Frank Milburn Howlett
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Francis "Frank" Milburn Howlett (5 January 1877 – 20 August 1920) was a British entomologist who served as a Second Imperial Entomologist and as Imperial Pathological Entomologist in India. He specialized in insects (mainly Diptera - sandflies) and parasitic ticks of medical and veterinary importance. A major discovery by him was the attractant methyl eugenol and its effect on flies of the genus '' Bactrocera''.


Life and work

Howlett was born in
Wymondham Wymondham ( ) is a market town and civil parishes in England, civil parish in the South Norfolk district of Norfolk, England, south-west of Norwich, England, Norwich off the A11 road (England), A11 road to London. The River Tiffey runs through ...
,
Norfolk Norfolk () is a ceremonial and non-metropolitan county in East Anglia in England. It borders Lincolnshire to the north-west, Cambridgeshire to the west and south-west, and Suffolk to the south. Its northern and eastern boundaries are the No ...
, the son of Francis John Howlett, a
solicitor A solicitor is a legal practitioner who traditionally deals with most of the legal matters in some jurisdictions. A person must have legally-defined qualifications, which vary from one jurisdiction to another, to be described as a solicitor and ...
, and Mary Jane née Milburn. He was educated at Wymondham Grammar School and
Bury St Edmunds Grammar School King Edward VI School is a co-educational Comprehensive school, comprehensive secondary school in Bury St Edmunds, Suffolk, England. The school in its present form was created in 1972 by the merging of King Edward VI Grammar School, with the Silv ...
, and then at
Christ's College, Cambridge Christ's College is a constituent college of the University of Cambridge. The college includes the Master, the Fellows of the College, and about 450 undergraduate and 170 graduate students. The college was founded by William Byngham in 1437 as ...
. He was an assistant master at
Edinburgh Academy The Edinburgh Academy is an Independent school (United Kingdom), independent day school in Edinburgh, Scotland, which was opened in 1824. The original building, on Henderson Row in the city's New Town, Edinburgh, New Town, is now part of the Se ...
from 1900 to 1903 and at Holt Grammar School before being posted as a professor of natural science (which included the teaching of chemistry) at
Muir Central College Muir Central College in Allahabad in northern India was a college of higher education founded by William Muir in 1872. It had a separate existence to 1921, when as a result of the Allahabad University Act it was merged into Allahabad University. ...
, Allahabad, from 1905 to 1908, initially in a temporary position (to replace E.G. Hill who was on furlough) which was then extended. He joined the Imperial Agricultural Research Institute at Pusa in December 1907 as Second (i.e. deputy) Imperial Entomologist under
Harold Maxwell-Lefroy Harold Maxwell-Lefroy (20 January 1877 – 14 October 1925) was an English entomologist. He served as a Professor of Entomology at Imperial College London and as the second Imperial Entomologist to India. He left India after the death of two of h ...
and from 1912 as Imperial Pathological Entomologist for the Government of India. In 1910 he was in England and Harold Maxwell-Lefroy deputed him to attend the first International Entomological Congress in Brussels, where he presented on the state of economic entomology in India and also on issues in preserving specimens in India. He left India during the
First World War World War I (28 July 1914 11 November 1918), often abbreviated as WWI, was one of the deadliest global conflicts in history. Belligerents included much of Europe, the Russian Empire, the United States, and the Ottoman Empire, with fightin ...
and worked with the
Royal Army Medical College The Royal Army Medical College (RAMC) was located on a site south of the Tate Gallery (now known as Tate Britain) on Millbank, in Westminster, London, overlooking the River Thames. The college moved from the site in 1999 and the buildings are no ...
, while also attending various meetings of learned societies, and returned to India only in 1917. In 1919 he presided over the zoological section of the 6th Indian Science Congress at Bombay, giving a talk on "tactics against insects." One of his most important findings was in noting the attraction of tephritid flies to
methyl eugenol Methyl eugenol (allylveratrol) is a natural chemical compound classified as a phenylpropene, a type of phenylpropanoid. It is the methyl ether of eugenol and is important to insect behavior and pollination. It is found in various essential oils. ...
, a component that he identified from several others present in
citronella oil Citronella oil Citronella oil is an essential oil obtained from the leaves and stems of different species of '' Cymbopogon'' (lemongrass). The oil is used extensively as a source of perfumery chemicals such as citronellal, citronellol, and gerani ...
. He also noted how fleas disliked wet grass and noted a decline in plague during the onset of the rains and suggested that this might be a reason for the plague being rare in Bengal. He published a technique to trap
thrips Thrips ( order Thysanoptera) are minute (mostly long or less), slender insects with fringed wings and unique asymmetrical mouthparts. Different thrips species feed mostly on plants by puncturing and sucking up the contents, although a few are ...
with attractant mixtures of
Benzaldehyde Benzaldehyde (C6H5CHO) is an organic compound consisting of a benzene ring with a formyl substituent. It is the simplest aromatic aldehyde and one of the most industrially useful. It is a colorless liquid with a characteristic almond-like odor. ...
,
Cinnamaldehyde Cinnamaldehyde is an organic compound with the formula(C9H8O) C6H5CH=CHCHO. Occurring naturally as predominantly the ''trans'' (''E'') isomer, it gives cinnamon its flavor and odor. It is a phenylpropanoid that is naturally synthesized by the shik ...
and
Anisaldehyde 4-Anisaldehyde, or ''p''-Anisaldehyde, is an organic compound with the formula CH3OC6H4CHO. The molecule consists of a benzene ring with an formyl and a methoxy group. It is a colorless liquid with a strong aroma. It provides sweet, floral and ...
. Howlett also discovered that he could induce ''Stomoxys calcitrans'' to oviposit on cotton impregnated with valerianic acid, which is a component of fermenting vegetable matter. He also noted the life history of mosquito larvae (''Stegomyia'', now ''
Anopheles ''Anopheles'' () is a genus of mosquito first described and named by J. W. Meigen in 1818. About 460 species are recognised; while over 100 can transmit human malaria, only 30–40 commonly transmit parasites of the genus ''Plasmodium'', which c ...
'') that could survive in dry soil. He also studied the biology of
sandflies Sandfly (or sand fly) is a colloquial name for any species or genus of flying, biting, blood-sucking dipteran (fly) encountered in sandy areas. In the United States, ''sandfly'' may refer to certain horse flies that are also known as "greenhe ...
. Howlett was also known for his humorous sketches at Pusa, but he later moved to the Agricultural Research Institute at Pune. Howlett was an athlete and artist but his health was poor during his service in India and he died a premature death due to complications following a surgical procedure at
Mussoorie Mussoorie is a hill station and a municipal board, near Dehradun city in the Dehradun district of the Indian state of Uttarakhand. It is about from the state capital of Dehradun and north of the national capital of New Delhi. The hill s ...
. At the time of his death he was working on a book titled ''The Control of Harmful Insects''. An obituarist writing in the ''Agricultural Journal of India'' noted that he was:
... a man of almost childlike simplicity and originality of outlook, and with many interests. ... He had the faults as well as the merits of the artistic temperament. ... He was a born schoolmaster, delighted in teaching, and could make all subjects interesting, and had the gift of implanting in his disciples some of his own enthusiasm. He was a combative apostle of pure research, and his disappearance from the ranks of scientists of this order will be a serious blow to the cause.
Howlett assisted Harold Maxwell-Lefroy in writing and illustrating the book ''Indian Insect Life''. He wrote the sections on the flies and also trained staff at Pusa in technical illustration. A species of tick, ''Haemaphysalis howletti'', was described by Warburton in 1913 from a pony in Pakistan and in 1962 was found on rodents and birds in Pune, Maharashtra. Howlett developed techniques for collecting and preserving insects and for marking insects (houseflies) to study dispersal. Brunetti, named a fly after Howlett as ''Howlettia'' (now considered a synonym of '' Platypalpus'' of family Hybotidae).


Publications

Apart from the publications cited, Howlett's works include: * * * * * * * * *


References

{{DEFAULTSORT:Howlett, Frank Milburn British entomologists 1877 births 1920 deaths Alumni of the University of Cambridge