Békésy worked for the Hungarian Post Office (1923 to 1946), where he did research on telecommunications signal quality. This research led him to become interested in the workings of the ear. In 1946, he left Hungary to follow this line of research at the
Karolinska Institute The Karolinska Institute (KI; ; sometimes known as the (Royal) Caroline Institute in English) is a research-led medical university in Solna within the Stockholm urban area of Sweden and one of the foremost medical research institutes globally ...
in Sweden. In 1947, he moved to the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
, working at
Harvard University Harvard University is a Private university, private Ivy League research university in Cambridge, Massachusetts, United States. Founded in 1636 and named for its first benefactor, the History of the Puritans in North America, Puritan clergyma ...
until 1966. In 1962 he was elected a Member of the
German Academy of Sciences Leopoldina The German National Academy of Sciences Leopoldina (), in short Leopoldina, is the national academy of Germany, and is located in Halle (Saale). Founded on 1 January 1652, based on academic models in Italy, it was originally named the ''Academi ...
. After his lab was destroyed by fire in 1965, he was invited to lead a research laboratory of sense organs in Honolulu, Hawaii. He became a professor at the
University of Hawaii A university () is an educational institution, institution of tertiary education and research which awards academic degrees in several Discipline (academia), academic disciplines. ''University'' is derived from the Latin phrase , which roughly ...
in 1966 and died in
Honolulu Honolulu ( ; ) is the List of capitals in the United States, capital and most populous city of the U.S. state of Hawaii, located in the Pacific Ocean. It is the county seat of the Consolidated city-county, consolidated City and County of Honol ...
. He became a well-known expert in
Asian art Asian art includes a vast range of arts from various cultures, regions, and religions across the continent of Asia. East Asian art includes works from China, Japan, and Korea, while Southeast Asian art includes the arts of Brunei, Cambodia, E ...
. He had a large collection which he donated to the Nobel Foundation in Sweden. His brother, Dr. Miklós Békésy (1903-1980), stayed in Hungary and became a famous agrobiologist who was awarded the
Kossuth Prize The Kossuth Prize (, ) is a state-sponsored award in Hungary, named after the Hungarian politician and revolutionist Lajos Kossuth. The Prize was established in 1936, by the Hungarian National Assembly, to acknowledge outstanding personal and grou ...
.


Research

Békésy contributed most notably to our understanding of the mechanism by which sound frequencies are registered in the inner ear. He developed a method for dissecting the inner ear of human
cadaver A cadaver, often known as a corpse, is a Death, dead human body. Cadavers are used by medical students, physicians and other scientists to study anatomy, identify disease sites, determine causes of death, and provide tissue (biology), tissue to ...
s while leaving the cochlea partly intact. By using
strobe A strobe light or stroboscopic lamp, commonly called a strobe, is a device used to produce regular flashes of light. It is one of a number of devices that can be used as a stroboscope. The word originated from the Ancient Greek ('), meaning ...
photography and
silver Silver is a chemical element; it has Symbol (chemistry), symbol Ag () and atomic number 47. A soft, whitish-gray, lustrous transition metal, it exhibits the highest electrical conductivity, thermal conductivity, and reflectivity of any metal. ...
flakes as a marker, he was able to observe that the
basilar membrane The basilar membrane is a stiff structural element within the cochlea of the inner ear which separates two liquid-filled tubes that run along the coil of the cochlea, the scala media and the scala tympani. The basilar membrane moves up and down ...
moves like a
surface wave In physics, a surface wave is a mechanical wave that propagates along the Interface (chemistry), interface between differing media. A common example is gravity waves along the surface of liquids, such as ocean waves. Gravity waves can also occu ...
when stimulated by
sound In physics, sound is a vibration that propagates as an acoustic wave through a transmission medium such as a gas, liquid or solid. In human physiology and psychology, sound is the ''reception'' of such waves and their ''perception'' by the br ...
. Because of the structure of the cochlea and the basilar membrane, different
frequencies Frequency is the number of occurrences of a repeating event per unit of time. Frequency is an important parameter used in science and engineering to specify the rate of oscillatory and vibratory phenomena, such as mechanical vibrations, audio ...
of sound cause the maximum amplitudes of the waves to occur at different places on the basilar membrane along the coil of the
cochlea The cochlea is the part of the inner ear involved in hearing. It is a spiral-shaped cavity in the bony labyrinth, in humans making 2.75 turns around its axis, the modiolus (cochlea), modiolus. A core component of the cochlea is the organ of Cort ...
.Goldstein, B. 2001. ''Sensation and Perception, 6th ed.'' London: Wadsworth. High frequencies cause more vibration at the base of the cochlea while low frequencies create more vibration at the apex.Lera Boroditsky. (1999) "Hearing I: Lecture Notes." pp. 3
/ref> Békésy concluded from these observations that by exciting different locations on the basilar membrane different sound wave frequencies excite different
nerve fiber An axon (from Greek ἄξων ''áxōn'', axis) or nerve fiber (or nerve fibre: see spelling differences) is a long, slender projection of a nerve cell, or neuron, in vertebrates, that typically conducts electrical impulses known as action pot ...
s that lead from the cochlea to the brain. He theorized that, due to its placement along the cochlea, each sensory cell (
hair cell Hair cells are the sensory receptors of both the auditory system and the vestibular system in the ears of all vertebrates, and in the lateral line organ of fishes. Through mechanotransduction, hair cells detect movement in their environment. ...
) responds maximally to a specific frequency of sound (the so-called
tonotopy In physiology, tonotopy (from Greek tono = frequency and topos = place) is the spatial arrangement of where sounds of different frequency are processed in the brain. Tones close to each other in terms of frequency are represented in topologically ...
). Békésy later developed a mechanical model of the cochlea, which confirmed the concept of frequency dispersion by the basilar membrane in the mammalian cochlea. In an article published posthumously in 1974, Békésy reviewed progress in the field, remarking "In time, I came to the conclusion that the dehydrated cats and the application of
Fourier analysis In mathematics, Fourier analysis () is the study of the way general functions may be represented or approximated by sums of simpler trigonometric functions. Fourier analysis grew from the study of Fourier series, and is named after Joseph Fo ...
to hearing problems became more and more a handicap for research in hearing," referring to the difficulties in getting animal preparations to behave as when alive, and the misleading common interpretations of Fourier analysis in hearing research.


Awards

Békésy's honours include: *The Denker Prize in Otology (1931), The Leibniz Medal of the Berlin Academy of Sciences (1937), The Guyot Prize for Speech and Otology of Groningen University (1939), The Academy Award of the Budapest Academy of Science (1946), Shambough Prize in Otology (1950). *Honorary doctorates (M.D.) were conferred on him by the University of Munster (1955), Bern (1959), Padua (1962), Buenos Aires (1968), Cordoba (1968), Hawaii (1969) and Semmelweiss University, Budapest (1969).


References


Further reading

* * * * * * * * * * * * * * * * * * * *


External links

* including the Nobel Lecture, December 11, 1961 ''Concerning the Pleasures of Observing, and the Mechanics of the Inner Ear''
Georg von Békésy
page at th
Pacific Biosciences Research Center

Békésy Laboratory of Neurobiology
website
Békésy art collection


autobiographical speech by von Békésy

{{DEFAULTSORT:Bekesy, Georg Von 1899 births 1972 deaths Scientists from Budapest Austrian untitled nobility Harvard University Department of Psychology faculty History of neuroscience Hungarian biophysicists Hungarian neuroscientists Hungarian Nobel laureates Nobel laureates from Austria-Hungary Hungarian nobility 20th-century Hungarian physicists Hungarian physicists Hungarian emigrants to the United States Members of the Hungarian Academy of Sciences Nobel laureates in Physiology or Medicine University of Hawaiʻi faculty Members of the German National Academy of Sciences Leopoldina ASA Gold Medal recipients Members of the United States National Academy of Sciences University of Bern alumni