Fauna of Scotland
   HOME

TheInfoList



OR:

The fauna of Scotland is generally typical of the northwest
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
an part of the
Palearctic realm The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa. The realm consists of several bioregions: the Euro-Sibe ...
, although several of the country's larger
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s were hunted to
extinction Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
in historic times and human activity has also led to various species of wildlife being introduced.
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
's diverse
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
environments support 62 species of wild mammals, including a population of
wild cat Felidae () is the family of mammals in the order Carnivora colloquially referred to as cats, and constitutes a clade. A member of this family is also called a felid (). The term "cat" refers both to felids in general and specifically to the dom ...
s, important numbers of
grey Grey (more common in British English) or gray (more common in American English) is an intermediate color between black and white. It is a neutral or achromatic color, meaning literally that it is "without color", because it can be composed o ...
and
harbour seal The harbor (or harbour) seal (''Phoca vitulina''), also known as the common seal, is a true seal found along temperate and Arctic marine coastlines of the Northern Hemisphere. The most widely distributed species of pinniped (walruses, eared se ...
s and the most northerly colony of
bottlenose dolphin Bottlenose dolphins are aquatic mammals in the genus ''Tursiops.'' They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus definitively contains two species: the common ...
s in the world.Matthews (1968) p. 254. Many populations of
moorland Moorland or moor is a type of habitat found in upland areas in temperate grasslands, savannas, and shrublands and montane grasslands and shrublands biomes, characterised by low-growing vegetation on acidic soils. Moorland, nowadays, generally ...
birds, including the
black Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white have o ...
and red grouse live here, and the country has internationally significant
nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materia ...
ing grounds for
seabirds Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine (ocean), marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ev ...
such as the
northern gannet The northern gannet (''Morus bassanus'') is a seabird, the largest species of the gannet family, Sulidae. It is native to the coasts of the Atlantic Ocean, breeding in Western Europe and Northeastern North America. It is the largest seabird in t ...
. The
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of p ...
has become a national icon, and
white-tailed eagle The white-tailed eagle (''Haliaeetus albicilla'') is a very large species of sea eagle widely distributed across temperate Palearctic, Eurasia. Like all eagles, it is a member of the family Accipitridae (or accipitrids) which includes other diur ...
s and
osprey The osprey (''Pandion haliaetus''), , also called sea hawk, river hawk, and fish hawk, is a diurnal, fish-eating bird of prey with a cosmopolitan range. It is a large raptor reaching more than in length and across the wings. It is brown o ...
s have recently re-colonised the land. The
Scottish crossbill The Scottish crossbill (''Loxia scotica'') is a small passerine bird in the finch family Fringillidae. It is endemic to the Caledonian Forests of Scotland, and is the only terrestrial vertebrate species endemic to the United Kingdom. The Scottish ...
is the only
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
in the UK. Scotland's seas are among the most biologically productive in the world; it is estimated that the total number of Scottish marine species exceeds 40,000. The
Darwin Mounds Darwin Mounds is a large field of undersea sand mounds situated off the north west coast of Scotland that were first discovered in May 1998. They provide a unique habitat for ancient deep water coral reefs and were found using remote sensing tech ...
are an important area of deep sea cold water coral reefs discovered in 1998. Inland, nearly 400 genetically distinct populations of
Atlantic salmon The Atlantic salmon (''Salmo salar'') is a species of ray-finned fish in the family Salmonidae. It is the third largest of the Salmonidae, behind Siberian taimen and Pacific Chinook salmon, growing up to a meter in length. Atlantic salmon are ...
live in Scottish rivers. Of the 42 species of fish found in the country's fresh waters, half have arrived by natural colonisation and half by human introduction. Only six amphibians and four land reptiles are
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to Scotland, but many species of
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s live there that are otherwise rare in the United Kingdom (UK). An estimated 14,000 species of insect, including rare bees and butterflies protected by conservation action plans, inhabit Scotland.
Conservation Conservation is the preservation or efficient use of resources, or the conservation of various quantities under physical laws. Conservation may also refer to: Environment and natural resources * Nature conservation, the protection and managem ...
agencies in the UK are concerned that climate change, especially its potential effects on mountain
plateau In geology and physical geography, a plateau (; ; ), also called a high plain or a tableland, is an area of a highland consisting of flat terrain that is raised sharply above the surrounding area on at least one side. Often one or more sides ha ...
s and marine life, threaten much of the fauna of Scotland.


Habitats

Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
enjoys diverse temperate environments, incorporating
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
and
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extant ...
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
s, and
moorland Moorland or moor is a type of habitat found in upland areas in temperate grasslands, savannas, and shrublands and montane grasslands and shrublands biomes, characterised by low-growing vegetation on acidic soils. Moorland, nowadays, generally ...
,
montane Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures fall as elevation increases, causing the ecosystem to stratify. This stratification is a crucial f ...
,
estuarine An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environment ...
,
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
,
ocean The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the wo ...
ic, and
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
landscape A landscape is the visible features of an area of land, its landforms, and how they integrate with natural or man-made features, often considered in terms of their aesthetic appeal.''New Oxford American Dictionary''. A landscape includes the ...
s. About 14% of Scotland is wooded, much of it in forestry plantations, but before humans cleared the land it supported much larger boreal Caledonian and
broad-leaved A broad-leaved, broad-leaf, or broadleaf tree is any tree within the diverse botanical group of angiosperms that has flat leaves and produces seeds inside of fruits. It is one of two general types of trees, the other being a conifer, a tree with ne ...
forests. Although much reduced, significant remnants of the native
Scots pine ''Pinus sylvestris'', the Scots pine (UK), Scotch pine (US) or Baltic pine, is a species of tree in the pine family Pinaceae that is native to Eurasia. It can readily be identified by its combination of fairly short, blue-green leaves and orang ...
woodlands can be found. Seventeen per cent of Scotland is covered by heather moorland and
peatland A mire, peatland, or quagmire is a wetland area dominated by living peat-forming plants. Mires arise because of incomplete decomposition of organic matter, usually litter from vegetation, due to water-logging and subsequent anoxia. All types ...
.
Caithness Caithness ( gd, Gallaibh ; sco, Caitnes; non, Katanes) is a historic county, registration county and lieutenancy area of Scotland. Caithness has a land boundary with the historic county of Sutherland to the west and is otherwise bounded by ...
and
Sutherland Sutherland ( gd, Cataibh) is a historic county, registration county and lieutenancy area in the Highlands of Scotland. Its county town is Dornoch. Sutherland borders Caithness and Moray Firth to the east, Ross-shire and Cromartyshire (later ...
have one of the world's largest and most intact areas of
blanket bog Blanket bog or blanket mire, also known as featherbed bog, is an area of peatland, forming where there is a climate of high rainfall and a low level of evapotranspiration, allowing peat to develop not only in wet hollows but over large expanses o ...
, which supports a distinctive wildlife community. Seventy-five per cent of Scotland's land is classed as agricultural (including some moorland) while urban areas account for around 3%. The coastline is long, and the number of islands with terrestrial vegetation is nearly 800, about 600 of them lying off the west coast. Scotland has more than 90% of the volume and 70% of the total surface area of
fresh water Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
in the United Kingdom. There are more than 30,000 freshwater
loch ''Loch'' () is the Scottish Gaelic, Scots language, Scots and Irish language, Irish word for a lake or sea inlet. It is Cognate, cognate with the Manx language, Manx lough, Cornish language, Cornish logh, and one of the Welsh language, Welsh w ...
s and 6,600 river systems. Under the auspices of the
European Union The European Union (EU) is a supranational political and economic union of member states that are located primarily in Europe. The union has a total area of and an estimated total population of about 447million. The EU has often been des ...
's
Habitats Directive The Habitats Directive (more formally known as Council Directive 92/43/EEC on the Conservation of natural habitats and of wild fauna and flora) is a directive adopted by the European Community in 1992 as a response to the Berne Convention. The E ...
, 244 sites in Scotland covering more than had been accepted by European Commission as Special Areas of Conservation (SAC). Quoting the Scottish Office. (1998). ''People and nature. A new approach to SSSI designations in Scotland''. The Scottish Office, Edinburgh. Retrieved 2 January 2007. Scotland's seas are among the most biologically productive in the world and contain 40,000 or more species. Twenty-four of the SACs are marine sites, and a further nine are coastal with marine and non-marine elements. These marine elements extend to an area of around . The
Darwin Mounds Darwin Mounds is a large field of undersea sand mounds situated off the north west coast of Scotland that were first discovered in May 1998. They provide a unique habitat for ancient deep water coral reefs and were found using remote sensing tech ...
, covering about , are being considered as the first offshore SAC.


Mammals

Scotland was entirely covered in ice during the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
glaciation A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate betw ...
s. As the post-glacial weather warmed and the ice retreated, mammals migrated through the landscape. However, the opening of the English Channel (as sea levels rose) prevented further migrations, so mainland
Britain Britain most often refers to: * The United Kingdom, a sovereign state in Europe comprising the island of Great Britain, the north-eastern part of the island of Ireland and many smaller islands * Great Britain, the largest island in the United King ...
has only two-thirds of the species that reached
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion#Europe, subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, ...
. The
Hebridean The Hebrides (; gd, Innse Gall, ; non, Suðreyjar, "southern isles") are an archipelago off the west coast of the Scottish mainland. The islands fall into two main groups, based on their proximity to the mainland: the Inner and Outer Hebri ...
islands off Scotland's west coast have only half those of Britain. Sixty-two species of mammal live wild in and around Scotland including 13 species found in coastal waters. The populations of a third of the land mammal species are thought to be in decline due to factors including environmental pollution, habitat fragmentation, changes in agricultural practices, particularly overgrazing, and competition from introduced species. No mammal species are unique to Scotland, although the St. Kilda field mouse, ''Apodemus sylvaticus hirtensi'', is an
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
subspecies of the
wood mouse The wood mouse (''Apodemus sylvaticus'') is a murid rodent native to Europe and northwestern Africa. It is closely related to the yellow-necked mouse (''Apodemus flavicollis'') but differs in that it has no band of yellow fur around the neck, ha ...
that reaches twice the size of its mainland cousins, and the
Orkney vole The Orkney vole (''Microtus arvalis orcadensis'') is a population of the common vole (''Microtus arvalis'') found in the Orkney Islands, off the northern coast of Scotland, as well as in the Channel Island of Guernsey. Orkney voles are about 10% ...
or cuttick, ''Microtus arvalis orcadensis'' found only in the
Orkney Orkney (; sco, Orkney; on, Orkneyjar; nrn, Orknøjar), also known as the Orkney Islands, is an archipelago in the Northern Isles of Scotland, situated off the north coast of the island of Great Britain. Orkney is 10 miles (16 km) north ...
archipelago, is a sub-species of the
common vole The common vole (''Microtus arvalis'') is a European rodent. Distribution and habitat The common vole is hardly restricted in means of distribution and habitat and inhabits large areas of Eurasia but, apart from the Orkney vole, not the Bri ...
. It may have been introduced by early settlers about 4,000 years ago. There are various notable domesticated Scottish mammal breeds including Highland Cattle, the
Shetland Pony The Shetland pony is a Scottish breed of pony originating in the Shetland Isles in the north of Scotland. It may stand up to at the withers. It has a heavy coat and short legs, is strong for its size, and is used for riding, driving, and pa ...
,
Eriskay Pony The Eriskay Pony ( gd, Each Beag nan Eilean or ''Each Èirisgeach'') is a breed of pony from Scotland. It is generally grey in colour, and has a dense, waterproof coat that protects it in harsh weather. The breed developed in ancient times in ...
,
Soay Sheep The Soay sheep is a breed of domestic sheep (''Ovis aries'') descended from a population of feral sheep on the island of Soay in the St Kilda Archipelago, about from the Western Isles of Scotland. It is one of the Northern European short-t ...
and
Scottish Terrier The Scottish Terrier ( gd, Abhag Albannach; also known as the Aberdeen Terrier), popularly called the Scottie, is a breed of dog. Initially one of the highland breeds of terrier that were grouped under the name of ''Skye Terrier'', it is one o ...
.


Carnivores

The representation of the weasel family (
Mustelidae The Mustelidae (; from Latin ''mustela'', weasel) are a family of carnivorous mammals, including weasels, badgers, otters, ferrets, martens, minks and wolverines, among others. Mustelids () are a diverse group and form the largest family in ...
) in Scotland is typical of Britain as a whole save that the
polecat Polecat is a common name for several mustelid species in the order Carnivora and subfamilies Ictonychinae and Mustelinae. Polecats do not form a single taxonomic rank (i.e. clade). The name is applied to several species with broad similarities ...
is absent and that Scotland is the UK's stronghold of the
pine marten The European pine marten (''Martes martes''), also known as the pine marten, is a mustelid native to and widespread in most of Europe, Asia Minor, the Caucasus and parts of Iran, Iraq and Syria. It is listed as Least Concern on the IUCN Red List. ...
, although the purity of the latter breed is threatened by a release of American martens in northern
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
. Scotland hosts the only populations of the
Scottish wildcat The Scottish wildcat is a European wildcat (''Felis silvestris silvestris'') Population (biology), population in Scotland. It was once widely distributed across Great Britain, but the population has declined drastically since the turn of the 20th ...
(''Felis silvestris'') in the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
with numbers estimated at between 400 and 2,000 animals, and of the
red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the Order (biology), order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe ...
subspecies ''Vulpes vulpes vulpes,'' a larger race than the more common ''V. v. crucigera'' and which has two distinct forms. The wild cat is at risk due to the inadequacy of protective legislation and is now considered at serious risk of extinction. In 2013 it was announced that the island of
Càrna Carna or Càrna is an island in Loch Sunart, an arm of the sea, close to the Ardnamurchan peninsula, on the west coast of Scotland. Geography Carna lies wedged across the mouth of Loch Teacuis in the middle of Loch Sunart, forming two narro ...
is to provide a sanctuary and breeding station in order to protect the species. Exterminations of the population of feral
American mink The American mink (''Neogale vison'') is a semiaquatic species of mustelid native to North America, though human intervention has expanded its range to many parts of Europe, Asia and South America. Because of range expansion, the American mink i ...
, which were brought to Britain for fur farms in the 1950s, have been undertaken under the auspices of the Hebridean Mink Project and the Scottish Mink Initiative, which hopes to create a mink-free zone in a large area stretching from
Wester Ross Wester Ross () is an area of the Northwest Highlands of Scotland in the council area of Highland. The area is loosely defined, and has never been used as a formal administrative region in its own right, but is generally regarded as lying to the ...
to
Tayside Tayside ( gd, Taobh Tatha) was one of the nine regions used for local government in Scotland from 15 May 1975 to 31 March 1996. The region was named for the River Tay. It was created by the Local Government (Scotland) Act 1973, following rec ...
. Other than occasional
vagrant Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
s, among the seals only the Phocidae, or earless seals, are represented. Two species, the
grey seal The grey seal (''Halichoerus grypus'') is found on both shores of the North Atlantic Ocean. In Latin Halichoerus grypus means "hook-nosed sea pig". It is a large seal of the family Phocidae, which are commonly referred to as "true seals" or " ...
and harbour or
common seal The harbor (or harbour) seal (''Phoca vitulina''), also known as the common seal, is a true seal found along temperate and Arctic marine coastlines of the Northern Hemisphere. The most widely distributed species of pinnipeds, pinniped (walruses ...
, are present around the coast of Scotland in internationally important numbers. In 2002 the Scottish grey seal population was estimated at 120,600 adult animals, which is around 36% of the world population and more than 90% of the UK's. The Scottish population of the common seal is 29,700, about 90% of the UK and 36% of the European total.


Rodents, insectivores and lagomorphs

Seventy-five per cent of the UK's
red squirrel The red squirrel (''Sciurus vulgaris'') is a species of tree squirrel in the genus ''Sciurus'' common throughout Europe and Asia. The red squirrel is an arboreal, primarily herbivorous rodent. In Great Britain, Ireland, and in Italy numbers ...
s are found in Scotland. This species faces threats that include competition from the introduced
grey squirrel The eastern gray squirrel (''Sciurus carolinensis''), also known, particularly outside of North America, as simply the grey squirrel, is a tree squirrel in the genus ''Sciurus''. It is native to eastern North America, where it is the most prodi ...
, and the 'Scottish Strategy for Red Squirrel Conservation' provides a framework for supporting its long-term conservation. Research in 2007 credited the growing population of pine martens with assisting this programme by preying selectively on the grey squirrels. Scotland has no population of the
edible An edible item is any item that is safe for humans to eat. "Edible" is differentiated from "eatable" because it does not indicate how an item tastes, only whether it is fit to be eaten. Nonpoisonous items found in nature – such as some mushroo ...
or
hazel dormouse The hazel dormouse or common dormouse (''Muscardinus avellanarius'') is a small mammal and the only living species in the genus ''Muscardinus''. Distribution and habitat The hazel dormouse is native to northern Europe and Asia Minor. It is the ...
, or of the
yellow-necked mouse The yellow-necked mouse (''Apodemus flavicollis''), also called yellow-necked field mouse, yellow-necked wood mouse, and South China field mouse, is closely related to the wood mouse, with which it was long confused. It was only recognised as a ...
, and the harvest mouse's range is limited to the southern part of the country. The St Kilda mouse and
Orkney vole The Orkney vole (''Microtus arvalis orcadensis'') is a population of the common vole (''Microtus arvalis'') found in the Orkney Islands, off the northern coast of Scotland, as well as in the Channel Island of Guernsey. Orkney voles are about 10% ...
(see above) are endemic, but otherwise population distributions are similar to the rest of mainland Britain. Colonies of
black rat The black rat (''Rattus rattus''), also known as the roof rat, ship rat, or house rat, is a common long-tailed rodent of the stereotypical rat genus ''Rattus'', in the subfamily Murinae. It likely originated in the Indian subcontinent, but is n ...
s remain only on the island of
Inchcolm Inchcolm (from the Scottish Gaelic "Innis Choluim", meaning Columba's Island) is an island in the Firth of Forth in Scotland. It was repeatedly attacked by English raiders during the Wars of Scottish Independence, and was fortified during both Wo ...
in the
Firth of Forth The Firth of Forth () is the estuary, or firth, of several Scottish rivers including the River Forth. It meets the North Sea with Fife on the north coast and Lothian on the south. Name ''Firth'' is a cognate of ''fjord'', a Norse word meani ...
and on the
Shiant Isles The Shiant Islands (; gd, Na h-Eileanan Mòra or ) or Shiant Isles are a privately owned island group in the Minch, east of Harris in the Outer Hebrides of Scotland. They are southeast of the Isle of Lewis.Keay, J. & Keay, J. (1994) ''Collins ...
. Mainland
insectivore A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores wer ...
populations are generally similar to the rest of Britain. Recent steps by
Scottish Natural Heritage NatureScot ( gd, NàdarAlba), which was formerly known as Scottish Natural Heritage, is an executive non-departmental public body of the Scottish Government responsible for the country's natural heritage, especially its natural, genetic and ...
, the Scottish Executive and the
Royal Society for the Protection of Birds The Royal Society for the Protection of Birds (RSPB) is a charitable organisation registered in England and Wales and in Scotland. It was founded in 1889. It works to promote conservation and protection of birds and the wider environment throug ...
to remove
European hedgehog The European hedgehog (''Erinaceus europaeus''), also known as the West European hedgehog or common hedgehog, is a hedgehog species native to Europe from Iberia and Italy northwards into Scandinavia and westwards into the British Isles.Harris, S. ...
s from the
Outer Hebrides The Outer Hebrides () or Western Isles ( gd, Na h-Eileanan Siar or or ("islands of the strangers"); sco, Waster Isles), sometimes known as the Long Isle/Long Island ( gd, An t-Eilean Fada, links=no), is an island chain off the west coast ...
, where their introduction has caused declines in internationally important breeding populations of wading seabird such as
dunlin The dunlin (''Calidris alpina'') is a small wader, formerly sometimes separated with the other "stints" in the genus ''Erolia''. The English name is a dialect form of "dunling", first recorded in 1531–1532. It derives from ''dun'', "dull brown ...
,
ringed plover The common ringed plover or ringed plover (''Charadrius hiaticula'') is a small plover that breeds in Arctic Eurasia. The genus name ''Charadrius'' is a Late Latin word for a yellowish bird mentioned in the fourth-century Vulgate. It derives from ...
and redshank, has caused considerable controversy, and hedgehog culls were halted in 2007. The trapped animals are now relocated to the mainland. The programme has reduced this population; only two individuals were caught in 2007. Of the
lagomorphs The lagomorphs are the members of the taxonomic order Lagomorpha, of which there are two living families: the Leporidae ( hares and rabbits) and the Ochotonidae (pikas). The name of the order is derived from the Ancient Greek ''lagos'' (λα ...
only hares and rabbits are represented in Scotland. The
mountain hare The mountain hare (''Lepus timidus''), also known as blue hare, tundra hare, variable hare, white hare, snow hare, alpine hare, and Irish hare, is a Palearctic hare that is largely adapted to polar and mountainous habitats. Evolution The mount ...
is the only native member of the hare family and is the dominant species throughout most of upland Scotland. The
European hare The European hare (''Lepus europaeus''), also known as the brown hare, is a species of hare native to Europe and parts of Asia. It is among the largest hare species and is adapted to temperate, open country. Hares are herbivorous and feed mainly ...
and
European rabbit The European rabbit (''Oryctolagus cuniculus'') or coney is a species of rabbit native to the Iberian Peninsula (including Spain, Portugal, and southwestern France), western France, and the northern Atlas Mountains in northwest Africa. It has ...
are both present, the latter having been brought to Britain by the
Roman Roman or Romans most often refers to: *Rome, the capital city of Italy *Ancient Rome, Roman civilization from 8th century BC to 5th century AD *Roman people, the people of ancient Rome *''Epistle to the Romans'', shortened to ''Romans'', a letter ...
s but not becoming widespread in Scotland until the 19th century.


Artiodactyls

Landseer's painting of a
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
stag Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
, '' Monarch of the Glen'', is one of the most notable images of Victorian Scotland. The species, a member of the biological order
artiodactyla The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
or "even-toed ungulates", is still 400,000 strong, although its existence in the pure form is threatened by hybridisation with introduced
sika deer The sika deer (''Cervus nippon''), also known as the Northern spotted deer or the Japanese deer, is a species of deer native to much of East Asia and introduced to other parts of the world. Previously found from northern Vietnam in the south to ...
. Very much a hill-dwelling species in Scotland (and so typically smaller in stature than its European forest-loving cousins), it is generally replaced by
roe deer The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
in lower-lying land. Although found elsewhere in the UK, no wild populations of
Chinese water deer The water deer (''Hydropotes inermis'') is a small deer superficially more similar to a musk deer than a true deer. Native to China and Korea, there are two subspecies: the Chinese water deer (''Hydropotes inermis inermis'') and the Korean wat ...
and no or very few Chinese muntjac exist in Scotland. It has isolated populations of
feral A feral () animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some ...
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s ''Capra hircus'' and feral
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated s ...
(''Ovis aries''), such as the herd of 1,000
Soay sheep The Soay sheep is a breed of domestic sheep (''Ovis aries'') descended from a population of feral sheep on the island of Soay in the St Kilda Archipelago, about from the Western Isles of Scotland. It is one of the Northern European short-t ...
on St Kilda. Since 1952 a herd of
reindeer Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subspe ...
have lived in the
Cairngorm National Park Cairngorms National Park ( gd, Pàirc Nàiseanta a' Mhonaidh Ruaidh) is a national park in northeast Scotland, established in 2003. It was the second of two national parks established by the Scottish Parliament, after Loch Lomond and The Tro ...
, the species having become extinct in Scotland after it was recorded as having been hunted in
Orkney Orkney (; sco, Orkney; on, Orkneyjar; nrn, Orknøjar), also known as the Orkney Islands, is an archipelago in the Northern Isles of Scotland, situated off the north coast of the island of Great Britain. Orkney is 10 miles (16 km) north ...
in the 12th century.


Other mammals

Only nine of the sixteen or seventeen bat species found elsewhere in Britain are present in Scotland. Widespread species are
common Common may refer to: Places * Common, a townland in County Tyrone, Northern Ireland * Boston Common, a central public park in Boston, Massachusetts * Cambridge Common, common land area in Cambridge, Massachusetts * Clapham Common, originally com ...
and
soprano pipistrelle The soprano pipistrelle (''Pipistrellus pygmaeus'') is a small species of bat. It is found in Europe and often roosts on buildings. Taxonomy Until 1999, the soprano pipistrelle was considered as conspecific with the common pipistrelle The co ...
s, the
brown long-eared bat The brown long-eared bat or common long-eared bat (''Plecotus auritus'') is a small Eurasian insectivorous bat. It has distinctive ears, long and with a distinctive fold. It is extremely similar to the much rarer grey long-eared bat which was o ...
,
Daubenton's bat Daubenton's bat or Daubenton's myotis (''Myotis daubentonii'') is a Eurasian bat with rather short ears. It ranges from Ireland to Japan (Hokkaido) and is considered to be increasing its numbers in many areas. This bat was first described in 181 ...
and
Natterer's bat Natterer's bat (''Myotis nattereri'') is a European vespertilionid bat with pale wings. It has brown fur tending to greyish-white on its underside. It is found across most of the continent of Europe, parts of the Near East and North Africa. It fe ...
. Those with a more restricted distribution are the
whiskered bat The whiskered bat (''Myotis mystacinus'') is a small European bat with long fur. Although uncommon, ''M. mystacinus'' is often found around human habitation and around water; it is similar to Brandt's bat Brandt's bat or Brandt's myotis (''M ...
,
noctule ''Nyctalus'' is a genus of vespertilionid bats commonly known as the noctule bats. They are distributed in the temperate and subtropical areas of Europe, Asia and North Africa. There are eight species within this genus: * Birdlike noctule, ''N ...
,
Leisler's bat The lesser noctule, Leisler's bat or the Irish bat (''Nyctalus leisleri''), is a species of insectivorous bat belonging to the vesper bat family, Vespertilionidae. The species was named to honour the naturalist Johann Philipp Achilles Leisler ...
and
Nathusius's pipistrelle Nathusius' pipistrelle (''Pipistrellus nathusii'') is a small bat in the genus ''Pipistrellus''. It is very similar to the common pipistrelle and has been overlooked in many areas until recently but it is widely distributed across Europe. It was ...
. Absences include the greater and lesser horseshoe bat, the
greater mouse-eared bat The greater mouse-eared bat (''Myotis myotis'') is a European species of bat in the family Vespertilionidae. Description ''Myotis myotis'' is a large bat with a long, broad muzzle and big, long ears. The body's dorsal side is brown to reddish-b ...
and
Bechstein's bat Bechstein's bat (''Myotis bechsteinii'') is a species of vesper bat found in Europe and western Asia, living in extensive areas of woodland. Description Bechstein's bat is a medium-sized and relatively long-eared bat. The adult has a long, fluf ...
. No bats reside in the Shetland Islands; the only records there are of migrants or vagrants. Twenty-one species of
cetacea Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel them ...
n have been recorded in Scottish waters within the last 100 years including
Cuvier's beaked whale The Cuvier's beaked whale, goose-beaked whale, or ziphius (''Ziphius cavirostris'') is the most widely distributed of all beaked whales in the family Ziphiidae. It is smaller than most baleen whales yet large among beaked whales. Cuvier's beaked ...
,
killer whale The orca or killer whale (''Orcinus orca'') is a toothed whale belonging to the oceanic dolphin family, of which it is the largest member. It is the only extant species in the genus ''Orcinus'' and is recognizable by its black-and-white pa ...
s,
sperm whale The sperm whale or cachalot (''Physeter macrocephalus'') is the largest of the toothed whales and the largest toothed predator. It is the only living member of the genus ''Physeter'' and one of three extant species in the sperm whale famil ...
s,
minke whale The minke whale (), or lesser rorqual, is a species complex of baleen whale. The two species of minke whale are the common (or northern) minke whale and the Antarctic (or southern) minke whale. The minke whale was first described by the Danish n ...
s and
common Common may refer to: Places * Common, a townland in County Tyrone, Northern Ireland * Boston Common, a central public park in Boston, Massachusetts * Cambridge Common, common land area in Cambridge, Massachusetts * Clapham Common, originally com ...
, white-beaked and
Risso's dolphin Risso's dolphin (''Grampus griseus'') is a dolphin, the only species of the genus ''Grampus''. Some of the closest related species to these dolphins include: pilot whales (''Globicephala'' spp.), pygmy killer whales (''Feresa attenuata''), melon ...
s. The
Moray Firth The Moray Firth (; Scottish Gaelic: ''An Cuan Moireach'', ''Linne Mhoireibh'' or ''Caolas Mhoireibh'') is a roughly triangular inlet (or firth) of the North Sea, north and east of Inverness, which is in the Highland council area of north of Scotl ...
colony of about 100
bottlenose dolphin Bottlenose dolphins are aquatic mammals in the genus ''Tursiops.'' They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus definitively contains two species: the common ...
s is the most northerly in the world. As recent dramatic television coverage indicated, this species preys on
harbour porpoise The harbour porpoise (''Phocoena phocoena'') is one of eight extant species of porpoise. It is one of the smallest species of cetacean. As its name implies, it stays close to coastal areas or river estuaries, and as such, is the most familiar ...
s; a third of the porpoise carcasses examined by pathologists from 1992 to 2002 indicated that death resulted from dolphin attacks. However, conservationists expressed dismay that the UK government decided to allow oil and gas prospecting in the Moray Firth, putting these populations of cetaceans at risk. In response, the government have placed
seismic Seismology (; from Ancient Greek σεισμός (''seismós'') meaning "earthquake" and -λογία (''-logía'') meaning "study of") is the scientific study of earthquakes and the propagation of elastic waves through the Earth or through other ...
surveys "on hold" during 2009 pending further research. The introduced
marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a po ...
, the
red-necked wallaby The red-necked wallaby or Bennett's wallaby (''Notamacropus rufogriseus'') is a medium-sized macropod marsupial (wallaby), common in the more temperate and fertile parts of eastern Australia, including Tasmania. Red-necked wallabies have been ...
, is confined to a colony on an island in
Loch Lomond Loch Lomond (; gd, Loch Laomainn - 'Lake of the Elms'Richens, R. J. (1984) ''Elm'', Cambridge University Press.) is a freshwater Scottish loch which crosses the Highland Boundary Fault, often considered the boundary between the lowlands of Ce ...
.


Extinctions and reintroductions

During the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
interglacials,
arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
animals that are no longer extant occupied Scotland, including the
woolly rhinoceros The woolly rhinoceros (''Coelodonta antiquitatis'') is an extinct species of rhinoceros that was common throughout Europe and Asia during the Pleistocene epoch and survived until the end of the last glacial period. The woolly rhinoceros was a me ...
,
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks and, ...
,
polar bear The polar bear (''Ursus maritimus'') is a hypercarnivorous bear whose native range lies largely within the Arctic Circle, encompassing the Arctic Ocean, its surrounding seas and surrounding land masses. It is the largest extant bear specie ...
,
lemming A lemming is a small rodent, usually found in or near the Arctic in tundra biomes. Lemmings form the subfamily Arvicolinae (also known as Microtinae) together with voles and muskrats, which form part of the superfamily Muroidea, which also include ...
,
Arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small fox native to the Arctic regions of the Northern Hemisphere and common throughout the Arctic tundra biome. It is well adapted to living in co ...
and the giant deer ''
Megaloceros giganteus The Irish elk (''Megaloceros giganteus''), also called the giant deer or Irish deer, is an extinct species of deer in the genus '' Megaloceros'' and is one of the largest deer that ever lived. Its range extended across Eurasia Eurasia ( ...
''. Other mammals that used to inhabit Scotland but became extinct in the wild during historic times include the
Eurasian lynx The Eurasian lynx (''Lynx lynx'') is a medium-sized wild cat widely distributed from Northern, Central and Eastern Europe to Central Asia and Siberia, the Tibetan Plateau and the Himalayas. It inhabits temperate and boreal forests up to an eleva ...
, which lived in Britain until 1,500 years ago, the
European brown bear The Eurasian brown bear (''Ursus arctos arctos'') is one of the most common subspecies of the brown bear, and is found in much of Eurasia. It is also called the European brown bear, common brown bear, common bear, and colloquially by many ot ...
, subspecies ''Ursus arctos caledoniensis'', which was taken to entertain the
Roman Roman or Romans most often refers to: *Rome, the capital city of Italy *Ancient Rome, Roman civilization from 8th century BC to 5th century AD *Roman people, the people of ancient Rome *''Epistle to the Romans'', shortened to ''Romans'', a letter ...
circus A circus is a company of performers who put on diverse entertainment shows that may include clowns, acrobats, trained animals, trapeze acts, musicians, dancers, hoopers, tightrope walkers, jugglers, magicians, ventriloquists, and unicyclist ...
es but died out in the 9th or 10th century, and the
elk The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The common ...
, which lasted until about 1300. The
wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The species is ...
and wild ox or urus died out in the subsequent two centuries, although the former's domesticated cousin, the
grice The grice was a breed of swine found in the Highlands and Islands of Scotland and in Ireland. Culley, George(1807), ''Observations on Livestock'', pub Wilkie, Robinson et al, p 176/ref> It became extinct, surviving longest in the Shetland I ...
, lasted until 1930 in
Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...
. The last known
wolf The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly u ...
was shot on
Mackintosh The Mackintosh or raincoat (abbreviated as mac) is a form of waterproof raincoat, first sold in 1824, made of rubberised fabric. The Mackintosh is named after its Scottish inventor Charles Macintosh, although many writers added a letter ''k' ...
land in
Inverness-shire Inverness-shire ( gd, Siorrachd Inbhir Nis) is a historic county, registration county and lieutenancy area of Scotland. Covering much of the Highlands and Outer Hebrides, it is Scotland's largest county, though one of the smallest in populatio ...
in 1743,Fraser Darling and Boyd (1969) p. 64 and the
walrus The walrus (''Odobenus rosmarus'') is a large pinniped, flippered marine mammal with a discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. The walrus is the only living species in ...
is now only an occasional vagrant. St Kilda also possessed an endemic subspecies of the
house mouse The house mouse (''Mus musculus'') is a small mammal of the order Rodentia, characteristically having a pointed snout, large rounded ears, and a long and almost hairless tail. It is one of the most abundant species of the genus '' Mus''. Althoug ...
, '' Mus musculus muralis'', which was longer, hairier, coloured differently and had a skull shape at variance to the norm. It became extinct in 1938, just eight years after the evacuation of the native St Kildans. A joint project of the
Royal Zoological Society of Scotland The Royal Zoological Society of Scotland is a learned society and registered charity based in Edinburgh, Scotland. It was founded by Thomas Haining Gillespie in 1909. In 1913, Edinburgh Town Council bought a large plot of land on Corstorphin ...
, the
Scottish Wildlife Trust The Scottish Wildlife Trust is a Office of the Scottish Charity Regulator, registered charity dedicated to conserving the Natural history of Scotland, wildlife and natural environment of Scotland. Description The Scottish Wildlife Trust has well ...
and Forestry Commission Scotland have successfully re-introduced the
European beaver The Eurasian beaver (''Castor fiber'') or European beaver is a beaver species that was once widespread in Eurasia, but was hunted to near-extinction for both its fur and castoreum. At the turn of the 20th century, only about 1,200 beavers surviv ...
to the wild in Scotland using Norwegian stock. The species was found in the
Highlands Highland is a broad term for areas of higher elevation, such as a mountain range or mountainous plateau. Highland, Highlands, or The Highlands, may also refer to: Places Albania * Dukagjin Highlands Armenia * Armenian Highlands Australia *Sou ...
until the 15th century, and although the then Scottish Government initially rejected the idea, a trial commenced in May 2009 in
Knapdale Knapdale ( gd, Cnapadal, IPA: kraʰpət̪əɫ̪ forms a rural district of Argyll and Bute in the Scottish Highlands, adjoining Kintyre to the south, and divided from the rest of Argyll to the north by the Crinan Canal. It includes two parishes, ...
. Separately, on Tayside, deliberate releases or escapes have led to up to 250 animals colonising the area. Although it was initially planned to remove these unofficially reintroduced beavers, in March 2012 the Scottish Government reversed the decision to remove beavers from the Tay, pending the outcome of studies into the suitability of re-introduction. Following receipt of the results of the studies, in November 2016 the Scottish Government announced that beavers could remain permanently, and would be given protected status as a native species within Scotland. Beavers will be allowed to extend their range naturally from Knapdale and along the River Tay, however to aid this process and improve the health and resilience of the population a further 28 beavers will be released in Knapdale between 2017 and 2020. By means of escapes or deliberate releases, wild boar (Sus scrofa) have been re-introduced to several places in Scotland including a wide area of Lochaber and West Inverness-shire. Various other schemes are under consideration. For example, the owner of the Alladale estate north of
Inverness Inverness (; from the gd, Inbhir Nis , meaning "Mouth of the River Ness"; sco, Innerness) is a city in the Scottish Highlands. It is the administrative centre for The Highland Council and is regarded as the capital of the Highlands. Histori ...
has expressed a desire to reintroduce
wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
as part of a wilderness reserve, the first of its kind in Britain.


Avifauna

The history of mammals suggests three broad overlapping phases: natural colonisation after the ice age, human-caused extinctions, and introduction by humans of non-native species.MacCormick, Finbar and Buckland, Paul C. ''The Vertebrate Fauna'' in Edwards, Kevin J. & Ralston, Ian B.M. (Eds) (2003) ''Scotland After the Ice Age: Environment, Archaeology and History, 8000 BC – AD 1000''. Edinburgh. Edinburgh University Press. pp. 83–103. The greater mobility of birds makes such generalisations hard to substantiate in their case. Modern humans have done great damage to bird species, especially the raptors, but natural variations in populations are complex. For example,
northern fulmar The northern fulmar (''Fulmarus glacialis''), fulmar, or Arctic fulmar is a highly abundant seabird found primarily in subarctic regions of the North Atlantic and North Pacific oceans. There has been one confirmed sighting in the Southern Hem ...
s were present at
Skara Brae Skara Brae is a stone-built Neolithic settlement, located on the Bay of Skaill on the west coast of Mainland, the largest island in the Orkney archipelago of Scotland. Consisting of ten clustered houses, made of flagstones, in earthen dams ...
during the
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several parts ...
period, but in
medieval In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the Post-classical, post-classical period of World history (field), global history. It began with t ...
times their breeding range was restricted to St Kilda. Since then they have spread throughout the British Isles. Most of about 250 species of bird regularly recorded in Britain venture into Scotland, and perhaps up to 300 more occur with varying degrees of rarity. A total of 247 species have been assessed and each placed onto one of three lists, red, amber or green, indicating the level of concern for their future. Forty species are red-listed, 121 are amber-listed and 86 are green-listed. The
Scottish crossbill The Scottish crossbill (''Loxia scotica'') is a small passerine bird in the finch family Fringillidae. It is endemic to the Caledonian Forests of Scotland, and is the only terrestrial vertebrate species endemic to the United Kingdom. The Scottish ...
, ''Loxia scotica'', which inhabits the coniferous forests of the
Highlands Highland is a broad term for areas of higher elevation, such as a mountain range or mountainous plateau. Highland, Highlands, or The Highlands, may also refer to: Places Albania * Dukagjin Highlands Armenia * Armenian Highlands Australia *Sou ...
, is Britain's only endemic bird and, with only 300 breeding pairs, one of Europe's most threatened species. Its shape, red/green hue and habit of hanging upside down has led to comparisons with
parrot Parrots, also known as psittacines (), are birds of the roughly 398 species in 92 genera comprising the order Psittaciformes (), found mostly in tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittacoid ...
s. St Kilda has a unique subspecies of wren, the
St Kilda wren The St Kilda wren (''Troglodytes troglodytes hirtensis'') is a small passerine bird in the wren family. It is a distinctive subspecies of the Eurasian wren endemic to the islands of the isolated St Kilda archipelago, in the Atlantic Ocean we ...
''Troglodytes troglodytes hirtensis'', which has adapted to perching on the rocks and cliffs of this treeless
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
island, and consequently has developed larger and stronger feet than the mainland variant. It is also slightly larger, has a longer beak, a drabber though more varied colouring, and a "peculiarly sweet and soft" song. The subspecies was recognised in 1884 and was protected by a special
Act of Parliament Acts of Parliament, sometimes referred to as primary legislation, are texts of law passed by the Legislature, legislative body of a jurisdiction (often a parliament or council). In most countries with a parliamentary system of government, acts of ...
in 1904 to prevent its destruction "at the hands of ornithologists, egg-collectors, taxidermists and tourists".


Raptors

All but a few pairs of Britain's approximately 600
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of p ...
s are found in Scotland as are most of the breeding
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a Cosmopolitan distribution, cosmopolitan bird of prey (Bird of prey, raptor) in the family (biology), family Falco ...
s. The
hobby A hobby is considered to be a regular activity that is done for enjoyment, typically during one's leisure time. Hobbies include collecting themed items and objects, engaging in creative and artistic pursuits, playing Sport, sports, or pursu ...
,
marsh harrier The marsh harriers are birds of prey of the harrier subfamily. They are medium-sized raptors and the largest and broadest-winged harriers. Most of them are associated with marshland and dense reedbeds. They are found almost worldwide, excluding ...
and
Montagu's harrier Montagu's harrier (''Circus pygargus'') is a migratory bird of prey of the harrier family. Its common name commemorates the British naturalist George Montagu. Taxonomy The first formal description of Montagu's harrier was by the Swedish na ...
although found in
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
and
Wales Wales ( cy, Cymru ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by England to the Wales–England border, east, the Irish Sea to the north and west, the Celtic Sea to the south west and the ...
are generally absent. In 1916 an English
vicar A vicar (; Latin: ''vicarius'') is a representative, deputy or substitute; anyone acting "in the person of" or agent for a superior (compare "vicarious" in the sense of "at second hand"). Linguistically, ''vicar'' is cognate with the English pref ...
stole the last native
white-tailed sea eagle The white-tailed eagle (''Haliaeetus albicilla'') is a very large species of sea eagle widely distributed across temperate Eurasia. Like all eagles, it is a member of the family Accipitridae (or accipitrids) which includes other diurnal raptor ...
eggs on
Skye The Isle of Skye, or simply Skye (; gd, An t-Eilean Sgitheanach or ; sco, Isle o Skye), is the largest and northernmost of the major islands in the Inner Hebrides of Scotland. The island's peninsulas radiate from a mountainous hub dominated b ...
, p. 35. and the last adult was shot in
Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...
two years later. However, the species was reintroduced to the island of
Rùm Rùm (), a Scottish Gaelic name often anglicised to Rum (), is one of the Small Isles of the Inner Hebrides, in the district of Lochaber, Scotland. For much of the 20th century the name became Rhum, a spelling invented by the former owner, Sir ...
in 1975. The bird spread successfully to various neighbouring islands, and 30 pairs were established by 2006. Despite fears expressed by local farmers, the
Royal Society for the Protection of Birds The Royal Society for the Protection of Birds (RSPB) is a charitable organisation registered in England and Wales and in Scotland. It was founded in 1889. It works to promote conservation and protection of birds and the wider environment throug ...
(RSPB) are in process of releasing up to 100 young eagles on the east coast in the
Forth Forth or FORTH may refer to: Arts and entertainment * ''forth'' magazine, an Internet magazine * ''Forth'' (album), by The Verve, 2008 * ''Forth'', a 2011 album by Proto-Kaw * Radio Forth, a group of independent local radio stations in Scotla ...
and
Tay Tay may refer to: People and languages * Tay (name), including lists of people with the given name, surname and nickname * Tay people, an ethnic group of Vietnam ** Tày language *Atayal language, an Austronesian language spoken in Taiwan (ISO 639 ...
estuaries. The
red kite The red kite (''Milvus milvus'') is a medium-large bird of prey in the family Accipitridae, which also includes many other diurnal raptors such as eagles, buzzards, and harriers. The species currently breeds in the Western Palearctic region o ...
was exterminated in Scotland in 1879, and a reintroduction programme was launched by the RSPB in the 1980s. Although the species has made significant advances, it is estimated that 38% of the 395 birds fledged between 1999 and 2003 were poisoned and a further 9% shot or otherwise killed by humans. The RSPB stated: "it may take a custodial sentence before people engaged with this activity begin to take the matter seriously". After an absence of nearly 40 years the
osprey The osprey (''Pandion haliaetus''), , also called sea hawk, river hawk, and fish hawk, is a diurnal, fish-eating bird of prey with a cosmopolitan range. It is a large raptor reaching more than in length and across the wings. It is brown o ...
successfully re-colonised Scotland in the early 1950s. In 1899 they had bred at the ruined Loch an Eilean castle near
Aviemore Aviemore (; gd, An Aghaidh Mhòr ) is a town and tourist resort, situated within the Cairngorms National Park in the Highlands of Scotland. It is in the Badenoch and Strathspey committee area, within the Highland council area. The town is popul ...
and at
Loch Arkaig Loch Arkaig (Scottish Gaelic: Loch Airceig) is a body of freshwater in Lochaber, Scotland, to the west of the Great Glen. It is approximately in length and lies above sea level, the maximum depth is around The main tributaries are the Dessarr ...
until 1908. In 1952 they claimed a new site at Loch Garten. There are now 150 breeding pairs. Other raptor species found in the UK such as the
kestrel The term kestrel (from french: crécerelle, derivative from , i.e. ratchet) is the common name given to several species of predatory birds from the falcon genus ''Falco''. Kestrels are most easily distinguished by their typical hunting behaviou ...
,
hen harrier The hen harrier (''Circus cyaneus'') is a bird of prey. It breeds in Eurasia. The term "hen harrier" refers to its former habit of preying on free-ranging fowl. It migrates to more southerly areas in winter. Eurasian birds move to southern Eur ...
,
goshawk Goshawk may refer to several species of birds of prey, mainly in the genus ''Accipiter'': * Northern goshawk, ''Accipiter gentilis'', often referred to simply as the goshawk, since it is the only goshawk found in much of its range (in Europe and N ...
,
sparrowhawk Sparrowhawk (sometimes sparrow hawk) may refer to several species of small hawk in the genus ''Accipiter''. "Sparrow-hawk" or sparhawk originally referred to ''Accipiter nisus'', now called "Eurasian" or "northern" sparrowhawk to distinguish it f ...
,
tawny owl The tawny owl (''Strix aluco''), also called the brown owl, is commonly found in woodlands across Europe to western Siberia, and has seven recognized subspecies. It is a stocky, medium-sized owl, whose underparts are pale with dark streaks, an ...
, and barn owl are widely distributed in Scotland, although the little owl is confined to the south.
Buzzards Buzzard is the common name of several species of birds of prey. ''Buteo'' species * Archer's buzzard (''Buteo archeri'') * Augur buzzard (''Buteo augur'') * Broad-winged hawk (''Buteo platypterus'') * Common buzzard (''Buteo buteo'') * Eastern ...
have displayed a remarkable resilience, having recovered from human persecution and the
myxomatosis Myxomatosis is a disease caused by ''Myxoma virus'', a poxvirus in the genus ''Leporipoxvirus''. The natural hosts are tapeti (''Sylvilagus brasiliensis'') in South and Central America, and brush rabbits (''Sylvilagus bachmani'') in North Ame ...
epidemic of the 1950s, which reduced their food supply. Numbers more than trebled between 1978 and 1998. At the other end of the population scale, a single pair of
snowy owl The snowy owl (''Bubo scandiacus''), also known as the polar owl, the white owl and the Arctic owl, is a large, white owl of the true owl family. Snowy owls are native to the Arctic regions of both North America and the Palearctic, breeding mos ...
s bred on
Fetlar Fetlar ( sco, Fetlar) is one of the North Isles of Shetland, Scotland, with a usually resident population of 61 at the time of the 2011 census. Its main settlement is Houbie on the south coast, home to the Fetlar Interpretive Centre. Fetlar i ...
from 1967 to 1975. In 2009 it was reported that the Scottish Government have decided to proceed with a controversial plan to relocate sparrowhawks found near pigeon lofts in Glasgow, Edinburgh, Kilmarnock, Stirling and Dumfries at a cost of £25,000.


Seabirds

Scotland's seas host almost half of the
European Union The European Union (EU) is a supranational political and economic union of member states that are located primarily in Europe. The union has a total area of and an estimated total population of about 447million. The EU has often been des ...
's breeding
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
s including about half of the world's
northern gannet The northern gannet (''Morus bassanus'') is a seabird, the largest species of the gannet family, Sulidae. It is native to the coasts of the Atlantic Ocean, breeding in Western Europe and Northeastern North America. It is the largest seabird in t ...
s and a third of the world's
Manx shearwater The Manx shearwater (''Puffinus puffinus'') is a medium-sized shearwater in the seabird family Procellariidae. The scientific name of this species records a name shift: Manx shearwaters were called Manks puffins in the 17th century. Puffin is an ...
s. Four seabird species have more than 95% of their combined British and Irish population in Scotland, while a further fourteen species have more than half of their breeding population in Scottish colonies. St Kilda, which is a
World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
, is a seabird haven of great significance. It has 60,000
northern gannet The northern gannet (''Morus bassanus'') is a seabird, the largest species of the gannet family, Sulidae. It is native to the coasts of the Atlantic Ocean, breeding in Western Europe and Northeastern North America. It is the largest seabird in t ...
s, amounting to 24% of the world population, 49,000 breeding pairs of Leach's storm petrel, up to 90% of the European population, 136,000 pairs of puffin and 67,000
northern fulmar The northern fulmar (''Fulmarus glacialis''), fulmar, or Arctic fulmar is a highly abundant seabird found primarily in subarctic regions of the North Atlantic and North Pacific oceans. There has been one confirmed sighting in the Southern Hem ...
pairs, about 30% and 13% of the respective UK totals. The island of
Mingulay Mingulay ( gd, Miughalaigh) is the second largest of the Bishop's Isles in the Outer Hebrides of Scotland. Located south of Barra, it is known for its important seabird populations, including puffins, black-legged kittiwakes, and razorbills, ...
also has a large seabird population and is an important breeding ground for
razorbill The razorbill, razor-billed auk, or lesser auk (''Alca torda'') is a colonial seabird and the only extant member of the genus '' Alca'' of the family Alcidae, the auks. It is the closest living relative of the extinct great auk (''Pinguinis impe ...
s, with 9,514 pairs, 6.3% of the European population. Sixty per cent of all breeding bonxies nest in Scotland, mostly in Orkney and Shetland, even though they did not arrive at all until the 18th century. Scotland is the breeding station for about 90% of the UK's
Arctic tern The Arctic tern (''Sterna paradisaea'') is a tern in the family Laridae. This bird has a circumpolar breeding distribution covering the Arctic and sub-Arctic regions of Europe (as far south as Brittany), Asia, and North America (as far south a ...
s, the majority of which make use of colonies in
Orkney Orkney (; sco, Orkney; on, Orkneyjar; nrn, Orknøjar), also known as the Orkney Islands, is an archipelago in the Northern Isles of Scotland, situated off the north coast of the island of Great Britain. Orkney is 10 miles (16 km) north ...
and
Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...
. A similar percentage of the UK's tysties breed on Scottish islands including
Unst Unst (; sco, Unst; nrn, Ønst) is one of the North Isles of the Shetland Islands, Scotland. It is the northernmost of the inhabited British Isles and is the third-largest island in Shetland after Mainland and Yell. It has an area of . Unst ...
,
Mingulay Mingulay ( gd, Miughalaigh) is the second largest of the Bishop's Isles in the Outer Hebrides of Scotland. Located south of Barra, it is known for its important seabird populations, including puffins, black-legged kittiwakes, and razorbills, ...
and
Iona Iona (; gd, Ì Chaluim Chille (IPA: iːˈxaɫ̪ɯimˈçiʎə, sometimes simply ''Ì''; sco, Iona) is a small island in the Inner Hebrides, off the Ross of Mull on the western coast of Scotland. It is mainly known for Iona Abbey, though there ...
. Scotland also hosts 1,000 pairs of
Arctic skua The parasitic jaeger (''Stercorarius parasiticus''), also known as the Arctic skua, Arctic jaeger or parasitic skua, is a seabird in the skua family Stercorariidae. It is a migratory species that breeds in Northern Scandinavia, Scotland, Iceland ...
and 21,000 breeding pairs of
shag Shag, or Shagged, or Shagger, or Shagging, or Shags may refer to: Animals * Shag or cormorant, a bird family ** European shag, a specific species of the shag or cormorant family ** Great cormorant another species of the family People Pseudonym ...
, 40% of the global population of the species. In excess of 130,000 birds inhabit
Fowlsheugh Fowlsheugh is a coastal nature reserve in Kincardineshire, northeast Scotland, known for its cliff formations and habitat supporting prolific seabird nesting colonies. Designated as a Site of Special Scientific Interest (SSSI) by Scottish Natura ...
nature reserve A nature reserve (also known as a wildlife refuge, wildlife sanctuary, biosphere reserve or bioreserve, natural or nature preserve, or nature conservation area) is a protected area of importance for flora, fauna, or features of geological or ...
in
Aberdeenshire Aberdeenshire ( sco, Aiberdeenshire; gd, Siorrachd Obar Dheathain) is one of the 32 Subdivisions of Scotland#council areas of Scotland, council areas of Scotland. It takes its name from the County of Aberdeen which has substantially differe ...
at the peak of the breeding season, making it one of the largest seabird colonies in Britain. There are significant numbers of
kittiwake The kittiwakes (genus ''Rissa'') are two closely related seabird species in the gull family Laridae, the black-legged kittiwake (''Rissa tridactyla'') and the red-legged kittiwake (''Rissa brevirostris''). The epithets "black-legged" and "red-l ...
, Atlantic puffin, razorbill, fulmar,
herring gull Herring gull is a common name for several birds in the genus ''Larus'', all formerly treated as a single species. Three species are still combined in some taxonomies: * American herring gull (''Larus smithsonianus'') - North America * European he ...
and
great black-backed gull The great black-backed gull (''Larus marinus'') is the largest member of the gull family. Described by the Cornell Lab of Ornithology as "the king of the Atlantic waterfront", it is a very aggressive hunter, pirate, and scavenger. It breeds on t ...
. The
Bass Rock The Bass Rock, or simply the Bass (), ( gd, Creag nam Bathais or gd, Am Bas) is an island in the outer part of the Firth of Forth in the east of Scotland. Approximately offshore, and north-east of North Berwick, it is a steep-sided volcan ...
in the
Firth of Forth The Firth of Forth () is the estuary, or firth, of several Scottish rivers including the River Forth. It meets the North Sea with Fife on the north coast and Lothian on the south. Name ''Firth'' is a cognate of ''fjord'', a Norse word meani ...
hosts upwards of 40,000 pairs of
northern gannet The northern gannet (''Morus bassanus'') is a seabird, the largest species of the gannet family, Sulidae. It is native to the coasts of the Atlantic Ocean, breeding in Western Europe and Northeastern North America. It is the largest seabird in t ...
s and is the largest single rock gannetry in the world. The bird's scientific name ''Morus bassanus'', derives from the rock.


Game birds, waders and water fowl

Red-listed
western capercaillie The western capercaillie (''Tetrao urogallus''), also known as the Eurasian capercaillie, wood grouse, heather cock, cock-of-the-woods, or simply capercaillie , is a heavy member of the grouse family and the largest of all extant grouse species. ...
and
ptarmigan ''Lagopus'' is a small genus of birds in the grouse subfamily commonly known as ptarmigans (). The genus contains three living species with numerous described subspecies, all living in tundra or cold upland areas. Taxonomy and etymology The ge ...
breed in Scotland and are absent elsewhere in the British Isles. The former became extinct in Scotland in 1785 but was successfully reintroduced from
Swedish Swedish or ' may refer to: Anything from or related to Sweden, a country in Northern Europe. Or, specifically: * Swedish language, a North Germanic language spoken primarily in Sweden and Finland ** Swedish alphabet, the official alphabet used by ...
stock in 1837. There are significant populations of other
Galliformes Galliformes is an order of heavy-bodied ground-feeding birds that includes turkeys, chickens, quail, and other landfowl. Gallinaceous birds, as they are called, are important in their ecosystems as seed dispersers and predators, and are often ...
including
blackcock The black grouse (''Lyrurus tetrix''), also known as northern black grouse, Eurasian black grouse, blackgame or blackcock, is a large Game (food), game Aves, bird in the grouse family. It is a Bird migration, sedentary species, spanning across t ...
and the famous red grouse. Common quail,
grey partridge The grey partridge (''Perdix perdix''), also known as the gray-legged partridge, English partridge, Hungarian partridge, or hun, is a gamebird in the pheasant family Phasianidae of the order Galliformes, gallinaceous birds. The scientific name ...
and
pheasant Pheasants ( ) are birds of several genera within the family (biology), family Phasianidae in the order (biology), order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera na ...
are well-distributed, although the
red-legged partridge The red-legged partridge (''Alectoris rufa'') is a gamebird in the pheasant family Phasianidae of the order Galliformes, gallinaceous birds. It is sometimes known as French partridge, to distinguish it from the English or grey partridge. The ge ...
is less so. A small colony of the introduced
golden pheasant The golden pheasant (''Chrysolophus pictus''), also known as the Chinese pheasant, and rainbow pheasant, is a gamebird of the order Galliformes (gallinaceous birds) and the family Phasianidae (pheasants). The genus name is from Ancient Greek ' ...
exists in the southwest. Among the waders,
avocet The four species of avocets are a genus, ''Recurvirostra'', of waders in the same avian family as the stilts. The genus name comes from Latin , 'curved backwards' and , 'bill'. The common name is thought to derive from the Italian ( Ferrarese) w ...
, stone-curlew,
little ringed plover The little ringed plover (''Charadrius dubius'') is a small plover. The genus name ''Charadrius'' is a Late Latin word for a yellowish bird mentioned in the fourth-century Vulgate. It derives from Ancient Greek ''kharadrios'' a bird found in river ...
and
Kentish plover The Kentish plover (''Charadrius alexandrinus'') is a small cosmopolitan shorebird (40-44 g) of the family Charadriidae that breeds on the shores of saline lakes, lagoons, and coasts, populating sand dunes, marshes, semi-arid desert, and tundra.S ...
are absent, but most of the 100 or so pairs of
dotterel The Eurasian dotterel (''Charadrius morinellus''), also known in Europe as just dotterel, is a small wader in the plover family of birds. The dotterel is a brown and black streaked bird with a broad white eye-stripe and an orange-red chest ban ...
in the UK spend their summers in Scotland as do all of the breeding Eurasian whimbrel,
greenshank The common greenshank (''Tringa nebularia'') is a wader in the large family Scolopacidae, the typical waders. The genus name ''Tringa'' is the New Latin name given to the green sandpiper by Aldrovandus in 1599 based on Ancient Greek ''trungas' ...
and
red-necked phalarope The red-necked phalarope (''Phalaropus lobatus''), also known as the northern phalarope and hyperborean phalarope, is a small wader. This phalarope breeds in the Arctic regions of North America and Eurasia. It is migratory, and, unusually for a ...
, (although the latter two species also breed in
Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...
). In summer the shallow lochs of the
machair A machair (; sometimes machar in English) is a fertile low-lying grassy plain found on part of the northwest coastlines of Ireland and Scotland, in particular the Outer Hebrides. The best examples are found on North and South Uist, Harri ...
lands in the
Uists "Uist" is a group of six islands and are part of the Outer Hebridean Archipelago, part of the Outer Hebrides of Scotland. North Uist and South Uist ( or ; gd, Uibhist ) are two of the islands and are linked by causeways running via the isles of ...
and
Benbecula Benbecula (; gd, Beinn nam Fadhla or ) is an island of the Outer Hebrides in the Atlantic Ocean off the west coast of Scotland. In the 2011 census, it had a resident population of 1,283 with a sizable percentage of Roman Catholics. It is in a ...
provide for a remarkable variety of waders and ducks including
shoveler The shovelers or shovellers are four species of dabbling ducks with long, broad spatula-shaped beaks: * Red shoveler, ''Anas platalea'' * Cape shoveler, ''Anas smithii'' * Australasian shoveler, ''Anas rhynchotis'' * Northern shoveler, ''Anas c ...
and
eider Eiders () are large seaducks in the genus ''Somateria''. The three extant species all breed in the cooler latitudes of the Northern Hemisphere. The down feathers of eider ducks, and some other ducks and geese, are used to fill pillows and quilt ...
. The rare
Slavonian grebe The horned grebe or Slavonian grebe (''Podiceps auritus'') is a relatively small waterbird in the family Podicipedidae. There are two known subspecies: ''P. a. auritus'', which breeds in the Palearctic, and ''P. a. cornutus'', which breeds i ...
and
common scoter The common scoter (''Melanitta nigra'') is a large sea duck, in length, which breeds over the far north of Europe and the Palearctic east to the Olenyok River. The genus name is derived from Ancient Greek ''melas'', "black", and ''netta'', "du ...
breed on a small number of lochs in Highland region.
Goldeneye ''GoldenEye'' is a 1995 spy film, the seventeenth in the ''James Bond'' series produced by Eon Productions, and the first to star Pierce Brosnan as the fictional MI6 agent James Bond. Directed by Martin Campbell, it was the first in the ser ...
have colonised an area centred around the
Cairngorms National Park Cairngorms National Park ( gd, Pàirc Nàiseanta a' Mhonaidh Ruaidh) is a national park in northeast Scotland, established in 2003. It was the second of two national parks established by the Scottish Parliament, after Loch Lomond and The Tro ...
since the 1970s, and about 100 pairs breed there. The majority of the roughly 25,000
whooper swan The whooper swan ( /ˈhuːpə(ɹ) swɒn/) (''Cygnus cygnus''), also known as the common swan, pronounced ''hooper swan'', is a large northern hemisphere swan. It is the Eurasian counterpart of the North American trumpeter swan, and the type specie ...
s in the British Isles winter in Scotland and About half of the 80,000
barnacle geese The barnacle goose (''Branta leucopsis'') is a species of goose that belongs to the genus ''Branta'' of black geese, which contains species with largely black plumage, distinguishing them from the grey ''Anser'' species. Despite its superficial s ...
, which breed in
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
, arrive on
Islay Islay ( ; gd, Ìle, sco, Ila) is the southernmost island of the Inner Hebrides of Scotland. Known as "The Queen of the Hebrides", it lies in Argyll just south west of Jura, Scotland, Jura and around north of the Northern Irish coast. The isl ...
for the winter, with further flocks wintering on other Scottish islands (e.g. Uists, Tiree, Colonsay) and many thousands wintering in Ireland. Tens of thousands of
pink-footed geese The pink-footed goose (''Anser brachyrhynchus'') is a goose which breeds in eastern Greenland, Iceland and Svalbard. It is migratory, wintering in northwest Europe, especially Ireland, Great Britain, the Netherlands, and western Denmark. The na ...
use the
Montrose Basin Montrose Basin is a nearly circular tidal basin which makes up part of the estuary of the River South Esk and which sits just inland of the town of Montrose in Angus on the east coast of Scotland. The basin is protected by a number of designati ...
as a winter roost in October and November as they do Loch Strathbeg and various lochs and reservoirs in Tayside and the Lothians. The amber-listed
black Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white have o ...
and red-throated diver's freshwater breeding strongholds in the British Isles are in the north and west of Scotland.


Other non-passerines

Considerable efforts have been taken to conserve the shy corncrake, and summer numbers of this red-listed species have recovered to over 1200 pairs. The
wryneck The wrynecks (genus ''Jynx'') are a small but distinctive group of small Old World woodpeckers. ''Jynx'' is from the Ancient Greek ''iunx'', the Eurasian wryneck. These birds get their English name from their ability to turn their heads almos ...
is now almost extinct in Scotland with one or two birds singing each summer, but not breeding. Of the Columbidae the turtle dove is largely absent, but in the British Isles the
rock dove The rock dove, rock pigeon, or common pigeon ( also ; ''Columba livia'') is a member of the bird family Columbidae (doves and pigeons). In common usage, it is often simply referred to as the "pigeon". The domestic pigeon (''Columba livia domes ...
is confined to the north and west coasts of Scotland and Ireland.


Passerines

Raven A raven is any of several larger-bodied bird species of the genus ''Corvus''. These species do not form a single taxonomic group within the genus. There is no consistent distinction between "crows" and "ravens", common names which are assigned t ...
s are typically forest-dwelling birds in much of Europe, but in Scotland they are generally associated with mountains and sea coasts. In 2002 the
hooded crow The hooded crow (''Corvus cornix''), also called the scald-crow or hoodie, is a Eurasian bird species in the genus ''Corvus''. Widely distributed, it is found across Northern, Eastern, and Southeastern Europe, as well as parts of the Middle Eas ...
was recognised as a separate species from the
carrion crow The carrion crow (''Corvus corone'') is a passerine bird of the family Corvidae and the genus ''Corvus'' which is native to western Europe and the eastern Palearctic. Taxonomy and systematics The carrion crow was one of the many species origi ...
. Scotland and Northern Ireland host all of the approximately 190,000 UK territories of the former. A recent survey suggest that raven numbers are increasing but that
hooded crow The hooded crow (''Corvus cornix''), also called the scald-crow or hoodie, is a Eurasian bird species in the genus ''Corvus''. Widely distributed, it is found across Northern, Eastern, and Southeastern Europe, as well as parts of the Middle Eas ...
s had declined by 59% while
carrion crow The carrion crow (''Corvus corone'') is a passerine bird of the family Corvidae and the genus ''Corvus'' which is native to western Europe and the eastern Palearctic. Taxonomy and systematics The carrion crow was one of the many species origi ...
numbers were essentially static. Concentrated on the islands of
Islay Islay ( ; gd, Ìle, sco, Ila) is the southernmost island of the Inner Hebrides of Scotland. Known as "The Queen of the Hebrides", it lies in Argyll just south west of Jura, Scotland, Jura and around north of the Northern Irish coast. The isl ...
and
Colonsay Colonsay (; gd, Colbhasa; sco, Colonsay) is an island in the Inner Hebrides of Scotland, located north of Islay and south of Mull. The ancestral home of Clan Macfie and the Colonsay branch of Clan MacNeil, it is in the council area of Argyll ...
, about 80 of Britain's 400 pairs of
red-billed chough The red-billed chough, Cornish chough or simply chough ( ; ''Pyrrhocorax pyrrhocorax''), is a bird in the crow family, one of only two species in the genus '' Pyrrhocorax''. Its eight subspecies breed on mountains and coastal cliffs from the we ...
nest in Scotland. In addition to
crossbill The crossbill is a genus, ''Loxia'', of birds in the finch family (Fringillidae), with six species. These birds are characterised by the mandibles with crossed tips, which gives the group its English name. Adult males tend to be red or orange in ...
s (see above),
crested tit The crested tit or European crested tit (''Lophophanes cristatus'') (formerly ''Parus cristatus''), is a passerine bird in the tit family Paridae. It is a widespread and common resident breeder in coniferous forests throughout central and north ...
s exist as a fragmented population of 2,400 breeding pairs in remnant patches of Caledonian Forest and in some larger plantations such as the Culbin Forest in
Moray Moray () gd, Moireibh or ') is one of the 32 local government council areas of Scotland. It lies in the north-east of the country, with a coastline on the Moray Firth, and borders the council areas of Aberdeenshire and Highland. Between 1975 ...
.
Ring ouzel The ring ouzel (''Turdus torquatus'') is a mainly European member of the thrush family Turdidae. It is a medium-sized thrush, in length and weighing . The male is predominantly black with a conspicuous white crescent across its breast. Females ...
s have declined to around 7,000 pairs, possibly due to disturbance from the growing number of human visitors to their upland habitat. There are fewer than 100 breeding pairs of
snow bunting The snow bunting (''Plectrophenax nivalis'') is a passerine bird in the family Calcariidae. It is an Arctic specialist, with a circumpolar Arctic breeding range throughout the northern hemisphere. There are small isolated populations on a few hig ...
, although in winter they are joined by migrants from continental Europe. A nest site near
Dumfries Dumfries ( ; sco, Dumfries; from gd, Dùn Phris ) is a market town and former royal burgh within the Dumfries and Galloway council area of Scotland. It is located near the mouth of the River Nith into the Solway Firth about by road from the ...
is thought to have been in use by
dipper Dippers are members of the genus ''Cinclus'' in the bird family Cinclidae, so-called because of their bobbing or dipping movements. They are unique among passerines for their ability to dive and swim underwater. Taxonomy The genus ''Cinclus'' ...
s since 1881. Scotland has 95% of the British breeding population of red-listed
twite The twite (''Linaria flavirostris'') is a small brown passerine bird in the finch family Fringillidae. It is similar in size and shape to a linnet, at long. It lacks the red head patch and breast shown by the linnet and the redpolls. It is bro ...
, about 64,000 pairs. However, a recent
RSPB The Royal Society for the Protection of Birds (RSPB) is a Charitable_organization#United_Kingdom, charitable organisation registered in Charity Commission for England and Wales, England and Wales and in Office of the Scottish Charity Regulator, ...
survey found a sudden and dramatic fall in winter numbers from 6,000 in 1998 to only 300 in 2006 in the counties of
Caithness Caithness ( gd, Gallaibh ; sco, Caitnes; non, Katanes) is a historic county, registration county and lieutenancy area of Scotland. Caithness has a land boundary with the historic county of Sutherland to the west and is otherwise bounded by ...
and
Sutherland Sutherland ( gd, Cataibh) is a historic county, registration county and lieutenancy area in the Highlands of Scotland. Its county town is Dornoch. Sutherland borders Caithness and Moray Firth to the east, Ross-shire and Cromartyshire (later ...
.


Vagrants

Scotland's position on the western seaboard of Europe means that a variety of birds not normally found in the country visit from time to time. These include accidental visits by
vagrant Vagrancy is the condition of homelessness without regular employment or income. Vagrants (also known as bums, vagabonds, rogues, tramps or drifters) usually live in poverty and support themselves by begging, scavenging, petty theft, temporar ...
birds that have wandered far from their normal habitations.
Fair Isle Fair Isle (; sco, Fair Isle; non, Friðarey; gd, Fara) is an island in Shetland, in northern Scotland. It lies about halfway between mainland Shetland and Orkney. It is known for its bird observatory and a traditional style of knitting. Th ...
is an internationally renowned site for the observation of migrant birds. Rarities have included
passerines A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by th ...
such as the thick-billed warbler,
white-throated sparrow The white-throated sparrow (''Zonotrichia albicollis'') is a passerine bird of the New World sparrow family Passerellidae. Etymology The genus name ''Zonotrichia'' is from Ancient Greek (, ) and (, ). The specific ''albicollis'' is from Lati ...
, yellow-rumped warbler and
collared flycatcher The collared flycatcher (''Ficedula albicollis'') is a small passerine bird in the Old World flycatcher family, one of the four species of Western Palearctic black-and-white flycatchers. It breeds in southeast Europe (isolated populations are p ...
. More than 345 species of bird have been recorded on this island, which measures only . Elsewhere, other rarities reported in 2006 include a white-billed diver at
Gairloch Gairloch ( ; gd, Geàrrloch , meaning "Short Loch") is a village, civil parish and community on the shores of Loch Gairloch in Wester Ross, in the North-West Highlands of Scotland. A tourist destination in the summer months, Gairloch has a go ...
, a
black-browed albatross The black-browed albatross (''Thalassarche melanophris''), also known as the black-browed mollymawk,Robertson, C. J. R. (2003) is a large seabird of the albatross family Diomedeidae; it is the most widespread and common member of its family. T ...
in the
Western Isles The Outer Hebrides () or Western Isles ( gd, Na h-Eileanan Siar or or ("islands of the strangers"); sco, Waster Isles), sometimes known as the Long Isle/Long Island ( gd, An t-Eilean Fada, links=no), is an island chain off the west coas ...
, a
laughing gull The laughing gull (''Leucophaeus atricilla'') is a medium-sized gull of North and South America. Named for its laugh-like call, it is an opportunistic omnivore and scavenger. It breeds in large colonies mostly along the Atlantic coast of North Am ...
in
Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...
and a
buff-breasted sandpiper The buff-breasted sandpiper (''Calidris subruficollis'') is a small wader, shorebird. The species name ''subruficollis'' is from Latin ''subrufus'', "reddish" (from ''sub'', "somewhat", and ''rufus'', "rufous") and ''collis'', "-necked/-throated" ...
at
Lossiemouth Lossiemouth ( gd, Inbhir Losaidh) is a town in Moray, Scotland. Originally the port belonging to Elgin, it became an important fishing town. Although there has been over 1,000 years of settlement in the area, the present day town was formed over ...
. Accidentals recorded in earlier years include an
American bittern The American bittern (''Botaurus lentiginosus'') is a species of wading bird in the heron family. It has a Nearctic distribution, breeding in Canada and the northern and central parts of the United States, and wintering in the U.S. Gulf Coast ...
in 1888 and a purple heron in the same year, a
Baikal teal The Baikal teal (''Sibirionetta formosa''), also called the bimaculate duck or squawk duck, is a dabbling duck that breeds in eastern Russia and winters in East Asia. Taxonomy The first formal description of the Baikal teal was by the German ...
in 1958, and a
black stork The black stork (''Ciconia nigra'') is a large bird in the stork family Ciconiidae. It was first described by Carl Linnaeus in the 10th edition of his ''Systema Naturae''. Measuring on average from beak tip to end of tail with a wingspan, t ...
in 1977. Birds are also presumed to have escaped from captivity, such as a
lanner falcon The lanner falcon (''Falco biarmicus'') is a medium-sized bird of prey that breeds in Africa, southeast Europe and just into Asia. It prefers open habitat and is mainly resident, but some birds disperse more widely after the breeding season. A l ...
in 1976,
Chilean flamingo The Chilean flamingo (''Phoenicopterus chilensis'') is a species of large flamingo at closely related to the American flamingo and the greater flamingo, with which it was sometimes considered conspecific. The species is listed as near threatened ...
s in 1976 and 1979, a
black-necked swan The black-necked swan (''Cygnus melancoryphus'') is a species of waterfowl in tribe Cygnini of subfamily Anserinae.HBW and BirdLife International (2021) Handbook of the Birds of the World and BirdLife International digital checklist of the bird ...
in 1988, and a
red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members with ...
in 1989. These records are but a small selection from two
counties A county is a geographic region of a country used for administrative or other purposesChambers Dictionary, L. Brookes (ed.), 2005, Chambers Harrap Publishers Ltd, Edinburgh in certain modern nations. The term is derived from the Old French ...
in the north-east and give only a flavour of the complexity and diversity of avian life in Scotland.


Extinctions

The
common crane The common crane (''Grus grus''), also known as the Eurasian crane, is a bird of the family Gruidae, the cranes. A medium-sized species, it is the only crane commonly found in Europe besides the demoiselle crane (''Grus virgo'') and the Siberi ...
and
great bittern The Eurasian bittern or great bittern (''Botaurus stellaris'') is a wading bird in the bittern subfamily (Botaurinae) of the heron family Ardeidae. There are two subspecies, the northern race (''B. s. stellaris'') breeding in parts of Europe a ...
were exterminated by hunters and the draining of marshes in the 18th century. The last
great auk The great auk (''Pinguinus impennis'') is a species of flightless alcid that became extinct in the mid-19th century. It was the only modern species in the genus ''Pinguinus''. It is not closely related to the birds now known as penguins, wh ...
seen in Britain was killed on
Stac an Armin Stac an Armin ( gd, Stac an Àrmainn), based on the proper Scottish Gaelic spelling (formerly ''àrmuinn''), is a sea stack in the St Kilda archipelago. It is 196 metres (643 ft.) tall, qualifying it as a Marilyn. It is the highest sea ...
, a rocky pinnacle in the St Kilda archipelago in July 1840.


Fish life in the sea

Of the 42 species of fish found in Scottish fresh waters, only half have arrived by natural colonisation. Native species include
allis shad The allis shad (''Alosa alosa'') is a widespread Northeast Atlantic species of fish in the herring family, Clupeidae. It is an anadromous fish which migrates into fresh water to spawn. It is found in the eastern Atlantic Ocean, the western Bal ...
,
brown trout The brown trout (''Salmo trutta'') is a European species of salmonid fish that has been widely introduced into suitable environments globally. It includes purely freshwater populations, referred to as the riverine ecotype, ''Salmo trutta'' morph ...
,
European eel The European eel (''Anguilla anguilla'') is a species of eel, a snake-like, catadromous fish. They are normally around and rarely reach more than , but can reach a length of up to in exceptional cases. Eels have been important sources of fo ...
and river lamprey. Scottish rivers support one of the largest
Atlantic salmon The Atlantic salmon (''Salmo salar'') is a species of ray-finned fish in the family Salmonidae. It is the third largest of the Salmonidae, behind Siberian taimen and Pacific Chinook salmon, growing up to a meter in length. Atlantic salmon are ...
resources in Europe, with nearly 400 rivers supporting genetically distinct populations. Five fish species are considered 'late arrivals' to Scotland, having colonised by natural means prior to 1790. They are the
northern pike The northern pike (''Esox lucius'') is a species of carnivorous fish of the genus '' Esox'' (the pikes). They are typical of brackish and fresh waters of the Northern Hemisphere (''i.e.'' holarctic in distribution). They are known simply as a ...
,
roach Roach may refer to: Animals * Cockroach, various insect species of the order Blattodea * Common roach (''Rutilus rutilus''), a fresh and brackish water fish of the family Cyprinidae ** ''Rutilus'' or roaches, a genus of fishes * California roach ...
,
stone loach The stone loach (''Barbatula barbatula'') is a European species of fresh water ray-finned fish in the family Nemacheilidae. It is one of nineteen species in the genus ''Barbatula''. Stone loaches live amongst the gravel and stones of fast flowin ...
,
European perch The European perch (''Perca fluviatilis''), also known as the common perch, redfin perch, big-scaled redfin, English perch, Euro perch, Eurasian perch, Eurasian river perch, Hatch, poor man’s rockfish or in Anglophone parts of Europe, simply th ...
, and
minnow Minnow is the common name for a number of species of small freshwater fish, belonging to several genera of the families Cyprinidae and Leuciscidae. They are also known in Ireland as pinkeens. Smaller fish in the subfamily Leusciscidae are co ...
. Rarer native species include the endemic '' Salvelinus killinensis'' and the powan, the latter found in only two locations and under threat from introduced
ruffe The Eurasian ruffe (''Gymnocephalus cernua''), also known as ruffe or pope, is a freshwater fish found in temperate regions of Europe and northern Asia. It has been introduced into the Great Lakes of North America, reportedly with unfortuna ...
and the
Arctic char The Arctic char or Arctic charr (''Salvelinus alpinus'') is a cold-water fish in the family Salmonidae, native to alpine lakes and arctic and subarctic coastal waters. Its distribution is Circumpolar North. It spawns in freshwater and populatio ...
r. The latter may have been the first fish species to re-enter fresh waters when the last ice age ended, and about 200 populations exist. The
freshwater pearl mussel The freshwater pearl mussel (''Margaritifera margaritifera'') is an endangered species of freshwater mussel, an aquatic bivalve mollusc in the family Margaritiferidae. Although the name "freshwater pearl mussel" is often used for this species ...
was once abundant enough to support commercial activities, and Scotland is the remaining European stronghold with about half the global number present. There are populations in more than 50 rivers, mainly in the
Highlands Highland is a broad term for areas of higher elevation, such as a mountain range or mountainous plateau. Highland, Highlands, or The Highlands, may also refer to: Places Albania * Dukagjin Highlands Armenia * Armenian Highlands Australia *Sou ...
, although illegal harvesting has seriously affected their survival. Scotland's seas, which constitute an area greater than that of the seas around the rest of the UK, are among the most biologically productive in the world. They are home to a third of the world's whale and dolphin species, most of the UK's maerl, (a collective term for several species of calcified red seaweed, and an important marine habitat), Horsemussel (''Modiolus modiolus'') and
seagrass Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families (Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the orde ...
beds, and distinctive species like the tall sea pen, ''Funiculina quadrangularis''. It is estimated that the total number of Scottish marine species exceeds 40,000. This includes 250 species of
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
, the most numerous inshore variety being
saithe The saithe ( or ) (''Pollachius virens'') is a species of marine fish in the Pollock genus ''Pollachius''. Together with '' P. pollachius'', it is generally referred to in the United States as pollock. Other names include the Boston blue (separa ...
, and deeper water creatures such as the dogfish,
porbeagle The porbeagle (''Lamna nasus'') is a species of mackerel shark in the family Lamnidae, distributed widely in the cold and temperate marine waters of the North Atlantic and Southern Hemisphere. In the North Pacific, its ecological equivalent is ...
and
blue shark The blue shark (''Prionace glauca''), also known as the great blue shark, is a species of requiem shark, in the family Carcharhinidae, which inhabits deep waters in the world's temperate and tropical oceans. Averaging around and preferring co ...
,
European eel The European eel (''Anguilla anguilla'') is a species of eel, a snake-like, catadromous fish. They are normally around and rarely reach more than , but can reach a length of up to in exceptional cases. Eels have been important sources of fo ...
,
sea bass Sea bass is a common name for a variety of different species of marine fish. Many fish species of various families have been called sea bass. In Ireland and the United Kingdom, the fish sold and consumed as sea bass is exclusively the European ba ...
, Atlantic halibut and various
rays Ray may refer to: Fish * Ray (fish), any cartilaginous fish of the superorder Batoidea * Ray (fish fin anatomy), a bony or horny spine on a fin Science and mathematics * Ray (geometry), half of a line proceeding from an initial point * Ray (gra ...
. There are four species of
sea turtle Sea turtles (superfamily Chelonioidea), sometimes called marine turtles, are reptiles of the order Testudines and of the suborder Cryptodira. The seven existing species of sea turtles are the flatback, green, hawksbill, leatherback, loggerhead, ...
, the
leatherback The leatherback sea turtle (''Dermochelys coriacea''), sometimes called the lute turtle or leathery turtle or simply the luth, is the largest of all living turtles and the heaviest non-crocodilian reptile, reaching lengths of up to and weights ...
, loggerhead,
Kemp's ridley Kemp's ridley sea turtle (''Lepidochelys kempii''), also called the Atlantic ridley sea turtle, is the rarest species of sea turtle and is the world's most endangered species of sea turtle. It is one of two living species in the genus '' Lepido ...
and
green turtle The green sea turtle (''Chelonia mydas''), also known as the green turtle, black (sea) turtle or Pacific green turtle, is a species of large sea turtle of the Family (biology), family Cheloniidae. It is the only species In biology, a spec ...
. Scottish waters contain around 2,500
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
species and 700
mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
s and in 2012 a bed of 100 million flame shells was found during a survey of
Loch Alsh Loch Alsh (from the Scottish Gaelic ''Loch Aillse'', "foaming lake") is a sea inlet between the isle of Skye in the Inner Hebrides and the Northwest Highlands of Scotland. The name is also used to describe the surrounding country and the feuda ...
. The
Darwin Mounds Darwin Mounds is a large field of undersea sand mounds situated off the north west coast of Scotland that were first discovered in May 1998. They provide a unique habitat for ancient deep water coral reefs and were found using remote sensing tech ...
, an important area of cold water
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. Co ...
s discovered in 1988, are about deep in the
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
, about north-west of
Cape Wrath Cape Wrath ( gd, Am Parbh, known as ' in Lewis) is a cape in the Durness parish of the county of Sutherland in the Highlands of Scotland. It is the most north-westerly point in mainland Britain. The cape is separated from the rest of the mai ...
in the north-east corner of the
Rockall Trough The Rockall Trough ( gd, Clais Sgeir Rocail) is a deep-water bathymetric feature to the northwest of Scotland and Ireland, running roughly from southwest to northeast, flanked on the north by the Rockall Plateau and to the south by the Porc ...
. The area covers approximately and contains hundreds of mounds of about in diameter and in height, many having a teardrop shaped 'tail' orientated south-west of the mound. This feature may be unique globally. The tops of the mounds have living stands of ''Lophelia'' corals and support significant populations of the single-celled ''
Syringammina fragilissima ''Syringammina'' is a xenophyophore found off the coast of Scotland, near Rockall. It is one of the largest single-celled organisms known, at up to across. It was first described in 1882 by the oceanographer John Murray, after being discovered ...
''. Fish have been observed in the vicinity but not at higher densities than the background environment. Damage from trawler fishing was visible over about a half of the eastern Darwin Mounds surveyed during summer 2000, and the UK government is taking steps to protect the area. In 2003 the
European Commission The European Commission (EC) is the executive of the European Union (EU). It operates as a cabinet government, with 27 members of the Commission (informally known as "Commissioners") headed by a President. It includes an administrative body o ...
provided emergency protection and banned damaging fishing activity in the locality. Further action on a much wider scale may be required. According to a recent report "Scotland's marine life could be almost wiped out within 50 years unless tough action is taken to manage the way humans use the seas". Fears were expressed by a consortium of environmental organisations that commercial fish stocks, including
Atlantic cod The Atlantic cod (''Gadus morhua'') is a benthopelagic fish of the family Gadidae, widely consumed by humans. It is also commercially known as cod or codling.Firth of Clyde The Firth of Clyde is the mouth of the River Clyde. It is located on the west coast of Scotland and constitutes the deepest coastal waters in the British Isles (it is 164 metres deep at its deepest). The firth is sheltered from the Atlantic ...
is affecting all forms of marine life and that the growth in off-shore tourism was deleterious to populations of, for example,
basking shark The basking shark (''Cetorhinus maximus'') is the second-largest living shark and fish, after the whale shark, and one of three plankton-eating shark species, along with the whale shark and megamouth shark. Adults typically reach in leng ...
. A call was made for a 'Scottish Marine Bill' to co-ordinate and manage human activity at sea and to provide more protected areas such as marine
national park A national park is a nature park, natural park in use for conservation (ethic), conservation purposes, created and protected by national governments. Often it is a reserve of natural, semi-natural, or developed land that a sovereign state dec ...
s. The
Marine (Scotland) Act 2010 On 10 March 2010, Scotland's Marine Bill received Royal Assent, making it thMarine (Scotland) Act 2010 The Marine (Scotland) Act is an Act of the Scottish Parliament which provides a framework which will help balance competing demands on Scotlan ...
was subsequently passed by the Scottish Parliament. '' Calyptraea chinensis'' (L.) is a
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
that has invaded the shores of Scotland and by 1998 had reached nearly as far north as Oban. One living specimen was found at Clachan Sound, and earlier records showed findings of gastropod shells.Smith, S. 1998. ''Calyptraea chinensis'' (L., 1758) (Mollusca: Gastropoda). ''Newsletter PMNHS'' No. 1. p.10.


Riverine extinctions

Pollution and predation led to the extinction of both species of vendace from its very restricted range in south-western Scottish freshwaters in 1980. In the 1990s a successful attempt to reintroduce '' Coregonus vandesius'' to the
Lochmaben Lochmaben ( Gaelic: ''Loch Mhabain'') is a small town and civil parish in Scotland, and site of a castle. It lies west of Lockerbie, in Dumfries and Galloway. By the 12th century the Bruce family had become the local landowners and, in the 14th ...
area began. ''
Coregonus albula ''Coregonus albula'', known as the vendace or as the European cisco, is a species of freshwater whitefish in the family Salmonidae. It is found in lakes in northern Europe, especially Finland, Latvia, Lithuania, Sweden, Russia and Estonia, and in ...
'' remains absent. ''
Salvelinus inframundus ''Salvelinus inframundus'', also known as Orkney charr is a cold-water fish in the family Salmonidae which is endemic to Scotland. Description ''Salvelinus inframundus'' has the following characteristics which in combination make this taxon dif ...
'', a rare Char (fish), char species that could be vulnerable species, vulnerable to extinct, has been found in Loch Mealt, Isle of Skye, Scottish Highlands.


Amphibians and land reptiles

Only six amphibians and four land reptiles are wiktionary:native, native to Scotland. The amphibians include three species of newt: the great crested newt, great crested, of which fewer than 1,000 individuals survive; the smooth newt, smooth, and the palmate newt, palmate. The other amphibians are the common toad, the natterjack toad, found in only four locations in the south-west, and the common frog. A single alien amphibian is known in Scotland, the Alpine newt, a recent escapee confined to the Edinburgh area. The reptiles include the Vipera berus, adder and the grass snake, the Anguis fragilis, slowworm, which is a legless lizard, and the viviparous lizard, common lizard. Coronella austriaca, Smooth snakes, found elsewhere in the UK are absent, and grass snakes are rarely reported.


Terrestrial invertebrates

Seventy-seven species of land snail and an estimated 14,000 species of insect live in Scotland, none of them "truly" endemic.Buckland, Paul C. and Sadler, Jon P. "Insects" in Edwards and Ralston (2003) pp. 105–08. These include ''Pardosa lugubris'', a species of wolf spider first found in the UK in 2000 at Abernethy Forest nature reserve, and the Formica aquilonia, Scottish wood ant. These ants, which are the most numerous residents of the Caledonian pine forest, build mounds from the pine cones and needles they find on the forest floor and may inhabit the mounds for decades. A single colony may collect 100,000 insects a day to feed its half million citizens and produce up to of Honeydew (secretion), honeydew per season. In addition to the Scottish wood ant, several Scottish species of invertebrate exist that are otherwise rare in the UK and important enough to have a specific "Action Plan" to provide protection. These are five species of ant and bee, six moths and butterfly, five fly, flies and a single beetle (the reed beetle) and snail (the Vertigo genesii, round-mouthed whorl snail, ''Vertigo genesii''). Northern colletes is a rare species of bee, the most significant British habitat for which is in the
Outer Hebrides The Outer Hebrides () or Western Isles ( gd, Na h-Eileanan Siar or or ("islands of the strangers"); sco, Waster Isles), sometimes known as the Long Isle/Long Island ( gd, An t-Eilean Fada, links=no), is an island chain off the west coast ...
, where there are more than ten colonies.Ross, David (13 September 2007 ) "Rare species of burrowing bee has flourished in Outer Hebrides". Glasgow. ''The Herald''. Scotland is also the UK stronghold of the bumblebee, Blaeberry bumblebee, and the Bumblebee Conservation Trust recently created the world's first sanctuary for this genus of insects at
RSPB The Royal Society for the Protection of Birds (RSPB) is a Charitable_organization#United_Kingdom, charitable organisation registered in Charity Commission for England and Wales, England and Wales and in Office of the Scottish Charity Regulator, ...
Vane Farm Nature Reserve near Loch Leven (Kinross), Loch Leven. The bumblebee Bombus jonellus, ''Bombus jonellus'' var. ''hebridensis'' is endemic to the Hebrides. In 2010, a colony of the beetle ''Meloe brevicollis'' was found on the island of Coll. The species is otherwise extinct in Scotland and is also flightless, raising the question of how the colony arrived on the island. The Northern February red, northern February red stonefly (''Brachyptera putata'') has recently lost its range elsewhere in Britain and is now it considered to be a Scottish endemic. Although many species of butterfly are in decline in the UK, recent research suggests that some, such as the pearl-bordered fritillary, marsh fritillary and Carterocephalus palaemon, chequered skipper, which are becoming rare in the rest of the UK, are moving north into Scotland in response to climate change. In June 2008 an adult ''Ethmia pyrausta'' moth was discovered in Easter Ross. This find was only the fifth sighting since its discovery in the UK at Loch Shin in 1853, and the species has gained "almost mythical status" according to Butterfly Conservation Scotland. The most well-known invertebrate may be a species of midge (''Highland midge, Culicoides impunctatus''), a tiny flying gnat that is the scourge of summer visitors and residents alike. Its predations result in the loss of up to 20% of summer working days in the forestry industry. Others of significance include the pine weevil, black pine beetle, Clytrinae, clytra beetle, and the timberman beetle, timberman, a long-horned beetle. The archaeological site at
Skara Brae Skara Brae is a stone-built Neolithic settlement, located on the Bay of Skaill on the west coast of Mainland, the largest island in the Orkney archipelago of Scotland. Consisting of ten clustered houses, made of flagstones, in earthen dams ...
provided the earliest known record of the human flea, ''flea, Pulex irritans'' in Europe. The islands of
Colonsay Colonsay (; gd, Colbhasa; sco, Colonsay) is an island in the Inner Hebrides of Scotland, located north of Islay and south of Mull. The ancestral home of Clan Macfie and the Colonsay branch of Clan MacNeil, it is in the council area of Argyll ...
and Oronsay, Inner Hebrides, Oronsay are home to about 50 colonies of the only native species of honeybee in Britain–''Apis mellifera mellifera''. The Scottish Government introduced the Bee Keeping (Colonsay and Oronsay) Order 2013 to protect the species from cross-breeding and disease as the species has suffered serious declines on the mainland.


Cryptozoology

A variety of exotic cats are rumoured to exist, including the 'Beast of Buchan'. The 'Kellas Cat' of
Moray Moray () gd, Moireibh or ') is one of the 32 local government council areas of Scotland. It lies in the north-east of the country, with a coastline on the Moray Firth, and borders the council areas of Aberdeenshire and Highland. Between 1975 ...
is a jet black, long-legged animal, and is probably the result of a modern wild cat/domestic cat Hybrid (biology), hybrid, or a melanistic wild cat. In earlier times it may have spawned the legend of the ''Cat Sidhe'' or "Fairy Cat". The fabulous Loch Ness Monster, possibly a form of "Each Uisge, water horse", has a long history; the first recorded sighting allegedly took place in 565 AD. More recently, the Stronsay Beast was an unidentified cryptid washed ashore in the Orkney islands in the 19th century.


Nature conservation in Scotland


Challenges

Conservation of the natural environment is well-developed in the United Kingdom. The resources of the organisations concerned may be insufficient to the challenge, but the contrast with earlier attitudes about the environment is striking. In Victorian times few animals became extinct in Scotland, but the scale of the slaughter on hunting estates was staggering. Richard Perry records that on a single estate in the Cairngorms between 1837 and 1840 the following "vermin" were exterminated by keepers purely in the interests of preserving the red grouse, grouse population:
246 Martens, 198 Wild Cats, 106 Polecats, 67 Badgers, 58 Otters, 475 Ravens, 462 Kestrels, 371 Rough-legged Buzzards, 285 Common Buzzards, 275 Kites, 98 Peregrine Falcons, 92 Hen Harriers, 78 Merlins, 71 Short-eared Owls, 63 Goshawks, 35 Long-eared Owls, 27 Sea Eagles, 18 Ospreys, 15 Golden Eagles, 11 Hobbys, 6 Gyrfalcons, 5 Marsh Harriers, 3 Honey Buzzards,
and for reasons apparently unconnected to grouse shooting, a further
11 Foxes, 301 Stoats and Weasels, 78 House Cats, 1,431 Hooded or Carrion Crow, 3 Barn Owls, 8 Magpies and 7 "Orange-legged Falcons".
Writing in 1947, Perry stated that his "first reaction to this dreadful black-list was that of amazed incredulity. I still find the details incredible. However, they were supplied by the lessee himself." In several instances these extermination totals are larger than the current resident numbers for the entire country. It remains to be seen if the destruction wrought by the Victorians continues to be the nadir for the fauna of Scotland. In addition to other difficulties the marine environment faces, climate change is a challenge facing all of Scotland's habitats. Among the birds,
ptarmigan ''Lagopus'' is a small genus of birds in the grouse subfamily commonly known as ptarmigans (). The genus contains three living species with numerous described subspecies, all living in tundra or cold upland areas. Taxonomy and etymology The ge ...
,
dotterel The Eurasian dotterel (''Charadrius morinellus''), also known in Europe as just dotterel, is a small wader in the plover family of birds. The dotterel is a brown and black streaked bird with a broad white eye-stripe and an orange-red chest ban ...
and
snow bunting The snow bunting (''Plectrophenax nivalis'') is a passerine bird in the family Calcariidae. It is an Arctic specialist, with a circumpolar Arctic breeding range throughout the northern hemisphere. There are small isolated populations on a few hig ...
in particular may be affected as they depend on high-altitude habitats, and populations are likely to decline if warmer weather brings competitors into their restricted ranges. Mammals and other vertebrates may fare better, although localised invertebrate populations are at risk. Marine life is already being affected. Planktonic species that prefer cold water are declining and are not able support the crucial food chains on which many seabirds depend. Further evidence of problems for marine species has been provided by the St Andrews University Sea Mammal Research Unit. An analysis suggests that
common seal The harbor (or harbour) seal (''Phoca vitulina''), also known as the common seal, is a true seal found along temperate and Arctic marine coastlines of the Northern Hemisphere. The most widely distributed species of pinnipeds, pinniped (walruses ...
populations in
Orkney Orkney (; sco, Orkney; on, Orkneyjar; nrn, Orknøjar), also known as the Orkney Islands, is an archipelago in the Northern Isles of Scotland, situated off the north coast of the island of Great Britain. Orkney is 10 miles (16 km) north ...
and
Shetland Shetland, also called the Shetland Islands and formerly Zetland, is a subarctic archipelago in Scotland lying between Orkney, the Faroe Islands and Norway. It is the northernmost region of the United Kingdom. The islands lie about to the no ...
fell by 40% from 2001 to 2006, prompting the then Scottish Executive to announce the likelihood of a new protective conservation order. The complexities involved in conserving Scottish wildlife are highlighted in an RSPB report, noting that pine martens have been found to be a significant predator of capercaillie nests. Both species are protected, providing conservation agencies with a challenging conundrum to address. In 2012 the Scottish Government published a "Code of Practice on Non-Native Species" to help people understand their responsibilities and provide guidance as to which public body has responsibility for the various habitats involved.


Conservation organisations

Various public sector organisations have an important role in the stewardship of the country's fauna.
Scottish Natural Heritage NatureScot ( gd, NàdarAlba), which was formerly known as Scottish Natural Heritage, is an executive non-departmental public body of the Scottish Government responsible for the country's natural heritage, especially its natural, genetic and ...
is the statutory body responsible for natural heritage management in Scotland. One of its duties is to establish national nature reserve (Scotland), national nature reserves (NNR)s. Until 2004 there were 73, but a review carried out in that year resulted in a significant number of sites losing their NNR status, and by 2006 there were 55. there are 43. Forestry and Land Scotland serves as the forestry department of the Scottish Government and is one of the country's largest landowners. The Joint Nature Conservation Committee is the statutory adviser to Government on UK and international nature conservation. The country has two
national park A national park is a nature park, natural park in use for conservation (ethic), conservation purposes, created and protected by national governments. Often it is a reserve of natural, semi-natural, or developed land that a sovereign state dec ...
s.
Cairngorms National Park Cairngorms National Park ( gd, Pàirc Nàiseanta a' Mhonaidh Ruaidh) is a national park in northeast Scotland, established in 2003. It was the second of two national parks established by the Scottish Parliament, after Loch Lomond and The Tro ...
includes the largest area of arctic mountain landscape in the UK. Sites designated as of importance to natural heritage take up 39% of the land area, two-thirds of which are of Europe-wide importance. Loch Lomond and the Trossachs National Park includes Britain's largest body of fresh water, the mountains of Breadalbane, Scotland, Breadalbane and the sea lochs of Argyll. Charitable and voluntary organisations also have important roles to play. The National Trust for Scotland is the conservation charity that protects and promotes Scotland's natural and cultural heritage. With more than 270,000 members it is the largest conservation charity in Scotland. The
Scottish Wildlife Trust The Scottish Wildlife Trust is a Office of the Scottish Charity Regulator, registered charity dedicated to conserving the Natural history of Scotland, wildlife and natural environment of Scotland. Description The Scottish Wildlife Trust has well ...
is a leading voluntary conservation organisation, working to protect Scotland's natural environment. The
Royal Zoological Society of Scotland The Royal Zoological Society of Scotland is a learned society and registered charity based in Edinburgh, Scotland. It was founded by Thomas Haining Gillespie in 1909. In 1913, Edinburgh Town Council bought a large plot of land on Corstorphin ...
is a learned society and registered charity that maintains Edinburgh Zoo and the Highland Wildlife Park (a safari park and zoo near Kingussie, which specialises in native fauna). The Society is also involved in various conservation programs around Scotland and the world. The
Royal Society for the Protection of Birds The Royal Society for the Protection of Birds (RSPB) is a charitable organisation registered in England and Wales and in Scotland. It was founded in 1889. It works to promote conservation and protection of birds and the wider environment throug ...
promotes conservation of birds and other wildlife through the protection and re-creation of habitats. The John Muir Trust is a charity whose main role is as a guardian of wild land and wildlife, through the ownership of land and the promotion of education and Conservation biology, conservation. The trust owns and manages estates in locations including Knoydart and Assynt, and on the isle of
Skye The Isle of Skye, or simply Skye (; gd, An t-Eilean Sgitheanach or ; sco, Isle o Skye), is the largest and northernmost of the major islands in the Inner Hebrides of Scotland. The island's peninsulas radiate from a mountainous hub dominated b ...
. It has links with the Sierra Club in the United States, which also celebrates the legacy of Dunbar-born John Muir. Trees for Life (Scotland), Trees for Life is a charity that aims to restore a "wild forest" in the Northwest Highlands and Grampian Mountains (Scotland), Grampian Mountains.


See also

*British avifauna *Climate of Scotland *Flora of Scotland *Geography of Scotland *Geology of Scotland *Nature of the Outer Hebrides *British Amphibians, List of British amphibians *List of birds of Great Britain, List of British birds *List of British butterflies *List of British mammals *List of British reptiles *List of extinct animals of Britain *List of fauna of the Scottish Highlands *Lists of insects recorded in Britain *List of moths of Great Britain (Arctiidae) *List of moths of Great Britain (Geometridae) *National nature reserve (Scotland) *:Royal Society for the Protection of Birds reserves in Scotland, Royal Society for the Protection of Birds reserves in Scotland


References


Cited texts

*Benvie, Neil (2004) ''Scotland's Wildlife''. London. Aurum Press. *Brown, Leslie (1989) ''British Birds of Prey''. London. Bloomsbury. *Corbet, Gordon and Ovenden, Denys (1984) ''The Mammals of Britain and Europe''. Glasgow. Collins. * Cook, Martin (1992). ''The Birds of Moray and Nairn''. Edinburgh: Mercat Press. *Edwards, Kevin J. & Ralston, Ian B.M. (Eds) (2003) ''Scotland After the Ice Age: Environment, Archaeology and History, 8000 BC - AD 1000''. Edinburgh. Edinburgh University Press. *Fraser Darling, F. & Boyd, J.M. (1969) ''Natural History in the Highlands and Islands.'' London. Bloomsbury. *Gooders, J. (1994) ''Field Guide to the Birds of Britain and Ireland''. London. Kingfisher. *Haswell-Smith, Hamish (2004) ''The Scottish Islands''. Edinburgh. Canongate. *Hull, Robin (2007) ''Scottish Mammals''. Edinburgh. Birlinn. *MacLean, Charles (1972) ''Island on the Edge of the World: the Story of St. Kilda''. Edinburgh. Canongate. *Matthews, L. Harrison (1968) ''British Mammals.'' London. Bloomsbury. *Miles, H. and Jackman, B. (1991) ''The Great Wood of Caledon''. Lanark. Colin Baxter Photography. *W.H. Murray, Murray, W. H. (1973) ''The Islands of Western Scotland: the Inner and Outer Hebrides.'' London. Eyre Methuen. SBN 413303802 *Perry, Richard (1948). ''In The High Grampians''. London. Lindsay Drummond. *Peterson, Roger Tory; Mountfort, Guy; and Hollom, P.A.D. (1993) ''Birds of Britain and Europe''. Glasgow. HarperCollins. *Quine, David (2000). ''St Kilda''. Grantown-on-Spey. Colin Baxter Island Guides. *Smout, T.C. MacDonald, R. and Watson, Fiona (2007) ''A History of the Native Woodlands of Scotland 1500–1920''. Edinburgh University Press. * Thompson, Francis (1968) ''Harris and Lewis, Outer Hebrides''. Newton Abbot. David & Charles. *


External links


Scottish Natural HeritageForestry Commission Scotland

Joint Nature Conservation CommitteeScottish Wildlife TrustRoyal Zoological Society of ScotlandScotland's National Nature ReservesRoyal Society for the Protection of BirdsScottish Seabird CentreCairngorms National Park AuthorityLoch Lomond & The Trossachs National Park AuthorityJohn Muir TrustTrees for LifeScottish Wildcat Association
{{DEFAULTSORT:Fauna Of Scotland Fauna of Scotland, Nature conservation in Scotland Natural history of Scotland Fauna of Great Britain, *