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A fossil track or ichnite (
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
"''ιχνιον''" (''ichnion'') – a track, trace or footstep) is a fossilized
footprint Footprints are the impressions or images left behind by a person walking or running. Hoofprints and pawprints are those left by animals with hooves or paws rather than feet, while "shoeprints" is the specific term for prints made by shoes. The ...
. This is a type of
trace fossil A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, ...
. A fossil trackway is a sequence of fossil tracks left by a single organism. Over the years, many ichnites have been found, around the world, giving important clues about the behaviour (and foot structure and stride) of the animals that made them. For instance, multiple ichnites of a single species, close together, suggest 'herd' or 'pack' behaviour of that species. Combinations of footprints of different species provide clues about the interactions of those species. Even a set of footprints of a single animal gives important clues, as to whether it was
biped Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' ...
al or
quadruped Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuor' ...
al. In this way, it has been suggested that some
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...
s, when on the ground, used their forelimbs in an unexpected quadrupedal action. Special conditions are required, in order to preserve a footprint made in soft ground (such as an alluvial plain or a formative
sedimentary Sedimentary rocks are types of rock that are formed by the accumulation or deposition of mineral or organic particles at Earth's surface, followed by cementation. Sedimentation is the collective name for processes that cause these particles ...
deposit). A possible scenario is a sea or lake shore that became dried out to a firm mud in hot, dry conditions, received the footprints (because it would only have been partially hardened and the animal would have been heavy) and then became silted over in a flash storm. The first ichnite found was in 1800 in Massachusetts, US, by a farmer named Pliny Moody, who found 1-foot (31 cm) long fossilized footprints. They were thought by Harvard and Yale scholars to be from "Noah's Raven". A famous group of ichnites was found in a limestone quarry at Ardley, 20 km Northeast of
Oxford Oxford () is a city in England. It is the county town and only city of Oxfordshire. In 2020, its population was estimated at 151,584. It is north-west of London, south-east of Birmingham and north-east of Bristol. The city is home to the ...
,
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
, in 1997. They were thought to have been made by ''
Megalosaurus ''Megalosaurus'' (meaning "great lizard", from Greek , ', meaning 'big', 'tall' or 'great' and , ', meaning 'lizard') is an extinct genus of large carnivorous theropod dinosaurs of the Middle Jurassic period (Bathonian stage, 166 million years ...
'' and possibly ''
Cetiosaurus ''Cetiosaurus'' () meaning 'whale lizard', from the Greek '/ meaning 'sea monster' (later, 'whale') and '/ meaning 'lizard', is a genus of herbivorous sauropod dinosaur from the Middle Jurassic Period, living about 168 million years ago in what ...
''. There are replicas of some of these footprints, set across the lawn of
Oxford University Museum of Natural History The Oxford University Museum of Natural History, sometimes known simply as the Oxford University Museum or OUMNH, is a museum displaying many of the University of Oxford's natural history specimens, located on Parks Road in Oxford, England. It a ...
(OUMNH). A creature named '' Cheirotherium'' was, for a long time and still may be, only known from its fossilized trail. Its footprints were first found in 1834, in
Thuringia Thuringia (; german: Thüringen ), officially the Free State of Thuringia ( ), is a state of central Germany, covering , the sixth smallest of the sixteen German states. It has a population of about 2.1 million. Erfurt is the capital and lar ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
, dating from the Late
Triassic The Triassic ( ) is a geologic period and system (stratigraphy), system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Year#Abbreviations yr and ya, Mya), to the beginning of the Jurassic Period 201.36 ...
Period Period may refer to: Common uses * Era, a length or span of time * Full stop (or period), a punctuation mark Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (or rhetorical period), a concept ...
. The largest known dinosaur footprints, belonging to
sauropods Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
and dating from the early
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
were found to the north of Broome on the
Dampier Peninsula The Dampier Peninsula is a peninsula located north of Broome and Roebuck Bay in Western Australia. It is surrounded by the Indian Ocean to the west and north, and King Sound to the east. It is named after the mariner and explorer William Damp ...
,
Western Australia Western Australia (commonly abbreviated as WA) is a state of Australia occupying the western percent of the land area of Australia excluding external territories. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to th ...
, with some footprints measuring 1.7 m. The 3D digital documentation of tracks has the benefit of being able to examine ichnite in detail remotely and distribute the data to colleagues and other interested personnel.


Fossil trackways

Many fossil trackways were made by dinosaurs, early
tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids ( pelycosaurs, extinct t ...
s, and other
quadruped Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuor' ...
s and
biped Bipedalism is a form of terrestrial locomotion where an organism moves by means of its two rear limbs or legs. An animal or machine that usually moves in a bipedal manner is known as a biped , meaning 'two feet' (from Latin ''bis'' 'double' ...
s on land. Marine organisms also made many ancient trackways (such as the trails of
trilobite Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the ...
s and
eurypterid Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 million years ago. The group is l ...
s like ''
Hibbertopterus ''Hibbertopterus'' is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of ''Hibbertopterus'' have been discovered in deposits ranging from the Devonian period in Belgium, Scotland and the United States to the Carboniferous ...
''). Some basic fossil trackway types: :#footprints :#tail drags :#belly drag marks – (e.g., tetrapods) :#chain of trace platforms – (example: ''
Yorgia ''Yorgia waggoneri'' is a discoid Ediacaran organism. It has a low, segmented body consisting of a short wide "head", no appendages, and a long body region, reaching a maximum length of . It is classified within the extinct animal phylum Proarti ...
'') :#body imprint – ( Monuron trackway, insect) The majority of fossil trackways are foot impressions on land, or subsurface water, but other types of creatures will leave distinctive impressions. Examples of creatures supported, or partially supported, in a
water Water (chemical formula ) is an Inorganic compound, inorganic, transparent, tasteless, odorless, and Color of water, nearly colorless chemical substance, which is the main constituent of Earth's hydrosphere and the fluids of all known living ...
environment are known. The fossil "
millipede Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a resu ...
-type" genus ''
Arthropleura ''Arthropleura'' () is a genus of extinct millipede arthropods that lived in what is now North America and Europe around 345 to 290 million years ago, from the Viséan stage of the lower Carboniferous Period to the Sakmarian stage of the lower ...
'' left its multi-legged/feet trackways on land.


Hominid trackways


Africa


=Tanzania

= Some of the earliest trackways for human ancestors have been discovered in
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands ...
. The
Laetoli Laetoli is a pre-historic site located in Enduleni ward of Ngorongoro District in Arusha Region, Tanzania. The site is dated to the Plio-Pleistocene and famous for its Hominina footprints, preserved in volcanic ash. The site of the Laetoli fo ...
trackway is famous for the
hominin The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The ...
footprints preserved in
volcanic ash Volcanic ash consists of fragments of rock, mineral crystals, and volcanic glass, created during volcanic eruptions and measuring less than 2 mm (0.079 inches) in diameter. The term volcanic ash is also often loosely used to refer ...
. After the footprints were made in powdery ash, soft rain cemented the ash layer into
tuff Tuff is a type of rock made of volcanic ash ejected from a vent during a volcanic eruption. Following ejection and deposition, the ash is lithified into a solid rock. Rock that contains greater than 75% ash is considered tuff, while rock ...
, preserving the prints. The hominid prints were produced by three individuals, one walking in the footprints of the other, making the original tracks difficult to discover. As the tracks lead in the same direction, they might have been produced by a group – but there is nothing else to support the common reconstruction of a nuclear family visiting the waterhole together.


=South Africa

= In
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
, two ancient trackways have been found containing footprints, one at Langebaan and one at Nahoon. Both trackways occur in calcareous eolianites or hardened sand dunes. At Nahoon, trackways of at least five species of vertebrates, including three hominid footprints, are preserved as casts. The prints at
Langebaan Langebaan is a town in the Western Cape province of South Africa on the eastern shore of Langebaan Lagoon. Langebaan is situated 120 km north of Cape Town, just off the R27, about 28 km from Vredenburg and 20 km from Saldanha Bay. ...
are the oldest human footprints, dated to approximately 117,000 years old.


Australia


=New South Wales

= Twenty six human fossil trackways have been found in the Willandra Lakes area adjacent to Lake Garnpung, consisting of 563 human footprints from 19,000 to 20,000 years ago.


Early Tetrapod

The earliest land creatures (actually land-marine coastal-riverine-marshland) left some of the first terrestrial trackways. They range from tetrapods to proto-reptilians and others. A possible first connection of a trackway with the vertebrate that left it was published by Drs. Sebastian Voigt and David Berman and Amy Henrici in the 12 September 2007 issue of ''
Journal of Vertebrate Paleontology The ''Journal of Vertebrate Paleontology'' is a bimonthly peer-reviewed scientific journal that was established in 1980 by Jiri Zidek (University of Oklahoma). It covers all aspects of vertebrate paleontology, including vertebrate origins, evolu ...
''. The paleontologists who made the connection were aided by unusually detailed trackways left in fine-grained
Lower Permian The Cisuralian is the first series/epoch of the Permian. The Cisuralian was preceded by the Pennsylvanian and followed by the Guadalupian. The Cisuralian Epoch is named after the western slopes of the Ural Mountains in Russia and Kazakhstan an ...
mud of the Tambach Formation in central Germany, together with exceptionally complete fossilised skeletons in the same 290-million-year-old strata. They matched the two most common trackways with the two most common fossils, two reptile-like herbivores known as '' Diadectes absitus'' (with the trackway pseudonym ''Ichniotherium cottae'') and '' Orobates pabsti'' (with the trackway pseudonym of ''Orobates pabsti''). The Permo- Carboniferous of
Prince Edward Island Prince Edward Island (PEI; ) is one of the thirteen provinces and territories of Canada. It is the smallest province in terms of land area and population, but the most densely populated. The island has several nicknames: "Garden of the Gulf", ...
, Canada contains trackways of tetrapods and stem-reptiles. Macrofloral and
palynological Palynology is the "study of dust" (from grc-gre, παλύνω, palynō, "strew, sprinkle" and ''-logy'') or of "particles that are strewn". A classic palynologist analyses particulate samples collected from the air, from water, or from deposit ...
information help date them. Ireland hosts late Middle Devonian tetrapod trackways at three sites on
Valentia Island Valentia Island () is one of Ireland's most westerly points. It lies off the Iveragh Peninsula in the southwest of County Kerry. It is linked to the mainland by the Maurice O'Neill Memorial Bridge at Portmagee. A car ferry also departs from ...
within the Valentia Slate Formation. The earliest fossil trackway of primitive tetrapods in Australia occurs in the Genoa River Gorge, Victoria, dating from the Devonian 350 million years ago.


Dinosaur trackways

Dinosaurs lived on the continents before
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns a ...
es evolved (the "Age of the Grasses" evolved with the "Age of the Mammals"); the dinosaurs lived in the
Triassic The Triassic ( ) is a geologic period and system (stratigraphy), system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Year#Abbreviations yr and ya, Mya), to the beginning of the Jurassic Period 201.36 ...
,
Jurassic The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...
, and
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
and left many trackways, both from plant-eaters and the meat-eaters, in various layers of mud and sand. With scientific analysis, dinosaur specialists are now analyzing tracks for the ''walking-speeds'', or ''sprint''-''running speeds'' for all categories of dinosaurs, even to the large plant eaters, but especially the faster 3-toed meat hunters. ''Evidence of'' ''herding'', as well as ''pack hunting'' are also being investigated.


Brazil

* Valley of the Dinosaurs, Paraíba, Brazil


Africa


= Namibia

= In north-central Namibia there is a dinosaur trackway in sandstone on what is now the private farm Otjihaenamparero. Larger footprints are of a
ceratosauria Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with ''Ceratosaurus'' than with birds. The oldest known ceratosaur, '' Saltriovenator'', dates to the earliest ...
and smaller ones of syntarsus. The prints are believed to be around 190 million years old.


=Zimbabwe

= In the Lower Zimbabwe Rift Valley there is a trackway in 140 Ma rose-coloured
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates ...
of Chewore Area. The small footprint size, with both manus and pes, implies that it is a trackway of a juvenile, a probable
carnosaur Carnosauria is an extinct large group of predatory dinosaurs that lived during the Jurassic and Cretaceous periods. Starting from the 1990s, scientists have discovered some very large carnosaurs in the carcharodontosaurid family, such as ''Gig ...
.


North America

The western regions of North America, especially the western border of the
Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient sea ...
, are common for dinosaur trackways.
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to the s ...
has dinosaur trackways from the Late Cretaceous, 65 ma. (A model example of this 3-toed Wyoming trackway was made for presentation) In the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
,
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
footprints and trackways are found in the
Glen Rose Formation The Glen Rose Formation is a shallow marine to shoreline geological formation from the lower Cretaceous period exposed over a large area from South Central to North Central Texas. The formation is most widely known for the dinosaur footprints ...
, the most famous of these being the Paluxy River site in
Dinosaur Valley State Park Dinosaur Valley State Park is a state park near Glen Rose, Texas, United States. History Dinosaur Valley State Park, located just northwest of Glen Rose in Somervell County, Texas, is a scenic park set astride the Paluxy River. The land fo ...
. These were the first
sauropoda Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
footprints scientifically documented, and were designated a US
National Natural Landmark The National Natural Landmarks (NNL) Program recognizes and encourages the conservation of outstanding examples of the natural history of the United States. It is the only national natural areas program that identifies and recognizes the best ...
in 1969. Some are as large as about 3 feet across. The prints are thought to have been preserved originally in a
tidal flat Mudflats or mud flats, also known as tidal flats or, in Ireland, slob or slobs, are coastal wetlands that form in intertidal areas where sediments have been deposited by tides or rivers. A global analysis published in 2019 suggested that tidal fl ...
or a
lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into ''coastal lagoons'' (or ''barrier lagoons'') ...
.Martin Lockley & Adrian P. Hunt, Dinosaur Tracks and Other Fossil Footprints of the Western United States, Columbia U. Press, New York (1995) There are tracks from two types of dinosaur. The first type of tracks are from a sauropod and were made by an animal of 30 to 50 feet in length, perhaps a
brachiosaurid The Brachiosauridae ("arm lizards", from Greek ''brachion'' (βραχίων) = "arm" and ''sauros'' = "lizard") are a family or clade of herbivorous, quadrupedal sauropod dinosaurs. Brachiosaurids had long necks that enabled them to access the le ...
such as '' Pleurocoelus'', and the second tracks by a theropoda, an animal of 20 to 30 feet in length, perhaps an ''
Acrocanthosaurus ''Acrocanthosaurus'' ( ; ) is a genus of carcharodontosaurid dinosaur that existed in what is now North America during the Aptian and early Albian stages of the Early Cretaceous, from 113 to 110 million years ago. Like most dinosaur genera, ' ...
''. A variety of scenarios was proposed to explain the tracks, but most likely represent twelve sauropods "probably as a herd, followed somewhat later by three theropods that may or may not have been stalking – but that certainly were not attacking." Other examples include: *Dinosaur tracks, near
Moab, Utah Moab () is the largest city and county seat of Grand County in eastern Utah in the western United States, known for its dramatic scenery. The population was 5,366 at the 2020 census. Moab attracts many tourists annually, mostly visitors to ...
* Dinosaur Footprints Reservation in Holyoke, Massachusetts, US * Red Gulch Dinosaur Tracksite, Wyoming *
Prehistoric Trackways National Monument Prehistoric Trackways National Monument is a national monument in the Robledo Mountains of Doña Ana County, New Mexico, United States, near the city of Las Cruces. The monument's Paleozoic Era fossils are on of land administered by the Bur ...
near
Las Cruces, New Mexico Las Cruces (; "the crosses") is the second-largest city in the U.S. state of New Mexico and the seat of Doña Ana County. As of the 2020 census the population was 111,385. Las Cruces is the largest city in both Doña Ana County and southern Ne ...
*
Connecticut River Valley trackways The Connecticut River Valley trackways are the fossilised footprints of a number of Early Jurassic dinosaurs or other archosauromorphs from the sandstone beds of Massachusetts and Connecticut. The finding has the distinction of being among the firs ...
, in New England * Clayton Lake State Park dinosaur trackway near Clayton, New Mexico


China

The Gansu dinosaur trackway located in the Liujiazia National Dinosaur Geopark in Yanguoxia, China contains hundreds of tracks including 245 dinosaur, 350 theropod, 364 sauropod and 628 ornithopod tracks among others.


Australia

The Lark Quarry Trackway in
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , establishe ...
contains three-toed tracks made by a heard of
ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous wo ...
dinosaurs crossing a river. It was once believed they respresented a large predator chasing doqn a mixed flock of small ornithopods and theropods, but this was contested in 2011.


Mammal trackways

Mammal trackways are among the least common trackways. Mammals were not often in mud, or riverine environments; they were more often in forestlands or grasslands. Thus the earlier tetrapods or proto-tetrapods would yield the most fossil trackways. The
Walchia ''Walchia'' is a fossil conifer, Cupressaceae, cypress-like genus found in upper Pennsylvanian (geology), Pennsylvanian (Carboniferous) and lower Permian (about 310-290 Mya (unit), Mya) rocks of Europe and North America. A forest of In situ, in-s ...
forest of
Brule, Nova Scotia Brule () is a rural community located in Colchester County, Nova Scotia, Canada. Located on Amet Sound an embayment of the Northumberland Strait 6 kilometres east of the village of Tatamagouche near the county boundary, the community is situat ...
has an example of an
in situ ''In situ'' (; often not italicized in English) is a Latin phrase that translates literally to "on site" or "in position." It can mean "locally", "on site", "on the premises", or "in place" to describe where an event takes place and is used in ...
Walchia forest, and tetrapod trackways that extended over some period of time through the forest area.


United States

*A 1.5km-long Late Pleistocene Age trackway of Human (child and adult) fossilized footprints, as well as
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks an ...
and
giant ground sloth ''Megatherium'' ( ; from Greek () 'great' + () 'beast') is an extinct genus of ground sloths endemic to South America that lived from the Early Pliocene through the end of the Pleistocene. It is best known for the elephant-sized type species ' ...
tracks have been found at White Sands National Park. Near Alamogordo, New Mexico


Australia

A recent
marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...
trackway site in the Colac district of Australia (west of Colac) contains marsupial trackways as well as kangaroo and wallaby tracks.Unrivalled fossil find, The Cola Herald
/ref>


Gallery of images

Image:Cheirotherium prints possibly Ticinosuchus.JPG, ''Cheirotherium'' trace fossil, displayed in
Oxford University Museum of Natural History The Oxford University Museum of Natural History, sometimes known simply as the Oxford University Museum or OUMNH, is a museum displaying many of the University of Oxford's natural history specimens, located on Parks Road in Oxford, England. It a ...
. Image:MammothFootImpressions25.jpg, Cross-section of
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in ...
mammoth A mammoth is any species of the extinct elephantid genus ''Mammuthus'', one of the many genera that make up the order of trunked mammals called proboscideans. The various species of mammoth were commonly equipped with long, curved tusks an ...
footprints at The Mammoth Site, Hot Springs, South Dakota. Image:Eubrontes01.JPG, ''Eubrontes'', a
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
footprint in the Lower
Jurassic The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...
Moenave Formation The Moenave Formation is a Mesozoic geologic formation, in the Glen Canyon Group. It is found in Utah and Arizona. The Moenave was deposited on an erosion surface on the Chinle Formation following an early Jurassic uplift and unconformity that re ...
at the
St. George Dinosaur Discovery Site The St. George Dinosaur Discovery Site is a fossil site and museum at Johnson Farm in Saint George, Utah. The museum preserves thousands of dinosaur footprints right at the original site of discovery. The site was discovered by accident on Febr ...
at Johnson Farm, southwestern Utah. Image:Gigandipus.JPG, ''Gigandipus'', a
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
footprint in the Lower
Jurassic The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...
Moenave Formation The Moenave Formation is a Mesozoic geologic formation, in the Glen Canyon Group. It is found in Utah and Arizona. The Moenave was deposited on an erosion surface on the Chinle Formation following an early Jurassic uplift and unconformity that re ...
at the St. George Dinosaur Discovery Site at Johnson Farm, southwestern Utah. Image:CamelFootprintBarstowMiocene.jpg, Cameloid footprint (''Lamaichnum alfi'' Sarjeant and Reynolds, 1999; convex hyporelief) from the
Barstow Formation The Barstow Formation is a series of limestones, conglomerates, sandstones, siltstones and shales exposed in the Mojave Desert near Barstow in San Bernardino County, California.Dibblee, T.W., Jr. (1967). Areal Geology of the Western Mojave De ...
(
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
) of
Rainbow Basin Rainbow Basin is a geological formation in the Calico Peaks range, located approximately north of Barstow in the Mojave Desert in San Bernardino County, California. The Rainbow Basin has been designated a National Natural Landmark and is in th ...
,
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
. Image:Saurichnites intermedius.jpg, ''Saurichnites intermedius'' Image:Enciso-dinosaur-footprints-track.jpg, A carnivorous
theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...
trackway, near Enciso,
La Rioja La Rioja () is an autonomous community and province in Spain, in the north of the Iberian Peninsula. Its capital is Logroño. Other cities and towns in the province include Calahorra, Arnedo, Alfaro, Haro, Santo Domingo de la Calzada, an ...
, Spain Image:Diplichnites.jpg, ''Petalichnus'',
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
walking traces. Devonian of northeastern
Ohio Ohio () is a state in the Midwestern region of the United States. Of the fifty U.S. states, it is the 34th-largest by area, and with a population of nearly 11.8 million, is the seventh-most populous and tenth-most densely populated. The sta ...
. File:Moa footprints.jpg,
Moa Moa are extinct giant flightless birds native to New Zealand. The term has also come to be used for chicken in many Polynesian cultures and is found in the names of many chicken recipes, such as Kale moa and Moa Samoa. Moa or MOA may also refe ...
footprints near the Manawatu River, New Zealand. Image:Hibbertopteroid track.jpg, ''
Hibbertopterus ''Hibbertopterus'' is a genus of eurypterid, a group of extinct aquatic arthropods. Fossils of ''Hibbertopterus'' have been discovered in deposits ranging from the Devonian period in Belgium, Scotland and the United States to the Carboniferous ...
'' trackway: ''negative relief image'', a groove infilled by sand appears as a ridgeline


See also

* ''
Ichnites ''Ichnites'' is an ichnogenus of dinosaur footprint. See also * List of dinosaur ichnogenera This list of dinosaur ichnogenera is a comprehensive listing of all ichnogenera that have been attributed to dinosaurs, excluding class Aves ( bird ...
'', a type of ichnite *
List of stratigraphic units with dinosaur tracks Indeterminate or unspecified dinosaur tracks Ornithischians Indeterminate or unspecified ornithischian tracks Saurischians Theropods Sauropodomorphs See also List of dinosaur-bearing rock formations This list of dinosaur-bearing rock f ...
* Formations with ichnites * List of non-Dinosauria fossil trackway articles *
List of fossil sites This list of fossil sites is a worldwide list of localities known well for the presence of fossils. Some entries in this list are notable for a single, unique find, while others are notable for the large number of fossils found there. Many of t ...


References


External links


Texts on natural casts of dinosaur tracks found in Utah coal mines
*

Dinosaur trackways:
Photo-High Res – (Outdoor photo)
Photo from
Dakota Formation The Dakota is a sedimentary rock, sedimentary geologic unit name of Formation (stratigraphy), formation and Group (stratigraphy), group rank in Midwestern North America. The Dakota units are generally composed of sandstones, mudstones, clays, and ...
, Colorado Early Tetrapods:
Earth History, (tetrapod trackways, etc.)
Australia
Marsupial trackways, Colac district.
{{DEFAULTSORT:Fossil Trackway Trace fossils