''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family

Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae. Species in ...

largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, anterior porefields, complete or nearly complete coxo-

pleural The pleural cavity, pleural space, or interpleural space is the potential space between the pleurae of the pleural sac that surrounds each lung. A small amount of serous pleural fluid is maintained in the pleural cavity to enable lubrication bet ...

sutures at the

prosternum The prothorax is the foremost of the three segments in the thorax of an insect, and bears the first pair of legs. Its principal sclerites (exoskeletal plates) are the pronotum (dorsal), the prosternum (ventral), and the propleuron (lateral) on ea ...

, and incomplete chitin-lines. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as ''

Geophilus electricus ''Geophilus electricus'' is a species of soil centipede in the family Geophilidae found across temperate Europe, especially Britain and Ireland. Originally discovered and named ''Scolopendra electrica'' in 1758, it was later moved to its own genu ...

''. Although centipedes in this genus can have as many as 89 pairs of legs, most species have a much smaller number of leg pairs. For example, two species in this genus include centipedes with only 29 pairs of legs, the lowest number found in the family Geophilidae: '' G. persephones'' (29 pairs in the only specimen, a male) and '' G. richardi'' (29 or 31 pairs in males and 33 pairs in females). This genus also includes other species with notably few legs, for example, ''G. hadesi'' (33 pairs in both males and females) and ''G. piae'' (35 pairs in males and 37 pairs in females).

Species

The genus contains the following species: *'' Geophilus admarinus'' Chamberlin, 1952 *''

Geophilus aenariensis Geophilus aenariensis is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Di ...

'' Verhoeff, 1942 *''

Geophilus aetnensis ''Geophilus aetnensis'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, ...

'' Verhoeff, 1928 *'' Geophilus alaskanus'' Cook, 1904 *''

Geophilus algarum ''Geophilus algarum'' is a species of centipede in the family Geophilidae found in the littoral zone on the French Atlantic and Channel Coasts. It has one subspecies, ''G. algarum var. decipiens'', which can be identified by lack of a dorsal coxa ...

'' Brölemann, 1909 *'' Geophilus alzonis'' Attems, 1952 *''

Geophilus ampyx ''Geophilus ampyx'' is a species of soil centipede in the family Geophilidae found in North America, especially South Carolina. It grows up to 52 millimeters in length, though it averages 30–40, has 49–53 leg pairs in males and 51–55 in fem ...

'' Crabill, 1954 *''

Geophilus angustatus ''Geophilus angustatus'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontida ...

'' Eschscholtz, 1823 *''

Geophilus anonyx ''Geophilus anonyx'' is a species of soil centipede in the family Geophilidae found in Oregon. It was originally placed in the genus Brachygeophilus based on the lack of sternal pores, a character shared with the type species ''B. truncorum'' (n ...

'' Chamberlin, 1941 *''

Geophilus arenarius ''Geophilus arenarius'' is a species of soil centipede in the family Geophilidae found in northwest Africa, specifically near Annaba, Algeria. It's frequently misidentified with '' G. electricus'', and as part of the carpophagus species-complex ...

'' Meinert, 1870 *''

Geophilus atopodon ''Geophilus atopodon'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, ...

'' Chamberlin, 1903 *''

Geophilus aztecus ''Geophilus aztecus'' is a species of soil centipede in the family Geophilidae found in Mexico and Guatemala. The original description of this species is based on a dark red specimen measuring 44 mm in length with antennae moniliform at the apex. ...

'' Humbert and Saussure, 1869 *''

Geophilus becki ''Geophilus becki'' is a species of soil centipede in the family Geophilidae found in Cabrillo Beach, California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Co ...

'' Chamberlin, 1951 *''

Geophilus bipartitus ''Geophilus bipartitus'' is a species of soil centipede in the family Geophilidae found in Japan. It grows up to 15 millimeters in length; the males have about 35 leg pairs, the females 39. It lives in Japanese white birch ''Betula platyphyl ...

'' Takakuwa, 1937 *''

Geophilus bluncki ''Geophilus bluncki'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, D ...

'' Verhoeff, 1928 *''

Geophilus bobolianus ''Geophilus bobolianus'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, ...

'' Verhoeff, 1928 *'' Geophilus bosniensis'' Verhoeff, 1895 *'' Geophilus brevicornis'' Wood, 1862 *'' Geophilus brunneus'' McNeill, 1887 *''

Geophilus carpophagus ''Geophilus carpophagus'' is a species of soil centipede in the family Geophilidae, widely considered to be a type species of the genus ''Geophilus'' (along with '' G. electricus''). It grows up to 60 millimeters in length, with an orange/tan bod ...

'' Leach, 1814 *'' Geophilus caucasicus'' Sseliwanoff, 1884 *'' Geophilus cayugae'' Chamberlin, 1904 *''

Geophilus chalandei ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Brölemann, 1909 *'' Geophilus challangeri'' Pocock, 1891 *'' Geophilus claremontus'' Chamberlin, 1909 *''

Geophilus compactus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Attems, 1934 *'' Geophilus crenulatus'' Silvestri, 1936 *'' Geophilus delotus'' Chamberlin, 1941 *'' Geophilus dentatus'' Takakuwa, 1936 *''

Geophilus duponti ''Geophilus duponti'' is a species of centipede in the Geophilidae family. It is endemic to Australia, and was first described in 1897 by Italian entomologist Filippo Silvestri. Description The original description is based on a female speci ...

'' Silvestri, 1897 *''

Geophilus easoni ''Geophilus easoni'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dig ...

'' Arthur et al. 2001 *'' Geophilus elazigus'' Chamberlin, 1952 *''

'' Linne,1758 *'' Geophilus embius'' Chamberlin, 1912 *'' Geophilus erzurumensis'' Chamberlin, 1952 *'' Geophilus eudontus'' Chamberlin, 1952 *''

Geophilus flavus ''Geophilus flavus'' is a terrestrial, soil-dwelling, species of centipede in the Geophilidae family. ''G. flavus'' occurs in a range of habitats across central Europe, North America, Australia and other tropical regions. Geophilomorph centipede ...

'' De Geer, 1778 *'' Geophilus fossularum'' Verhoeff, 1943 *'' Geophilus fossuliferus'' Karsch, 1884 *'' Geophilus foveatus'' McNeill, 1887 *'' Geophilus frigidanus'' Verhoeff, 1928 *'' Geophilus fruitanus'' Chamberlin, 1928 *''

Geophilus fucorum ''Geophilus fucorum'' is a halophilic species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphil ...

'' Brölemann, 1900 *''

Geophilus gavoyi ''Geophilus gavoyi'' is a species of soil centipede in the family Geophilidae. It is closely related to both ''Geophilus aetnensis ''Geophilus aetnensis'' is a species of soil centipede in the family Geophilidae The Geophilidae are a pol ...

'' Chalande, 1910 *'' Geophilus geronimo'' Chamberlin, 1912 *'' Geophilus gigas'' Attems, 1951 *'' Geophilus glaber'' Bollman, 1887 *''

Geophilus glyptus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Chamberlin, 1902 *''

Geophilus gracilis ''Geophilus gracilis'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, ...

'' Meinert, 1870 *'' Geophilus guanophilus'' Verhoeff, 1939 *''

Geophilus hadesi ''Geophilus hadesi'' is a species of soil centipede in the family Geophilidae, named after Hades, king of the underworld in Greek mythology. This species is found in the caves of the Velebit Mountains of Croatia and characterized by relatively ...

'' Stoev et al. 2015 *'' Geophilus honozus'' Chamberlin, 1952 *'' Geophilus ibericus'' Attems, 1952 *''

Geophilus impressus ''Geophilus impressus'' is a species of soil centipede in the family Geophilidae found all over Europe, and has also been recorded in North Africa (Maghreb, Morocco, and Tunisia). It lives frequently in endogean habitats; in Sardinia it's found ...

'' C.L. Koch, 1847 *'' Geophilus indianae'' McNeill, 1887 *'' Geophilus infossulatus'' Attems, 1901 *''

Geophilus intermissus ''Geophilus intermissus'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontid ...

'' Silvestri, 1935 *'' Geophilus joyeuxi'' Léger and Duboscq, 1903 *'' Geophilus judaicus'' Verhoeff, 1934 *'' Geophilus kobelti'' Attems, 1903 *'' Geophilus koreanus'' Takakuwa, 1936 *''

Geophilus labrofissus ''Geophilus labrofissus'' is a species of soil centipede in the family Geophilidae found on the Balkan Peninsula The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geogra ...

'' Verhoeff, 1938 *''

Geophilus lanius ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Brölemann, 1896 *'' Geophilus leionyx'' Chamberlin, 1938 *'' Geophilus lemuricus'' Verhoeff, 1939 *'' Geophilus longicapillatus'' Verhoeff, 1937 *'' Geophilus madeirae'' Latzel, 1895 *'' Geophilus marginatus'' Lucas, 1849 *''

Geophilus minimus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Verhoeff, 1928 *'' Geophilus monoporus'' Takakuwa, 1934 *''

Geophilus mordax ''Geophilus mordax'', also called the pitted soil centipede, is a species of soil centipede in the family Geophilidae found in North America, especially Indiana, Arkansas, Louisiana, North and South Carolina, and Florida. It grows up to 50 mill ...

'' Meinert, 1886 *'' Geophilus multiporus'' Miyosi, 1955 *'' Geophilus mustiquensis'' Pocock, 1893 *'' Geophilus nanus'' Attems, 1952 *'' Geophilus nasintus'' Chamberlin, 1909 *'' Geophilus naxius'' Verhoeff, 1901 *'' Geophilus nealotus'' Chamberlin, 1902 *'' Geophilus nesiotes'' Attems, 1903 *'' Geophilus nicolanus'' Chamberlin, 1940 *'' Geophilus occidentalis'' Linnaeus, 1758 *'' Geophilus okolonae'' Bollman, 1888 *'' Geophilus oligopus'' Attems, 1895 *'' Geophilus orae'' Verhoeff, 1943 *'' Geophilus oregonus'' Chamberlin, 1941 *'' Geophilus orientalis'' Seliwanoff, 1881 *'' Geophilus orientis'' Chamberlin, 1952 *''

Geophilus osquidatum ''Geophilus osquidatum'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontida ...

'' Brölemann, 1909 *'' Geophilus oweni'' Brölemann, 1887 *''

Geophilus parki ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Auerbach, 1954 *'' Geophilus pauciporus'' Machado, 1952 *'' Geophilus pellekanus'' Attems, 1903 *''

Geophilus persephones ''Geophilus persephones'' is a species of soil centipede in the family Geophilidae discovered in 1999. This species is named after Persephone, the queen of the underworld in Greek mythology, and found in caves in the Gouffre de la Pierre Saint- ...

'' Foddai & Minelli, 1999 *'' Geophilus phanus'' Chamberlin, 1943 *'' Geophilus piae'' Minelli, 1983 *''

Geophilus piedus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Chamberlin, 1930 *'' Geophilus pinivagus'' Verhoeff, 1928 *''

Geophilus polyporus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Takakuwa, 1942 *''

Geophilus procerus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' C. L. Koch, 1878 *'' Geophilus promontorii'' Verhoeff, 1928 *''

Geophilus proximus ''Geophilus proximus'' is a species of soil centipede in the family Geophilidae found in the northern part of the Palearctic and widespread across the entire Baltic basin, though it reaches as far as the Arctic Circle and has been introduced th ...

'' C.L.Koch, 1847 *''

Geophilus punicus ''Geophilus punicus'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, ...

'' Silvestri, 1896 *''

Geophilus pusillifrater ''Geophilus pusillifrater'' is a species of soil centipede in the family Geophilidae found in Bosnia-Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and Pars pro toto#Ge ...

'' Verhoeff, 1898 *'' Geophilus pusillus'' Meinert, 1870 *'' Geophilus pygmaeus'' Latzel, 1880 *'' Geophilus pyrenaicus'' Chalande, 1909 *'' Geophilus readae'' Jones, 2001 *'' Geophilus regnans'' Chamberlin, 1904 *'' Geophilus rex'' Chamberlin, 1912 *'' Geophilus rhomboideus'' Takakuwa, 1937 *'' Geophilus ribauti'' Brölemann, 1908 *''

Geophilus richardi ''Geophilus richardi'' is a species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which a ...

'' Brölemann, 1904 *''

Geophilus ridleyi ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Pocock, 1890 *'' Geophilus rouncei'' Jones, 2001 *'' Geophilus secundus'' Chamberlin, 1912 *'' Geophilus serbicus'' Stojanović, Mitić, Antić, 2019 *'' Geophilus setiger'' Bollman, 1887 *'' Geophilus seurati'' Brölemann, 1924 *'' Geophilus shoshoneus'' Chamberlin, 1925 *''

Geophilus sibiricus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Stuxberg, 1876 *'' Geophilus silesiacus'' Haase, 1881 *'' Geophilus simoporus'' Chamberlin, 1952 *''

Geophilus smithi ''Geophilus smithi'' is a species of soil centipede in the family Geophilidae found in Washington, D.C. It grows up to 28 millimeters in length, has 49 leg pairs (in female specimens), 25-30 coxal In vertebrate anatomy, hip (or "coxa"Latin ...

'' Bollman, 1889 *'' Geophilus sounkyoensis'' Takakuwa, 1937 *'' Geophilus strictus'' Latzel, 1880 *''

Geophilus strigosus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' McNeill, 1887 *'' Geophilus studeri'' Rothenbühler, 1899 *'' Geophilus tampophor'' Chamberlin, 1953 *'' Geophilus tenellus'' L. Koch, 1882 *''

Geophilus tenuiculus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' C. L. Koch, 1878 *'' Geophilus terranovae Palmén, 1954 *''

Geophilus transitus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Chamberlin, 1941 *'' Geophilus trichopus'' Muralewicz, 1926 *''

Geophilus truncorum ''Geophilus truncorum'' is a species of soil centipede in the family Geophilidae found across Western Europe, though it reaches as far as Poland, Italy, and Morocco. This centipede is relatively small, growing up to 20mm in length, with a ye ...

'' Bergsøe and Meinert, 1866 *''

Geophilus ungviculatus ''Geophilus '' is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with '' Brachygeophilus''. It is a mostly holarctic genus characterized by a claw-shaped ultimate pretarsus, ant ...

'' Daday, 1889 *'' Geophilus varians'' McNeill, 1887 *'' Geophilus venezuelae'' Silvestri, 1897 *'' Geophilus vinciguerrae'' Silvestri, 1895 *''

Geophilus virginiensis ''Geophilus virginiensis'' is a species of soil centipede in the family Geophilidae The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodonti ...

'' Bollman, 1889 *'' Geophilus vittatus'' Raffinesque, 1820 *'' Geophilus winnetui'' Attems, 1947 *'' Geophilus yavapainus'' Chamberlin, 1941

References

{{Taxonbar, from=Q4039347 Centipede genera Geophilomorpha Taxa named by William Elford Leach