Galago (Otolicnus Galago)
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Galagos , also known as bush babies, or ''nagapies'' (meaning "night monkeys" in
Afrikaans Afrikaans (, ) is a West Germanic language that evolved in the Dutch Cape Colony from the Dutch vernacular of Holland proper (i.e., the Hollandic dialect) used by Dutch, French, and German settlers and their enslaved people. Afrikaans gra ...
), are small nocturnal
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians ( monkeys and apes, the latter including ...
s native to continental, sub-Sahara
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, and make up the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Galagidae (also sometimes called Galagonidae). They are considered a sister group of the
Lorisidae Lorisidae (or sometimes Loridae) is a family of strepsirrhine primates. The lorisids are all slim arboreal animals and comprise the lorises, pottos and angwantibos. Lorisids live in tropical, central Africa as well as in south and southeast As ...
. According to some accounts, the name "bush baby" comes from either the animal's cries or its appearance. The Ghanaian name ''aposor'' is given to them because of their firm grip on branches. In both variety and abundance, the bush babies are the most successful
strepsirrhine Strepsirrhini or Strepsirhini (; ) is a suborder of primates that includes the lemuriform primates, which consist of the lemurs of Madagascar, galagos ("bushbabies") and pottos from Africa, and the lorises from India and southeast Asia. Colle ...
primates in Africa, according to the
African Wildlife Foundation The African Wildlife Foundation (AWF) is the leading international conservation organization focused exclusively on Africa's wildlife and wild lands. AWF's programs and conservation strategies are designed to protect the wildlife and wild lands of ...
.


Taxonomic classification and phylogeny

Galagos are currently grouped into six genera. ''
Euoticus The needle-clawed bushbabies are the two species in the genus ''Euoticus'', which is in the family Galagidae. Galagidae is sometimes included as a subfamily within the Lorisidae (or Loridae). * Genus ''Euoticus'' ** Southern needle-clawed bush ...
'' is a basal
sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
to all the other galagids. The 'dwarf' galagids recently grouped under the genus ''
Galagoides The western dwarf galagos are a group of three species of strepsirrhine primates, native to western and central Africa. They are classified in the genus ''Galagoides'' of the family Galagidae. The eastern dwarf galagos (''P. cocos, P. granti, P. ...
'' have been found, based on genetic data, and supported by analysis of vocalisations and morphology, to actually consist of two clades, which are not sister taxa, in eastern and western/central Africa (separated by the
rift valley A rift valley is a linear shaped lowland between several highlands or mountain ranges created by the action of a geologic rift. Rifts are formed as a result of the pulling apart of the lithosphere due to extensional tectonics. The linear d ...
). The latter are basal to all the other non-''Euoticus'' galagids. The former group is sister to ''Galago'' and has been elevated to full genus status as ''
Paragalago The eastern dwarf galagos are a group of seven species of strepsirrhine primates of the family Galagidae, native to East Africa. They were formerly classified in the genus ''Galagoides'' but have been moved to their own genus, ''Paragalago'', ...
''. The genera ''
Otolemur The greater galagos or thick-tailed bushbabies are three species of strepsirrhine primates. They are classified in the genus ''Otolemur'' in the family Galagidae. Historical classification and species discovery The diversity of galago species h ...
'' and ''
Sciurocheirus The squirrel galagos are a group of four species of strepsirrhine primates. They are classified in the genus ''Sciurocheirus'' of the family Galagidae. Originally a single species was described, ''Galago alleni'', by Waterhouse (1838), and the ...
'' are also sisters. Family Galagidae - galagos, or bushbabies * Genus ''
Euoticus The needle-clawed bushbabies are the two species in the genus ''Euoticus'', which is in the family Galagidae. Galagidae is sometimes included as a subfamily within the Lorisidae (or Loridae). * Genus ''Euoticus'' ** Southern needle-clawed bush ...
'', needle-clawed bushbabies **
Southern needle-clawed bushbaby The southern needle-clawed bushbaby (''Euoticus elegantulus'') is a species of strepsirrhine primate in the family Galagidae. Found in Cameroon, Central African Republic, Republic of the Congo, and possibly Democratic Republic of the Congo, its n ...
, ''E. elegantulus'' **
Northern needle-clawed bushbaby The northern needle-clawed bushbaby (''Euoticus pallidus'') is a species of strepsirrhine primate in the family Galagidae. It is found in the coastal region of Cameroon and Nigeria, and on the island of Bioko, Equatorial Guinea in lower-elevation ...
, ''E. pallidus'' *Genus ''
Galago Galagos , also known as bush babies, or ''nagapies'' (meaning "night monkeys" in Afrikaans), are small nocturnal primates native to continental, sub-Sahara Africa, and make up the family Galagidae (also sometimes called Galagonidae). They are ...
'', lesser galagos, or lesser bushbabies **''Galago senegalensis'' group ***
Somali bushbaby The Somali bushbaby (''Galago gallarum''), or the Somali lesser galago, as it is also known, is a species of nocturnal, arboreal primate in the family Galagidae. It is found in Ethiopia, Kenya, and Somalia. It is threatened by habitat loss. Dis ...
, ''G. gallarum'' ***
Mohol bushbaby The Mohol bushbaby (''Galago moholi'') is a species of primate in the family Galagidae which is native to mesic woodlands of the southern Afrotropics. It is physically very similar to the Senegal bushbaby, and was formerly considered to be its s ...
, ''G. moholi'' ***
Senegal bushbaby The Senegal bushbaby (''Galago senegalensis''), also known as the Senegal galago, the lesser galago or the lesser bush baby, is a small, nocturnal primate, a member of the galago family Galagidae. The name "bush baby" may come either from the a ...
, ''G. senegalensis'' **''Galago matschiei'' group ***
Dusky bushbaby The dusky bushbaby (''Galago matschiei'') is a species of primate in the family Galagidae. It is also known as Matschie's galago, in honour of the German zoologist Paul Matschie, curator of mammals at the Museum für Naturkunde in Berlin. Native ...
, ''G. matschiei'' *Genus ''
Galagoides The western dwarf galagos are a group of three species of strepsirrhine primates, native to western and central Africa. They are classified in the genus ''Galagoides'' of the family Galagidae. The eastern dwarf galagos (''P. cocos, P. granti, P. ...
'', western dwarf galagos **
Prince Demidoff's bushbaby Prince Demidoff's bushbaby (''Galagoides demidovii''), also known as Prince Demidoff's galago, is a species of primate in the family Galagidae. It is native to parts of tropical West and Central Africa. Description Prince Demidoff's bushbaby gro ...
, ''Gs. demidovii'' ** Angolan dwarf galago, ''Gs. kumbirensis'' **
Thomas's bushbaby Thomas's bushbaby (''Galagoides thomasi'') is a species of primate in the family Galagidae. It is found in Angola, Burundi, Cameroon, Democratic Republic of the Congo, Equatorial Guinea, Gabon, Kenya, Nigeria, Rwanda, Tanzania, Uganda, and Zam ...
, ''Gs. thomasi'' *Genus †''
Laetolia ''Laetolia'' is an extinct genus of galagid primates from the Pliocene of Tanzania. It contains one species, ''L. sadimanensis'', which is known from several dentary fragments discovered in the Upper Laetolil Beds of Laetoli. It was originally d ...
'' **†'' Laetolia sadimanensis'' *Genus ''
Otolemur The greater galagos or thick-tailed bushbabies are three species of strepsirrhine primates. They are classified in the genus ''Otolemur'' in the family Galagidae. Historical classification and species discovery The diversity of galago species h ...
'', greater galagos, or thick-tailed bushbabies **
Brown greater galago The brown greater galago (''Otolemur crassicaudatus''), also known as the large-eared greater galago or thick-tailed galago, is a nocturnal primate, the largest in the family of galagos. As opposed to smaller galago species it would climb, walk o ...
, ''O. crassicaudatus'' **
Northern greater galago The northern greater galago (''Otolemur garnettii''), also known as Garnett's greater galago, Garnett's galago, or the small-eared greater galago, is a nocturnal, arboreal primate endemic to Africa. Subspecies Four subspecies of ''Otolemur garne ...
, ''O. garnettii'' ** Silvery greater galago, ''O. monteiri'' *Genus ''
Paragalago The eastern dwarf galagos are a group of seven species of strepsirrhine primates of the family Galagidae, native to East Africa. They were formerly classified in the genus ''Galagoides'' but have been moved to their own genus, ''Paragalago'', ...
'', eastern dwarf galagos **''Paragalago zanzibaricus'' group ***
Kenya coast galago The Kenya coast galago (''Paragalago cocos'') is a species of primate in the family Galagidae. It is found across Kenya and Tanzania, particularly around the northern coastal area of each of the countries, and has been located living from sea ...
, ''P. cocos'' ***
Grant's bushbaby Grant's bushbaby (''Paragalago granti''), also known as Grant's lesser bushbaby or the Mozambique lesser bushbaby, is a species of primate in the family Galagidae. It is found in Malawi, Mozambique, Tanzania, and Zimbabwe. Its natural habitat is ...
, ''P. granti'' ***
Malawi bushbaby The Malawi bushbaby (''Paragalago nyasae'') is a species of primate in the family Galagidae. It lives in southern Malawi and the neighboring region in Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ...
, ''P. nyasae'' ***
Zanzibar bushbaby The Zanzibar bushbaby, Matundu dwarf galago, Udzungwa bushbaby, or Zanzibar galago (''Paragalago zanzibaricus'') is a primate of the family Galagidae. An adult typically weighs , its head-body length is and its tail is between long. Like other ...
, ''P. zanzibaricus'' **''Paragalago orinus'' group ***
Uluguru bushbaby The Uluguru bushbaby (''Paragalago orinus''), also known as the mountain dwarf galago or the Amani dwarf galago, is a species of primate in the family Galagidae. Like all galagos, it is a strepsirrhine primate. It is endemic to the Eastern Arc Mo ...
, ''P. orinus'' *** Rondo bushbaby, ''P. rondoensis'' **Uncertain ***
Rungwe dwarf galago The Rungwe dwarf galago is a newly identified species of eastern dwarf galago. Specimens were first collected in the 1930s, but were identified as different species. A formal description of the species is presently being made. Description The Ru ...
, ''P.'' sp. nov. *Genus ''
Sciurocheirus The squirrel galagos are a group of four species of strepsirrhine primates. They are classified in the genus ''Sciurocheirus'' of the family Galagidae. Originally a single species was described, ''Galago alleni'', by Waterhouse (1838), and the ...
'', squirrel galagos **
Bioko Allen's bushbaby Bioko Allen's bushbaby (''Sciurocheirus alleni''), also known as the Bioko squirrel galago, is a species of primate in the galago family found in Bioko, Equatorial Guinea. Its natural habitat is subtropical or tropical dry forests. The bushbaby ...
, ''S. alleni'' **
Cross River bushbaby The Cross River bushbaby, also known as the Cross River squirrel galago, (''Sciurocheirus cameronensis'') is a species of prosimian primate in the family Galagidae which is endemic to a restricted area of West Africa. It is one of four species of ...
, ''S. cameronensis'' **
Gabon bushbaby The Gabon bushbaby (''Sciurocheirus gabonensis'') is a species of primate in the family Galagidae found in Cameroon, Gabon, and the Republic of the Congo. Its head and body length is 8.5 in with a 10-in tail, and it weighs about 10 oz. It lives ...
, ''S. gabonensis'' ** Makandé squirrel galago, ''S. makandensis'' The phylogeny of Galagidae according to Masters ''et al.'', 2017 is as follows:


Characteristics

Galagos have large eyes that give them good night vision in addition to other characteristics, like strong hind limbs, acute
hearing Hearing, or auditory perception, is the ability to perceive sounds In physics, sound is a vibration that propagates as an acoustic wave, through a transmission medium such as a gas, liquid or solid. In human physiology and psycholog ...
, and long tails that help them balance. Their ears are bat-like and allow them to track insects in the dark. They catch insects on the ground or snatch them out of the air. They are fast, agile creatures. As they bound through the thick bushes, they fold their delicate ears back to protect them. They also fold them during rest. They have nails on most of their digits, except for the second toe of the hind foot, which bears a
grooming claw A grooming claw (or toilet claw) is the specialized claw or nail on the foot of certain primates, used for personal grooming. All prosimians have a grooming claw, but the digit that is specialized in this manner varies. Tarsiers have a grooming ...
. Their diet is a mixture of insects and other small animals, fruit, and tree gums. They have pectinate (comb-like)
incisors Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wh ...
called
toothcomb A toothcomb (also tooth comb or dental comb) is a dental structure found in some mammals, comprising a group of front teeth arranged in a manner that facilitates grooming, similar to a hair comb. The toothcomb occurs in lemuriform primates ( ...
s, and the
dental formula Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
: They are active at night. After a
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregna ...
period of 110–133 days, young galagos are born with half-closed eyes and are initially unable to move about independently. After a few (6–8) days, the mother carries the infant in her mouth, and places it on branches while feeding. Females may have singles, twins, or triplets, and may become very aggressive. Each newborn weighs less than half an ounce. For the first three days, the infant is kept in constant contact with the mother. The young are fed by the mother for six weeks and can feed themselves at two months. The young grow rapidly, often causing the mother to walk awkwardly as she transports them. Females maintain a
territory A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
shared with their offspring, while males leave their mothers' territories after puberty. Thus social groups consist of closely related females and their young. Adult males maintain separate territories, which overlap with those of the female social groups; generally, one adult male mates with all the females in an area. Males that have not established such territories sometimes form small bachelor groups. Bush-babies are sometimes kept as pets, although this is not advised because, like many other nonhuman primates, they are a likely sources of diseases that can cross species barriers. Equally, they are very likely to attract attention from customs officials on importation into many countries. Reports from veterinary and zoological sources indicate captive lifetimes of 12.0 to 16.5 years, suggesting a natural lifetime over a decade. Galagos communicate by calling to each other and by marking their paths with urine. By following the scent of urine, they can land on exactly the same branch every time. Each species produces a unique set of loud calls that have different functions. One function is to identify individuals as members of a particular species across distances. Scientists can recognize all known galago species by their 'loud calls'. At the end of the night, group members use a special rallying call and gather to sleep in a nest of leaves, a group of branches, or a hole in a tree.


Jumping

Galagos have remarkable
jumping Jumping or leaping is a form of locomotion or movement in which an organism or non-living (e.g., robotics, robotic) mechanical system propels itself through the air along a ballistic trajectory. Jumping can be distinguished from running, gallo ...
abilities. The highest reliably reported jump for a galago is 2.25 m. According to a study published by the Royal Society, given the body mass of each animal and the fact that the leg muscles amount to about 25% of this, galago's jumping muscles should perform six to nine times better than those of a frog. This is thought to be due to elastic energy storage in tendons of the lower leg, allowing far greater jumps than would otherwise be possible for an animal of their size. In mid-flight, they tuck their arms and legs close to the body; they bring them out at the last second to grab a branch. In a series of leaps, a galago can cover ten yards in mere seconds. The tail, which is longer than the length of the head and body combined, assists the legs in powering the jumps. They may also hop like a kangaroo or simply run or walk on four legs. Such strong, complicated, and coordinated movements are due to the rostral half of the posterior parietal cortex that is linked to the motor, premotor, and visuomotor areas of the frontal cortex.


Behaviour

Generally, the social structure of the galago has components of both social life and solitary life. This can be seen in their play. They swing off branches or climb high and throw things. Social play includes play fights, play grooming, and following-play. During following-play, two galagos jump sporadically and chase each other through the trees. The older galagos in a group prefer to rest alone, while younger ones are in constant contact with one another. This is observed in the ''Galago garnetti'' species. Mothers often leave infants alone for long periods and do not try to stop them from leaving. On the other hand, the offspring tries to stay close to, and initiate social interactions with the mother. Grooming is a very important part of galago daily life. They often groom themselves before, during, and after rest.
Social grooming Social grooming is a behavior in which social animals, including humans, clean or maintain one another's body or appearance. A related term, allogrooming, indicates social grooming between members of the same species. Grooming is a major soci ...
is done more often by males in the group. Females often reject attempts by males to groom them.


Relationship with humans

The bush baby also refers to a myth that is used to scare children to stay indoors at night. Most likely arising from the baby-like cry, the unusual nature evolved into a myth about a powerful animal that can kidnap humans. It is also said that wild bush babies/galagos in Nigeria can never be found dead on plain ground. Rather, they make a nest of sticks, leaves or branches to die in. Endangerment of the species in sub-Saharan Africa has made this claim difficult to verify.


References


External links

{{Taxonbar, from=Q185239 01 . Primates of Africa Taxa named by John Edward Gray Taxa described in 1825