The fulmars are tubenosed seabirds of the
family
Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Procellariidae
The family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prions, and the shearwaters. This family is part of the bird order Procellariiformes (or tubenoses), which also ...
. The family consists of two
extant
Extant is the opposite of the word extinct. It may refer to:
* Extant hereditary titles
* Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English
* Extant taxon, a taxon which is not extinct, ...
species and two extinct fossil species from the
Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
.
Fulmars superficially resemble gulls, but are readily distinguished by their flight on stiff wings, and their tube noses. They breed on
cliffs
In geography and geology, a cliff is an area of rock which has a general angle defined by the vertical, or nearly vertical. Cliffs are formed by the processes of weathering and erosion, with the effect of gravity. Cliffs are common on co ...
, laying one or rarely two eggs on a ledge of bare rock or on a grassy cliff. Outside the breeding season, they are
pelagic
The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...
, feeding on fish,
squid
True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ...
and
shrimp
Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
in the open ocean. They are long-lived for birds, living for up to 40 years.
Historically, the northern fulmar lived on the
Isle of St Kilda
St Kilda ( gd, Hiort) is an isolated archipelago situated west-northwest of North Uist in the North Atlantic Ocean. It contains the westernmost islands of the Outer Hebrides of Scotland. The largest island is Hirta, whose sea cliffs are the ...
, where it was extensively hunted. The species has expanded its breeding range southwards to the coasts of England and northern France.
Taxonomy
The genus ''Fulmarus'' was introduced in 1826 by the English naturalist
James Stephens. The name comes from the
Old Norse
Old Norse, Old Nordic, or Old Scandinavian, is a stage of development of North Germanic languages, North Germanic dialects before their final divergence into separate Nordic languages. Old Norse was spoken by inhabitants of Scandinavia and t ...
''Fúlmár'' meaning "foul-mew" or "foul-gull" because of the birds' habit of ejecting a foul-smelling oil. The
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
was designated by
George Gray in 1855 as the
northern fulmar
The northern fulmar (''Fulmarus glacialis''), fulmar, or Arctic fulmar is a highly abundant seabird found primarily in subarctic regions of the North Atlantic and North Pacific oceans. There has been one confirmed sighting in the Southern Hem ...
.
As members of
Procellaridae and then the
order Procellariiformes
Procellariiformes is an order of seabirds that comprises four families: the albatrosses, the petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still called tubenoses in English, procellariiforms are of ...
, they share certain traits. First, they have nasal passages that attach to the upper bill called
naricorns
A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbi ...
. The bills of Procellariiformes are unique in being split into between seven and nine horny plates. Finally, they produce a
stomach oil
Stomach oil is the light oil composed of neutral dietary lipids found in the proventriculus (fore-gut) of birds in the order Procellariiformes. All albatrosses, procellarids (gadfly petrels and shearwaters) and northern and austral storm petrel ...
made up of
wax ester
A wax ester (WE) is an ester of a fatty acid and a fatty alcohol. Wax esters comprise the main components of three commercially important waxes: carnauba wax, candelilla wax, and beeswax..
Wax esters are formed by combining one fatty acid with o ...
s and
triglycerides
A triglyceride (TG, triacylglycerol, TAG, or triacylglyceride) is an ester derived from glycerol and three fatty acids (from ''tri-'' and ''glyceride'').
Triglycerides are the main constituents of body fat in humans and other vertebrates, as we ...
that is stored in the
proventriculus The proventriculus is part of the digestive system of birds.Encarta World English Dictionary orth American Edition(2007). ''Proventriculus''. Source: (accessed: December 18, 2007) An analogous organ exists in invertebrates and insects.
Birds
Th ...
. This can be sprayed out of their mouths as a defence against predators and as an energy rich food source for chicks and for the adults during their long flights. It will mat the
plumage
Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
of avian predators, which can lead to their death. Fulmars have a
salt gland
The salt gland is an organ for excreting excess salts. It is found in the cartilaginous fishes subclass elasmobranchii (sharks, rays, and skates), seabirds, and some reptiles. Salt glands can be found in the rectum of sharks. Birds and reptiles ...
that is situated above the nasal passage and helps desalinate their bodies, due to the high amount of ocean water that they imbibe. It excretes a strong saline solution from their nose.
Extant species
The genus contains the following two species.
Fossils
Two prehistoric species have been described from
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
bones found on the
Pacific
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
coast of
California
California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
: ''Fulmarus miocaenus'' (
Temblor Formation
The Temblor Formation is a geologic formation in California. It preserves fossils dating back from the Late Oligocene to the Middle Miocene of the Neogene period. It is notable for the famous Sharktooth Hill deposit (otherwise known as Ernst Q ...
) and ''Fulmarus hammeri'' from the
Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
.
Description
The two fulmars are closely related
seabird
Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...
s occupying the same niche in different
oceans
The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the worl ...
. The
northern fulmar
The northern fulmar (''Fulmarus glacialis''), fulmar, or Arctic fulmar is a highly abundant seabird found primarily in subarctic regions of the North Atlantic and North Pacific oceans. There has been one confirmed sighting in the Southern Hem ...
(''Fulmarus glacialis'') or just fulmar lives in the North
Atlantic
The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...
and North
Pacific
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
, whereas the
southern fulmar
The southern fulmar (''Fulmarus glacialoides'') is a seabird of the Southern Hemisphere. Along with the northern fulmar, ''F. glacialis'', it belongs to the fulmar genus ''Fulmarus'' in the family Procellariidae, the true petrels. It is also kno ...
, (''Fulmarus glacialoides'') is, as its name implies, a bird of the
Southern Ocean
The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the World Ocean, generally taken to be south of 60° S latitude and encircling Antarctica. With a size of , it is regarded as the second-small ...
. These
birds
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
look superficially like
gull
Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century, m ...
s, but are not closely related, and are in fact
petrel
Petrels are tube-nosed seabirds in the bird order Procellariiformes.
Description
The common name does not indicate relationship beyond that point, as "petrels" occur in three of the four families within that group (all except the albatross f ...
s. The northern
species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
is grey and white with a yellow bill, in length with a wingspan.
[Maynard, B. J. (2003)] The southern form is a paler bird with dark wing tips, long, with a wingspan.
Behavior
Breeding
![Fulmar Yesnaby 2010](https://upload.wikimedia.org/wikipedia/commons/c/c8/Fulmar_Yesnaby_2010.jpg)
Both recent species breed on
cliffs
In geography and geology, a cliff is an area of rock which has a general angle defined by the vertical, or nearly vertical. Cliffs are formed by the processes of weathering and erosion, with the effect of gravity. Cliffs are common on co ...
, laying a single white
egg
An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the a ...
.
Unlike many small to medium birds in the
Procellariiformes
Procellariiformes is an order of seabirds that comprises four families: the albatrosses, the petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still called tubenoses in English, procellariiforms are of ...
, they are neither
nocturnal
Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite.
Nocturnal creatures generally have highly developed sens ...
breeders, nor do they use
burrow
An Eastern chipmunk at the entrance of its burrow
A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of sh ...
s; their eggs are laid on the bare rock or in shallow depressions lined with plant material.
In Britain, northern fulmars historically bred on
St. Kilda (where their harvesting for oil, feathers and meat was central to the islands' economy). They spread into northern Scotland in the 19th century, and to the rest of the
United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
by 1930. The expansion has continued further south; the fulmar can now often be seen in the English Channel and in France along the northern and western coasts, with breeding pairs or small colonies in Nord,
Picardy
Picardy (; Picard and french: Picardie, , ) is a historical territory and a former administrative region of France. Since 1 January 2016, it has been part of the new region of Hauts-de-France. It is located in the northern part of France.
Hi ...
,
Normandy
Normandy (; french: link=no, Normandie ; nrf, Normaundie, Nouormandie ; from Old French , plural of ''Normant'', originally from the word for "northman" in several Scandinavian languages) is a geographical and cultural region in Northwestern ...
and along the Atlantic coast in
Brittany
Brittany (; french: link=no, Bretagne ; br, Breizh, or ; Gallo language, Gallo: ''Bertaèyn'' ) is a peninsula, Historical region, historical country and cultural area in the west of modern France, covering the western part of what was known ...
.
[Yeatman, L (1976)]
Feeding
![Bird's Eggs from Sea-Cliffs tail-piece in Bewick British Birds 1804](https://upload.wikimedia.org/wikipedia/commons/1/1f/Bird%27s_Eggs_from_Sea-Cliffs_tail-piece_in_Bewick_British_Birds_1804.jpg)
Fulmars are highly
pelagic
The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or wa ...
outside the breeding season, like most
tubenoses, feeding on
fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
, small
squid
True squid are molluscs with an elongated soft body, large eyes, eight arms, and two tentacles in the superorder Decapodiformes, though many other molluscs within the broader Neocoleoidea are also called squid despite not strictly fitting t ...
,
shrimp
Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are refer ...
,
crustacean
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s,
marine worm
Any worm that lives in a marine environment is considered a marine worm. Marine worms are found in several different phyla, including the Platyhelminthes, Nematoda, Annelida (segmented worms), Chaetognatha, Hemichordata, and Phoronida. For a ...
s, and
carrion
Carrion () is the decaying flesh of dead animals, including human flesh.
Overview
Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
. The range of these species increased greatly in the 20th century due to the availability of fish offal from commercial fleets, but may contract because of less food from this source and climatic change.
The population increase has been especially notable in the
British Isles
The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
.
[Bull, J. & Farrand Jr., J. (1993)]
Like other
petrels
Petrels are tube-nosed seabirds in the bird order Procellariiformes.
Description
The common name does not indicate relationship beyond that point, as "petrels" occur in three of the four families within that group (all except the albatross f ...
, their walking ability is limited, but they are strong fliers, with a stiff
wing
A wing is a type of fin that produces lift while moving through air or some other fluid. Accordingly, wings have streamlined cross-sections that are subject to aerodynamic forces and act as airfoils. A wing's aerodynamic efficiency is expres ...
action quite unlike the
gulls
Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari (bird), Lari. They are most closely related to the terns and Skimmer (bird), skimmers and only distantly related to auks, and even more distantly to waders ...
. They look bull-necked compared to gulls, and have short stubby bills. They are long-lived, the longest recorded lifespan for ''F. glacialis'' being 40 years, 10 months and 16 days.
Relationship with humans
Fulmars have for centuries been hunted for food. The engraver
Thomas Bewick
Thomas Bewick (c. 11 August 17538 November 1828) was an English wood-engraver and natural history author. Early in his career he took on all kinds of work such as engraving cutlery, making the wood blocks for advertisements, and illustrating ch ...
wrote in 1804 that "
Pennant, speaking of those
irdswhich breed on, or inhabit, the
Isle of St Kilda
St Kilda ( gd, Hiort) is an isolated archipelago situated west-northwest of North Uist in the North Atlantic Ocean. It contains the westernmost islands of the Outer Hebrides of Scotland. The largest island is Hirta, whose sea cliffs are the ...
, says—'No bird is of so much use to the islanders as this: the Fulmar supplies them with oil for their lamps, down for their beds, a delicacy for their tables, a balm for their wounds, and a medicine for their distempers. A photograph by
George Washington Wilson taken about 1886 shows a "view of the men and women of St Kilda on the beach dividing up the catch of Fulmar".
James Fisher, author of ''The Fulmar'' (1952) calculated that every person on St Kilda consumed over 100 fulmars each year; the meat was their staple food, and they caught around 12,000 birds annually.
Fulmar eggs were collected until the late 1920s in the
St Kilda islands by their men scaling the cliffs. The eggs were buried in St Kilda
peat
Peat (), also known as turf (), is an accumulation of partially decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, moors, or muskegs. The peatland ecosystem covers and is the most efficien ...
ash to be eaten through the cold, northern winters. The eggs were considered to taste like
duck eggs
Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especia ...
in taste and nourishment.
However, when the human population left St Kilda in 1930, the fulmar population did not suddenly increase.
Both the southern fulmar and the northern fulmar are listed as of
Least Concern
A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
by the
International Union for Conservation of Nature
The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN).
Gallery
File:Northern Fulmar, Finstown, Orkney.JPG, Northern fulmar on the nest in Orkney, Scotland
File:Southern Fulmar closeup.jpg, Southern fulmar in Drake's Passage.
File:Fulmarus glacialis 1 8.jpg, Northern fulmar, breeding on Bear Island (Norway)
Bear Island ( no, Bjørnøya, ) is the southernmost island of the Norway, Norwegian Svalbard, Svalbard archipelago. The island is located in the western part of the Barents Sea, approximately halfway between Spitsbergen and the North Cape, Norwa ...
File:Northern Fulmar (Fulmarus glacialis) flying, side-on.jpg, Northern fulmar in flight over Faxaflói
Faxaflói (sometimes Faxa Bay, Faxe Bay or Faxi Bay,Thorstein Thorsteinsson. 1930. ''Iceland, 1930: A Handbook Published on the Fortieth Anniversary of Landsbanki Íslands (National Bank of Iceland)''. Reykjavík: Ríkisprentsmidjan Gutenberg. i ...
(Iceland
Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its s ...
)
File:Fulmarus glacialis 1 2.jpg, Northern fulmar, breeding on Bear Island
File:Eissturmvogel02.jpg, Northern fulmar, at the Norwegian bird-island Runde
File:Northern Fulmar from the Crossley ID Guide Britain and Ireland.jpg, Composite image of northern fulmars in different plumages
References
Sources
*
*
*
*
* See also more recent publication(s) with similar title.
Further reading
*
External links
*
Northern fulmar profileas part of BTO BirdFacts
Northern fulmar videos photos and soundson the Internet Bird Collection
Southern fulmar videos photos and soundson the Internet Bird Collection
{{Taxonbar, from=Q311224
Fulmarus
Seabirds
Taxa named by James Francis Stephens