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''Gasosaurus'' () is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
tetanuran Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans (including birds). Tetanurans ar ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
that lived approximately 171.6 to 161.2 
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago) ...
during the middle of the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
. The name "''Gasosaurus''" is derived from the English "''
gasoline Gasoline (; ) or petrol (; ) (see ) is a transparent, petroleum-derived flammable liquid that is used primarily as a fuel in most spark-ignited internal combustion engines (also known as petrol engines). It consists mostly of organic co ...
''" and the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
() ("lizard / generic reptile"). Only one species is currently recognised, ''G. constructus'', from which the specific name honours the
gasoline Gasoline (; ) or petrol (; ) (see ) is a transparent, petroleum-derived flammable liquid that is used primarily as a fuel in most spark-ignited internal combustion engines (also known as petrol engines). It consists mostly of organic co ...
company that found the Dashanpu fossil quarry in
Sichuan Sichuan (; zh, c=, labels=no, ; zh, p=Sìchuān; alternatively romanized as Szechuan or Szechwan; formerly also referred to as "West China" or "Western China" by Protestant missions) is a province in Southwest China occupying most of the ...
Province,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, now named as the
Lower Shaximiao Formation The Shaximiao Formation () is a Middle Jurassic, Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in ...
.


Discovery and naming

The first and to date only
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s, albeit
postcranial Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated s ...
(missing the
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
), were recovered in 1985 during the construction of a gas facility, which explains the dinosaur's unusual name. It consists of laterally compressed teeth, 4 cervical, 7 dorsal, 5 sacral, and 7 caudal vertebrae, both humeri, and highly abraded and reconstructed pelvic and hind limb material that includes the left ilium, left ischium, left pubis, left femur, left tibia and left fibula. The fossils were defined as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
''Gasosaurus constructus'' by the
paleontologist Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
s
Dong Zhiming Dong Zhiming (Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; born January 1937) is a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) in Beijing. He began working at the ...
and Tang Zilu.


Description

''Gasosaurus'' is a carnivorous
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
with strong legs but short arms. It measured between in length, with a weight of around . However, some estimates put its weight as high as , as very little is known about it. Many potentially informative features of the holotype skeleton (IVPP V7264) are difficult to assess based on published descriptions and images. The taxon represents a tetanuran based on the presence of a pubic peduncle of the ilium that is substantially larger than the ischial peduncle. Examination of casts reveals that the lesser trochanter does not rise above the level of the femoral head, and instead, the proximal portion of the femoral head is broken and the lesser trochanter reaches approximately midlevel of the head as in non-coelurosaurian tetanurans. Furthermore, it is recognized by an amphiplatyan cervical centra with incipient weak ventral keel, dorsals with amphiplatyan centra, low neural arches and plate-like neural spines, dorsal neural spines lacking expanded bulks at tips, five sacral centra and arches firmly fused while neural spines not fused, presence of a humeral foramen, low ilium with a less developed anterior process, and distal ends of pubis and ischium expanded but lacking foot-like processes.


Classification

Traditionally thought to be a
megalosauroid Megalosauroidea (meaning 'great/big lizard forms') is a Taxonomic rank, superfamily (or clade) of Tetanurae, tetanuran theropod dinosaurs that lived from the Middle Jurassic to the Late Cretaceous period. The group is defined as ''Megalosaurus, M ...
, Holtz (2000) found it to be a basal
coelurosauria Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, t ...
n, although later Holtz ''et al.'' (2004) suggested it was a basal
carnosaur Carnosauria is an extinct large group of predatory dinosaurs that lived during the Jurassic and Cretaceous periods. Starting from the 1990s, scientists have discovered some very large carnosaurs in the carcharodontosaurid family, such as ''Gig ...
(possibly a
metriacanthosaurid Metriacanthosauridae is an extinct family (biology), family of Allosauroidea, allosauroid theropod dinosaurs that lived from the Middle Jurassic to the Early Cretaceous. When broken down into its Greek roots, it means "moderately-spined lizards". ...
) on the basis of data from undescribed specimens.Holtz TR Jr, Molnar RE, Currie PJ. 2004. Basal Tetanurae. In: Weishampel DB, Dodson P, Osmólska H, eds. The dinosauria, 2nd edn. Berkeley, CA: University of California Press, 71–110. It may in fact be the most basal coelurosaurian yet known, or may even be close to the
common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. All living beings are in fact descendants of a unique ancestor commonly referred to as the last universal comm ...
of the two groups; in any case, it represents one of the oldest definitive
tetanuran Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans (including birds). Tetanurans ar ...
theropods. Some paleontologists have speculated that ''Gasosaurus'' and ''
Kaijiangosaurus ''Kaijiangosaurus'' (meaning "Kaijiang lizard") is a genus of carnivorous tetanuran theropod dinosaur from the Middle Jurassic of China. In 1984 He Xinlu named and described the type species ''Kaijiangosaurus lini''. The generic name refers to ...
'' may be one and the same species. Because of the fragmentary nature of the known ''Gasosaurus'' fossils, it has an uncertain position within
Tetanurae Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans (including birds). Tetanurans ar ...
, and probably lays outside
Coelurosauria Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs. Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, t ...
. A detailed restudy of the holotype is underway that may change its classification.


Paleoecology

''Gasosarurus'' lived during the mid-
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
period (
Bathonian In the geologic timescale the Bathonian is an age and stage of the Middle Jurassic. It lasted from approximately 168.3 Ma to around 166.1 Ma (million years ago). The Bathonian Age succeeds the Bajocian Age and precedes the Callovian Age. Strat ...
and/ or
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 166.1 ± 4.0 Ma (million years ago) and 163.5 ± 4.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
stages), around 164 million years ago. It coexisted with other mid-range theropods such as '' Chuandongocoelurus'' and ''
Kaijiangosaurus ''Kaijiangosaurus'' (meaning "Kaijiang lizard") is a genus of carnivorous tetanuran theropod dinosaur from the Middle Jurassic of China. In 1984 He Xinlu named and described the type species ''Kaijiangosaurus lini''. The generic name refers to ...
'', two other dinosaurs found from the
Lower Shaximiao Formation The Shaximiao Formation () is a Middle Jurassic, Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in ...
. These three taxa pertain to medium to large bodied theropods and are known from associated prostcranial skeletons. China, boasts the highest taxic diversity of Middle Jurassic theropods of all body sizes from anywhere in the world. Most Middle
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
theropods from china are 'medium-sized', as are the majority of Middle
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The J ...
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
s globally. These similar dinosaurs show an intriguing combination of derived tetanuran synapomorphies and primitive features shared with non-tetanurans, which suggest they occupy a basal position within
Tetanurae Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, tyrannosauroids, ornithomimosaurs, compsognathids and maniraptorans (including birds). Tetanurans ar ...
. Understanding the anatomy of Chinese Jurassic taxa may reveal further details of primitive tetanuran anatomy and help resolve the early evolution of this successful theropod clade. Future exploration of the Chinese Middle Jurassic theropod record may bring new insights into the diversification of Tetanurae, the origin of larger body sizes among more derived theropods, and Middle Jurassic paleobiogeography.


References


External links

{{Taxonbar, from=Q131771 Carnosaurs Middle Jurassic dinosaurs of Asia Jurassic China Fossils of China Paleontology in Sichuan Fossil taxa described in 1985 Taxa named by Dong Zhiming