Toothcomb
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Toothcomb
A toothcomb (also tooth comb or dental comb) is a dental structure found in some mammals, comprising a group of front teeth arranged in a manner that facilitates grooming, similar to a hair comb. The toothcomb occurs in lemuriform primates (which includes lemurs and lorisoids), treeshrews, colugos, hyraxes, and some African antelopes. The structures evolved independently in different types of mammals through convergent evolution and varies both in dental composition and structure. In most mammals the comb is formed by a group of teeth with fine spaces between them. The toothcombs in most mammals include incisors only, while in lemuriform primates they include incisors and canine teeth that tilt forward at the front of the lower jaw, followed by a canine-shaped first premolar. The toothcombs of colugos and hyraxes take a different form with the individual incisors being serrated, providing multiple tines per tooth. The toothcomb is usually used for grooming. While licking ...
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Toothcomb - Jones 1918
A toothcomb (also tooth comb or dental comb) is a dental structure found in some mammals, comprising a group of front teeth arranged in a manner that facilitates grooming, similar to a hair comb. The toothcomb occurs in lemuriform primates (which includes lemurs and lorisoids), treeshrews, colugos, hyraxes, and some African antelopes. The structures evolved independently in different types of mammals through convergent evolution and varies both in dental composition and structure. In most mammals the comb is formed by a group of teeth with fine spaces between them. The toothcombs in most mammals include incisors only, while in lemuriform primates they include incisors and canine teeth that tilt forward at the front of the lower jaw, followed by a canine-shaped first premolar. The toothcombs of colugos and hyraxes take a different form with the individual incisors being serrated, providing multiple tines per tooth. The toothcomb is usually used for grooming. While licking ...
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Lemur
Lemurs ( ) (from Latin ''lemures'' – ghosts or spirits) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea (), divided into 8 Family (biology), families and consisting of 15 genera and around 100 existing species. They are endemic to the island of Madagascar. Most existing lemurs are small, have a pointed snout, large eyes, and a long tail. They arboreal, chiefly live in trees and nocturnal, are active at night. Lemurs share resemblance with other primates, but evolved independently from monkeys and apes. Due to Madagascar's highly seasonal climate, Evolution of lemurs, lemur evolution has produced a level of species diversity rivaling that of any other primate group. Until shortly after humans arrived on the island around 2,000 years ago, there were lemurs as large as a male gorilla. Most species have been discovered or promoted to full species status since the 1990s; however, lemur Taxonomy (biology), taxonomic classification is ...
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Sublingua
The sublingua ("under-tongue") is a muscular secondary tongue found below the primary tongue in tarsiers and living Strepsirrhini, strepsirrhine primates, which includes lemurs and Lorisoidea, lorisoids (collectively called "Lemuriformes, lemuriforms"). Although it is most fully developed in these primates, similar structures can be found in some other mammals, such as marsupials, treeshrews, and colugos. This "second tongue" lacks taste buds, and in lemuriforms, it is thought to be used to remove hair and other debris from the toothcomb, a specialized dental structure used to comb the fur during oral grooming. A rigid structure called the plica mediana or lytta runs from the front to the back, down the center of the sublingua to give it support. The plica mediana is usually made of cartilage and attaches the sublingua to the underside of the tongue. In lemuriforms, the sublingua mostly consists of two plicae fimbriatae (singular: plica fimbriata), which run along the sides of the ...
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Evolutionary History Of Lemurs
Lemurs, primates belonging to the suborder Strepsirrhini which branched off from other primates less than 63 million years ago, evolved on the island of Madagascar, for at least 40 million years. They share some traits with the most basal primates, and thus are often confused as being ancestral to modern monkeys, apes, and humans. Instead, they merely resemble ancestral primates. Lemurs are thought to have evolved during the Eocene or earlier, sharing a closest common ancestor with lorises, pottos, and galagos (lorisoids). Fossils from Africa and some tests of nuclear DNA suggest that lemurs made their way to Madagascar between 40 and 52 mya. Other mitochondrial and nuclear DNA sequence comparisons offer an alternative date range of 62 to 65 mya. An ancestral lemur population is thought to have inadvertently rafted to the island on a floating mat of vegetation, although hypotheses for land bridges and island hopping have also been proposed. The timing and numb ...
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Hair Comb
A comb is a tool consisting of a shaft that holds a row of teeth for pulling through the hair to clean, untangle, or style it. Combs have been used since prehistoric times, having been discovered in very refined forms from settlements dating back to 5,000 years ago in Persia. Weaving combs made of whalebone dating to the middle and late Iron Age have been found on archaeological digs in Orkney and Somerset. Description Combs consist of a shaft and teeth that are placed at a perpendicular angle to the shaft. Combs can be made out of a number of materials, most commonly plastic, metal, or wood. In antiquity, horn and whalebone was sometimes used. Combs made from ivory and tortoiseshell were once common but concerns for the animals that produce them have reduced their usage. Wooden combs are largely made of boxwood, cherry wood, or other fine-grained wood. Good quality wooden combs are usually handmade and polished. Combs come in various shapes and sizes depending on what t ...
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Lemuriformes
Lemuriformes is an infraorder of primate that falls under the suborder Strepsirrhini. It includes the lemurs of Madagascar, as well as the galagos and lorisids of Africa and Asia, although a popular alternative taxonomy places the lorisoids in their own infraorder, Lorisiformes. Lemuriform primates are characterized by a toothcomb, a specialized set of teeth in the front, lower part of the mouth mostly used for combing fur during grooming. Evolutionary history Lemuriform origins are unclear and debated. American paleontologist Philip Gingerich proposed that lemuriform primates evolved from one of several genera of European adapids based on similarities between the front lower teeth of adapids and the toothcomb of extant lemuriforms; however, this view is not strongly supported due to a lack of clear transitional fossils. Instead, lemuriforms may be descended from a very early branch of Asian cercamoniines or sivaladapids that immigrated to northern Africa. Until discov ...
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Djebelemur
''Djebelemur'' is an extinct genus of early strepsirrhine primate from the late early or early middle Eocene period from the Chambi locality in Tunisia. Although they probably lacked a toothcomb, a specialized dental structure found in living lemuriforms (lemurs and lorisoids), they are thought to be a related stem group. The one recognized species, ''Djebelemur martinezi'', was very small, approximately . ''Djebelemur'' is one of five genera of djebelemurids, others including '' 'Anchomomys' milleri'', a fossil primate formerly thought to be related to other members of the genus '' Anchomomys'', a group of Eocene European adapiform primates. Following its discovery and description in 1992, ''Djebelemur'' was thought to be either a relative of European adapiforms or an early simian, mostly due to the fragmentary nature of the lower jaw and two isolated upper molars that may not belong with the jaw. By 2006, it was viewed as a stem lemuriform. Etymology The name "''Djebe ...
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Primate
Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians (monkeys and apes, the latter including humans). Primates arose 85–55 million years ago first from small terrestrial mammals, which adapted to living in the trees of tropical forests: many primate characteristics represent adaptations to life in this challenging environment, including large brains, visual acuity, color vision, a shoulder girdle allowing a large degree of movement in the shoulder joint, and dextrous hands. Primates range in size from Madame Berthe's mouse lemur, which weighs , to the eastern gorilla, weighing over . There are 376–524 species of living primates, depending on which classification is used. New primate species continue to be discovered: over 25 species were described in the 2000s, 36 in the 2010s, and three in the 2020s. Primates have large bra ...
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Fork-marked Lemur
Fork-marked lemurs or fork-crowned lemurs are strepsirrhine primates; the four species comprise the genus ''Phaner''. Like all lemurs, they are native to Madagascar, where they are found only in the west, north, and east sides of the island. They are named for the two black stripes which run up from the eyes, converge on the top of the head, and run down the back as a single black stripe. They were originally placed in the genus ''Lemur'' in 1839, later moved between the genera '' Cheirogaleus'' and ''Microcebus'', and given their own genus in 1870 by John Edward Gray. Only one species ('' Phaner furcifer'') was recognized, until three subspecies described in 1991 were promoted to species status in 2001. New species may yet be identified, particularly in northeast Madagascar. Fork-marked lemurs are among the least studied of all lemurs and are some of the largest members of the family Cheirogaleidae, weighing around or more. They are the most phylogenetically distinct of the ch ...
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Colugo
Colugos () are arboreal gliding mammals that are native to Southeast Asia. Their closest evolutionary relatives are primates. There are just two living species of colugos: the Sunda flying lemur (''Galeopterus variegatus'') and the Philippine flying lemur (''Cynocephalus volans''). These two species make up the entire family Cynocephalidae () and order Dermoptera (not to be confused with Dermaptera, an order of insects known as earwigs). Although they are called "flying lemurs", the colugos are not lemurs and do not fly. Instead, they glide as they leap among trees. They are the most capable gliders of all gliding mammals. A fur-covered membrane, called a patagium, connects to the face, paws, and tail. This enables them to glide in the air for distances of up to between trees. They are also known as cobegos. Flying lemurs and regular lemurs (Lemuriformes) form a mirorder together (called primatomorpha). However, notably, the haplorhini are sister to the regular lemurs within t ...
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Lorisoidea
Lorisoidea is a superfamily of nocturnal primates found throughout Africa and Asia. Members include the galagos and the lorisids. As strepsirrhines, lorisoids are related to the lemurs of Madagascar and are sometimes included in the infraorder Lemuriformes, although they are also sometimes placed in their own infraorder, Lorisiformes Gregory, 1915. Classification * Order Primates ** Suborder Strepsirrhini *** Infraorder †Adapiformes *** Infraorder Lemuriformes **** Superfamily Lemuroidea: lemurs **** Superfamily Lorisoidea ***** Family Lorisidae: lorises, pottos, and angwantibos ***** Family Galagidae: galagos ** Suborder Haplorhini Haplorhini (), the haplorhines (Greek for "simple-nosed") or the "dry-nosed" primates, is a suborder of primates containing the tarsiers and the simians (Simiiformes or anthropoids), as sister of the Strepsirrhini ("moist-nosed"). The name is some ...: tarsiers, monkeys, and apes Notes References * . Mammal superfamilies Taxa na ...
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Indriidae
The Indriidae (sometimes incorrectly spelled Indridae) are a family of strepsirrhine primates. They are medium- to large-sized lemurs, with only four teeth in the toothcomb instead of the usual six. Indriids, like all lemurs, live exclusively on the island of Madagascar. Classification The 19 living species in the family are divided into three genera. Family Indriidae *Genus ''Indri'' **Indri, ''Indri indri'' *Genus '' Avahi'', woolly lemurs **Bemaraha woolly lemur, ''Avahi cleesei'' **Eastern woolly lemur, ''Avahi laniger'' **Moore's woolly lemur, ''Avahi mooreorum'' **Western woolly lemur, ''Avahi occidentalis'' **Sambirano woolly lemur, ''Avahi unicolor'' **Peyrieras's woolly lemur, ''Avahi peyrierasi'' **Southern woolly lemur, ''Avahi meridionalis'' **Ramanantsoavana's woolly lemur, ''Avahi ramanantsoavani'' **Betsileo woolly lemur, ''Avahi betsileo'' *Genus ''Propithecus'', sifakas **''Propithecus diadema'' group ***Diademed sifaka, ''Propithecus diadema'' ***Silky sifaka, ...
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