W. Ford Doolittle (born February 21, 1942, in

Urbana, Illinois Urbana ( ) is a city in and the county seat of Champaign County, Illinois, United States. As of the 2020 census, Urbana had a population of 38,336. As of the 2010 United States Census, Urbana is the List of municipalities in Illinois, 38th-most pop ...

) is an

evolutionary Evolution is change in the heredity, heritable Phenotypic trait, characteristics of biological populations over successive generations. These characteristics are the Gene expression, expressions of genes, which are passed on from parent to ...

and

molecular biologist Molecular biology is the branch of biology that seeks to understand the molecular basis of biological activity in and between cells, including biomolecular synthesis, modification, mechanisms, and interactions. The study of chemical and physi ...

. He is a member of the US

National Academy of Sciences The National Academy of Sciences (NAS) is a United States nonprofit, non-governmental organization. NAS is part of the National Academies of Sciences, Engineering, and Medicine, along with the National Academy of Engineering (NAE) and the Nati ...

and a Fellow of the

Royal Society of Canada The Royal Society of Canada (RSC; french: Société royale du Canada, SRC), also known as the Academies of Arts, Humanities and Sciences of Canada (French: ''Académies des arts, des lettres et des sciences du Canada''), is the senior national, bil ...

and the Norwegian Academy of Science and Letters. He is also the winner of the 2013

Herzberg Medal The Gerhard Herzberg Canada Gold Medal for Science and Engineering is awarded by the Natural Sciences and Engineering Research Council (NSERC) of Canada to recognize "research contributions characterized by both excellence and influence." Prior ...

of the

Natural Sciences and Engineering Research Council of Canada The Natural Sciences and Engineering Research Council of Canada (NSERC; french: Conseil de recherches en sciences naturelles et en génie du Canada, CRSNG) is the major federal agency responsible for funding natural sciences and engineering rese ...

and the 2017

Killam Prize The Izaak Walton Killam Memorial Prize was established according to the will of Dorothy J. Killam to honour the memory of her husband Izaak Walton Killam. Five Killam Prizes, each having a value of $100,000, are annually awarded by the Canada Cou ...

. Doolittle has made significant contributions to the study of

cyanobacteria Cyanobacteria (), also known as Cyanophyta, are a phylum of gram-negative bacteria that obtain energy via photosynthesis. The name ''cyanobacteria'' refers to their color (), which similarly forms the basis of cyanobacteria's common name, blu ...

. He found evidence for the

endosymbiont An ''endosymbiont'' or ''endobiont'' is any organism that lives within the body or cells of another organism most often, though not always, in a mutualistic relationship. (The term endosymbiosis is from the Greek: ἔνδον ''endon'' "within" ...

origins of

chloroplasts A chloroplast () is a type of membrane-bound organelle known as a plastid that conducts photosynthesis mostly in plant cell, plant and algae, algal cells. The photosynthetic pigment chlorophyll captures the energy from sunlight, converts it, ...

, and developed a theoretical basis for the initial evolution of

eukaryotes Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bacte ...

. He has shown the importance of

horizontal gene transfer Horizontal gene transfer (HGT) or lateral gene transfer (LGT) is the movement of genetic material between Unicellular organism, unicellular and/or multicellular organisms other than by the ("vertical") transmission of DNA from parent to offsprin ...

prokaryotic A prokaryote () is a Unicellular organism, single-celled organism that lacks a cell nucleus, nucleus and other membrane-bound organelles. The word ''prokaryote'' comes from the Greek language, Greek wikt:πρό#Ancient Greek, πρό (, 'before') a ...

evolution. , he has been

Professor Emeritus ''Emeritus'' (; female: ''emerita'') is an adjective used to designate a retired chair, professor, pastor, bishop, pope, director, president, prime minister, rabbi, emperor, or other person who has been "permitted to retain as an honorary title ...

Dalhousie University Dalhousie University (commonly known as Dal) is a large public research university in Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the fou ...

Halifax, Nova Scotia Halifax is the capital and largest municipality of the Canadian province of Nova Scotia, and the largest municipality in Atlantic Canada. As of the 2021 Census, the municipal population was 439,819, with 348,634 people in its urban area. The ...

. He received his BA in

Biochemical Biochemistry or biological chemistry is the study of chemical processes within and relating to living organisms. A sub-discipline of both chemistry and biology, biochemistry may be divided into three fields: structural biology, enzymology an ...

Sciences from

Harvard University Harvard University is a private Ivy League research university in Cambridge, Massachusetts. Founded in 1636 as Harvard College and named for its first benefactor, the Puritan clergyman John Harvard, it is the oldest institution of higher le ...

in 1963 and his PhD from

Stanford University Stanford University, officially Leland Stanford Junior University, is a private research university in Stanford, California. The campus occupies , among the largest in the United States, and enrolls over 17,000 students. Stanford is consider ...

in 1967, under

Charles Yanofsky Charles Yanofsky (April 17, 1925 – March 16, 2018) was an American geneticist on the faculty of Stanford University who contributed to the establishment of the one gene-one enzyme hypothesis and discovered attenuation, a riboswitch mechanis ...

. He went on to do postdoctoral fellowships with

Sol Spiegelman Sol Spiegelman (December 14, 1914 – January 21, 1983) was an American molecular biologist. He developed the technique of nucleic acid hybridization, which helped to lay the groundwork for advances in recombinant DNA technology. Early life and ...

and

Norman R. Pace Norman Richard Pace Jr. (born 1942) is an American biochemist, and is Distinguished Professor Emeritus of Molecular, Cellular and Developmental Biology at the University of Colorado. He is principal investigator at the Pace lab. Early life and ...

. In 1981, Doolittle received some level of notoriety for his article in ''The CoEvolution Quarterly'' entitled "Is Nature Really Motherly?". This was a sharp rebuttal of J. E. Lovelock's formulation of the

Gaia Theory The Gaia hypothesis (), also known as the Gaia theory, Gaia paradigm, or the Gaia principle, proposes that living organisms interact with their inorganic surroundings on Earth to form a synergistic and self-regulating, complex system that helps ...

. Doolittle's article is often cited by Lovelock's critics. Because of his philosophical musings on the non-existence of an all-encompassing

Tree of life The tree of life is a fundamental archetype in many of the world's mythological, religious, and philosophical traditions. It is closely related to the concept of the sacred tree.Giovino, Mariana (2007). ''The Assyrian Sacred Tree: A History ...

, Doolittle has occasionally been cited on

Intelligent Design Intelligent design (ID) is a pseudoscientific argument for the existence of God, presented by its proponents as "an evidence-based scientific theory about life's origins". Numbers 2006, p. 373; " Dcaptured headlines for its bold attempt to ...

blogs. However, though Doolittle argues that a bifurcating tree is not an adequate metaphor for the evolution of life on earth, he is not a supporter of

. A single common ancestor and tree relating all of life on earth is not a necessary component of the theory of descent with modification, the essence of evolution. Doolittle is currently involved in a debate about the proper use of

function (biology) In evolutionary biology, function is the reason some object or process occurred in a system that evolved through natural selection. That reason is typically that it achieves some result, such as that chlorophyll helps to capture the energy of sun ...

within evolutionary biology sparked by controversy over the results of the

ENCODE The Encyclopedia of DNA Elements (ENCODE) is a public research project which aims to identify functional elements in the human genome. ENCODE also supports further biomedical research by "generating community resources of genomics data, software ...

consortium stating that 80% of the genome is "functional". He is a supporter of the concept of

junk DNA Non-coding DNA (ncDNA) sequences are components of an organism's DNA that do not genetic code, encode protein sequences. Some non-coding DNA is Transcription (genetics), transcribed into functional non-coding RNA molecules (e.g. transfer RNA, micro ...

. In addition to his contributions to evolutionary biology, Doolittle is an artist who studied at

NSCAD University NSCAD University, also known as the Nova Scotia College of Art and Design or NSCAD, is a public art university in Halifax, Nova Scotia, Canada. The university is a co-educational institution that offers bachelor's and master's degrees. The uni ...

, achieving a BA in photography.

References

External links

Personal homepage
at

* * * *
W.F.Doolittle on Google Scholar

an article describing Doolittle's views on updating Darwin's theory. 1941 births Living people Harvard University alumni Members of the Norwegian Academy of Science and Letters NSCAD University alumni Members of the United States National Academy of Sciences American biochemists Fellows of the Royal Society of Canada People from Urbana, Illinois Dalhousie University faculty {{US-biochemist-stub