Geokichla Citrina
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The orange-headed thrush (''Geokichla citrina'') is a
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
in the
thrush ''The Man from U.N.C.L.E.'' is an American spy fiction television series produced by Metro-Goldwyn-Mayer Television and first broadcast on NBC. The series follows secret agents, played by Robert Vaughn and David McCallum, who work for a secret ...
family. It is common in well-wooded areas of the
Indian Subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
and
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
. Most populations are resident. The species shows a preference for shady damp areas, and like many '' Geokichla'' and '' Zoothera'' thrushes, can be quite secretive. The orange-headed thrush is omnivorous, eating a wide range of insects,
earthworm An earthworm is a terrestrial invertebrate that belongs to the phylum Annelida. They exhibit a tube-within-a-tube body plan; they are externally segmented with corresponding internal segmentation; and they usually have setae on all segments. Th ...
s and fruit. It nests in trees but does not form flocks. The male of this small thrush has uniform grey upperparts, and an orange head and underparts. The females and young birds have browner upper parts.


Taxonomy

This species was first described by John Latham in 1790 as ''Turdus citrinus'', the species name meaning " citrine" and referencing the colour of the head and underparts. It has about 12 subspecies. Rasmussen and
Anderton Anderton may refer to: People *Anderton (surname) *The Anderton baronets of England Places *Anderton with Marbury, Cheshire *Anderton, Cornwall Anderton is a settlement in south-east Cornwall, England, United Kingdom, located at . There are als ...
(2005) suggest that this complex may consist of more than one species. *''G. c. citrina'', the nominate subspecies breeds from northern
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
east along the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
to eastern
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
and possibly in western and northern Burma. It winters further south in
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
,
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
and
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
. *''G. c. cyanota'' is mainly resident in Peninsular
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
south to
Kerala Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South ...
. It has a white throat and face sides, with two black stripes running downwards from below the eyes. The spelling emendation ''cyanota'' is suggested by Rasmussen and Anderton. *''G. c. amadoni'' (not always recognized) found in northeastern part of peninsular
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
(Madhya Pradesh and Orissa) has brighter orange crown and longer wings than ''cyanota''. *''G. c. innotata'' breeds through most of South-East Asia from southern Burma and southwestern China to northwest
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
, central and southern
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist ...
,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand t ...
and southern
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
. It winter further south in southern Burma, and much of the rest of Thailand into Malaysia. It is very similar to the nominate but the male is brighter or deeper orange and lacks white tips to the median coverts; the female is duller on head and underparts, with an olive tinge to the grey of the mantle and back. *''G. c. melli'' breeds in southeastern China, and is partially migratory, regularly wintering in
Hong Kong Hong Kong ( (US) or (UK); , ), officially the Hong Kong Special Administrative Region of the People's Republic of China ( abbr. Hong Kong SAR or HKSAR), is a city and special administrative region of China on the eastern Pearl River Delt ...
. *''G. c. courtoisi'' breeds in eastern-central China; its wintering range is unknown. *''G. c. aurimacula'' breeds in southern Vietnam,
Hainan Hainan (, ; ) is the smallest and southernmost province of the People's Republic of China (PRC), consisting of various islands in the South China Sea. , the largest and most populous island in China,The island of Taiwan, which is slightly l ...
and possibly northern Laos. It resembles ''G. c. cyanota'', but with a less defined head pattern. The face and neck-sides are whitish but flecked with orange or brownish and with weaker face stripes. The orange breast and flanks become paler orange on the belly and lower flanks. *''G. c. andamensis'' is resident in the
Andaman Islands The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between th ...
. *''G. c. albogularis'' is resident in the
Nicobar Islands The Nicobar Islands are an archipelagic island chain in the eastern Indian Ocean. They are located in Southeast Asia, northwest of Aceh on Sumatra, and separated from Thailand to the east by the Andaman Sea. Located southeast of the Indian s ...
. *''G. c. gibsonhilli'' breeds from southern Burma to southern Thailand, and winters further south at lower levels in Peninsular Thailand, on islands in the
Gulf of Thailand The Gulf of Thailand, also known as the Gulf of Siam, is a shallow inlet in the southwestern South China Sea, bounded between the southwestern shores of the Indochinese Peninsula and the northern half of the Malay Peninsula. It is around in l ...
, and into
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
. It is similar to the nominate subspecies, but averages slightly brighter or deeper orange on head and upperparts and also has a slightly longer, heavier bill, and white tips to the median coverts. *''G. c. aurata'' is resident in the mountains of northern
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and eas ...
. *''G. c. rubecula'' is resident in Western Java. *''G. c. orientis'' is resident in Eastern Java and
Bali Bali () is a province of Indonesia and the westernmost of the Lesser Sunda Islands. East of Java and west of Lombok, the province includes the island of Bali and a few smaller neighbouring islands, notably Nusa Penida, Nusa Lembongan, and Nu ...
and intergrades with ''G. c. rubecula'' in the west of its range. The separation of this form from the western Javan subspecies has been questioned.


Measurements

The following table summarises selected physical measurements for those subspecies for which data is available.


Distribution and habitat

The orange-headed thrush breeds in much of the
Indian Subcontinent The Indian subcontinent is a list of the physiographic regions of the world, physiographical region in United Nations geoscheme for Asia#Southern Asia, Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian O ...
, including
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the eighth-most populous country in the world, with a population exceeding 165 million people in an area of . Bangladesh is among the mos ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
and
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
, and through
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
to
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's List ...
and southern China. Its habitat is moist broadleaved evergreen woodlands, with a medium-density undergrowth of bushes and
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s, but it also utilises
bamboo Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, bu ...
forests for secondary growth. ''G. c. cyanota'' also occurs in large gardens and orchards. This species is often found in damp areas, near streams or in shady ravines. It occurs between 250–1830 metres (825– 6040 ft) in the Himalayas and up to about 1500 metres (5000 ft) in Malaysia, Thailand and Java. ''G. c. aurata'' is resident between 1000–1630 metres (3300–5400 ft) on Mt Kinabalu and Mt Trus Madi, northern Borneo. Some of the subspecies are completely or partially migratory; their wintering habitat is similar to the breeding forests, but more likely to be at lower altitudes.


Description

The orange-headed thrush is 205–235 milliimetres (8.1–9.25 in) long and weighs 47–60 grammes (1.7–2.1 oz). The adult male of the nominate subspecies of this small thrush has an entirely orange head and underparts, uniformly grey upperparts and wings, and white median and undertail coverts. It has a slate-coloured bill and the legs and feet have brown fronts and pink or yellowish rears. The female resembles the male but has browner or more olive upperparts and warm brown wings, but some old females are almost identical to the male. The juvenile is dull brown with buff streaks on its back, and a rufous tone to the head and face; it has grey wings. The bill is brownish horn, and the legs and feet are brown. This species' orange and grey plumage is very distinctive, and it is unlikely to be confused with any other species. Differences between the subspecies, as described above, can be quite striking, as with the strong head pattern on ''G. c. cyanota'', but may be less obvious variations in plumage tone, or whether there is white on the folded wing. As with many other thrushes, all forms of this species shows a distinctive underwing pattern, with a strong white band.


Voice

Calls of the orange-headed thrush include a soft ' or ', a screeching ', and a thin ' or ' given in flight. However, this bird is generally silent especially in winter. The song is a loud clear series of variably sweet lilting musical notes, recalling the quality of the
common blackbird The common blackbird (''Turdus merula'') is a species of true thrush. It is also called the Eurasian blackbird (especially in North America, to distinguish it from the unrelated New World blackbirds), or simply the blackbird where this does not ...
, but with the more repetitive structure of the
song thrush The song thrush (''Turdus philomelos'') is a Thrush (bird), thrush that breeds across the West Palearctic. It has brown upper-parts and black-spotted cream or buff underparts and has three recognised subspecies. Its distinctive Birdsong, song, ...
. It also includes imitations of other birds like
bulbul The bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropical As ...
s,
babbler Babbler may refer to: * Old World babbler, a large family of mostly Old World passerine birds * Australo-Papuan babbler, passerine birds endemic to Australia-New Guinea * Babbler (software) Babbler is a French software company founded in Septem ...
s and
common tailorbird The common tailorbird (''Orthotomus sutorius'') is a songbird found across tropical Asia. Popular for its nest made of leaves "sewn" together and immortalized by Rudyard Kipling as ''Darzee'' in his ''Jungle Book'', it is a common resident in ur ...
. It sings from a perch in a leafy tree, mostly early morning and late afternoon.


Behaviour

The orange-headed thrush is a shy, secretive bird usually occurring alone or in pairs, but is comparatively more easily seen than many other ''Zoothera'' thrushes, and several birds may congregate outside the breeding season at a good food source. It has a swift, silent flight, but when disturbed will often sit motionless until the threat has passed.


Breeding

The nest, built by both sexes, is a wide but shallow cup of twigs, bracken and rootlets lined with softer plant material like leaves, moss and
conifer Conifers are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single ...
needles. It is constructed at a height of up to 4.5 metres (15 ft) in a small tree or bush, with
mango A mango is an edible stone fruit produced by the tropical tree ''Mangifera indica''. It is believed to have originated in the region between northwestern Myanmar, Bangladesh, and northeastern India. ''M. indica'' has been cultivated in South a ...
trees and
coffee Coffee is a drink prepared from roasted coffee beans. Darkly colored, bitter, and slightly acidic, coffee has a stimulant, stimulating effect on humans, primarily due to its caffeine content. It is the most popular hot drink in the world. S ...
bushes being preferred. Three or four, occasionally five, eggs are laid; they are cream or tinted with pale blue, grey or green, and have pale lilac blotches and reddish brown spots. They are incubated for 13–14 days to hatching, with another 12 days until the young birds leave the nest. This species is a host of the
pied cuckoo The Jacobin cuckoo (''Clamator jacobinus''), also pied cuckoo or pied crested cuckoo, is a member of the cuckoo order of birds that is found in Africa and Asia. It is partially migratory and in India, it has been considered a harbinger of the mons ...
, ''Clamator jacobinus'', a
brood parasite Brood parasites are animals that rely on others to raise their young. The strategy appears among birds, insects and fish. The brood parasite manipulates a host, either of the same or of another species, to raise its young as if it were its own ...
which lay a single egg in the nest. Unlike the common cuckoo, neither the hen nor the hatched chick evict the host's eggs, but the host's young often die because they cannot compete successfully with the
cuckoo Cuckoos are birds in the Cuculidae family, the sole taxon in the order Cuculiformes . The cuckoo family includes the common or European cuckoo, roadrunners, koels, malkohas, couas, coucals and anis. The coucals and anis are sometimes separ ...
for food. The
chestnut-winged cuckoo The chestnut-winged cuckoo or red-winged crested cuckoo (''Clamator coromandus'') is a cuckoo found in Southeast Asia and parts of South Asia. It has dark glossy upperparts, a black head with long crest chestnut wings, a long graduated glossy bla ...
, ''Clamator coromandus'', and, very rarely, the common cuckoo, ''Cuculus canorus'' have also been claimed as parasites on this species.


Feeding

The orange-headed thrush feeds on the ground in dense undergrowth or other thick cover. It is most active at dawn and dusk, probing the leaf litter for insects and their larvae, spiders, other
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s and fruit. In Malaysia, wintering birds regularly feed on figs.


Status

The orange-headed thrush has an extensive range, estimated at . The population size has not been quantified, but it is believed to be large due to its extensive range; it also reported as being locally common. The species is not believed to approach the thresholds for the global population decline criterion of the IUCN Red List (i.e., declining more than 30% in ten years or three generations), and is therefore evaluated as
Least Concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
. It is very popular as cage-bird on Java, and numbers have severely declined in recent years owing to trapping for aviculture. Against the trend in Southeast Asia where loss or fragmentation of woodland poses a threat to forest birds, the orange-headed thrush has colonized Hong Kong, where it was first recorded in 1956, thanks to forest maturation.


Gallery

File:Geokichla citrina, orange-headed thrush - Khao Yai National Park.webm, Geokichla citrina, orange-headed thrush -
Khao Yai National Park Khao Yai National Park ( th, อุทยานแห่งชาติเขาใหญ่, , ) is a national park in Thailand. Established in 1962 as Thailand's first national park, it is the third largest national park in Thailand. Descri ...
,
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
Image:Geokichla citrina, orange-headed thrush.jpg, Geokichla citrina, orange-headed thrush -
Khao Yai National Park Khao Yai National Park ( th, อุทยานแห่งชาติเขาใหญ่, , ) is a national park in Thailand. Established in 1962 as Thailand's first national park, it is the third largest national park in Thailand. Descri ...
,
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is bo ...
File:Orange headed thrush (Z. c. cyanotus).jpg, An orange-headed thrush (ssp. cyanota) at Dandeli Wildlife Sanctuary,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
File:Orange-headed thrush (Geokichla citrina cyanota).jpg, Orange-headed Thrush (Geokichla citrina cyanota) at Bhadra Wildlife Sanctuary,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...


References


External links


Videos on the Internet Bird Collection
{{Taxonbar, from=Q26833749, from2=Q599334
orange-headed thrush The orange-headed thrush (''Geokichla citrina'') is a bird in the thrush family. It is common in well-wooded areas of the Indian Subcontinent and Southeast Asia. Most populations are resident. The species shows a preference for shady damp areas ...
Birds of China Birds of Hainan Birds of India Birds of the Himalayas Birds of Java Birds of East Malaysia Birds of Southeast Asia
orange-headed thrush The orange-headed thrush (''Geokichla citrina'') is a bird in the thrush family. It is common in well-wooded areas of the Indian Subcontinent and Southeast Asia. Most populations are resident. The species shows a preference for shady damp areas ...
orange-headed thrush The orange-headed thrush (''Geokichla citrina'') is a bird in the thrush family. It is common in well-wooded areas of the Indian Subcontinent and Southeast Asia. Most populations are resident. The species shows a preference for shady damp areas ...
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