Ceratosauria
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Ceratosauria
Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with '' Ceratosaurus'' than with birds. The oldest known ceratosaur, '' Saltriovenator'', dates to the earliest part of the Jurassic, around 199 million years ago. According to the majority of the latest research, Ceratosauria includes three major clades: Ceratosauridae, Noasauridae, and Abelisauridae, found primarily (though not exclusively) in the Southern Hemisphere. Originally, Ceratosauria included the above dinosaurs plus the Late Triassic to Early Jurassic Coelophysoidea and Dilophosauridae, implying a much earlier divergence of ceratosaurs from other theropods. However, most recent studies have shown that coelophysoids and dilophosaurids do not form a natural group with other ceratosaurs, and are excluded from this group. Ceratosauria derives its names from the type species, '' Ceratosaurus nasicornis'', described by O.C. Marsh in 1 ...
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Ceratosaurus Nasicornis
''Ceratosaurus'' (from Greek κέρας/κέρατος, ' meaning "horn" and σαῦρος ' meaning "lizard") was a carnivorous theropod dinosaur in the Late Jurassic period (Kimmeridgian to Tithonian). The genus was first described in 1884 by American paleontologist Othniel Charles Marsh based on a nearly complete skeleton discovered in Garden Park, Colorado, in rocks belonging to the Morrison Formation. The type species is ''Ceratosaurus nasicornis''. The Garden Park specimen remains the most complete skeleton known from the genus, and only a handful of additional specimens have been described since. Two additional species, ''Ceratosaurus dentisulcatus'' and ''Ceratosaurus magnicornis'', were described in 2000 from two fragmentary skeletons from the Cleveland-Lloyd Quarry of Utah and from the vicinity of Fruita, Colorado. The validity of these additional species has been questioned, however, and all three skeletons possibly represent different growth stages of the same spe ...
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Ceratosaurus
''Ceratosaurus'' (from Greek κέρας/κέρατος, ' meaning "horn" and σαῦρος ' meaning "lizard") was a carnivorous theropod dinosaur in the Late Jurassic period (Kimmeridgian to Tithonian). The genus was first described in 1884 by American paleontologist Othniel Charles Marsh based on a nearly complete skeleton discovered in Garden Park, Colorado, in rocks belonging to the Morrison Formation. The type species is ''Ceratosaurus nasicornis''. The Garden Park specimen remains the most complete skeleton known from the genus, and only a handful of additional specimens have been described since. Two additional species, ''Ceratosaurus dentisulcatus'' and ''Ceratosaurus magnicornis'', were described in 2000 from two fragmentary skeletons from the Cleveland-Lloyd Quarry of Utah and from the vicinity of Fruita, Colorado. The validity of these additional species has been questioned, however, and all three skeletons possibly represent different growth stages of the same spe ...
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Theropoda
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic period 231.4 million years ago ( Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of diets was historically considered a characteris ...
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Berberosaurus
''Berberosaurus'' (meaning " Berber lizard", in reference to the Berbers of Morocco) is a genus of neotheropod dinosaur, possibly a ceratosaur, from the Early Toarcian-age (Lower Jurassic) Azilal Formation found in the High Atlas of Toundoute, Ouarzazate, Morocco. The type species of the genus ''Berberosaurus'' is ''B. liassicus'', in reference to the Lias epoch. ''Berberosaurus'' might be the oldest known ceratosaur, and is based on partial postcranial remains. Classification Ronan Allain and colleagues, who described ''Berberosaurus'', performed a phylogenetic analysis and found their new genus to be the most basal known abelisauroid, more derived than ''Elaphrosaurus'', ''Ceratosaurus'', and ''Spinostropheus'', but less so than ''Xenotarsosaurus'' and abelisaurians. ''Berberosaurus'' is distinguished from other theropods by anatomical details found in its vertebrae, metacarpals, and hindlimb bones. Its assignment as an abelisauroid pushes back the record of this group and sho ...
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Abelisauridae
Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are found on the modern continents of Africa and South America, as well as on the Indian subcontinent and the island of Madagascar. Isolated teeth were found in the Late Jurassic of Portugal, and the Late Cretaceous genera ''Tarascosaurus'' and '' Arcovenator'' have been described in France. Abelisaurids first appear in the fossil record of the early middle Jurassic period, and at least two genera (the Moroccan '' Chenanisaurus'' and the Madagascan '' Majungasaurus'') survived until the end of the Mesozoic era 66 million years ago. Like most theropods, abelisaurids were carnivorous bipeds. They were characterized by stocky hind limbs and extensive ornamentation of the skull bones, with grooves and pits. In many abelisaurids, such as '' Carnotauru ...
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Theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally carnivorous, although a number of theropod groups evolved to become herbivores and omnivores. Theropods first appeared during the Carnian age of the late Triassic period 231.4 million years ago ( Ma) and included all the large terrestrial carnivores from the Early Jurassic until at least the close of the Cretaceous, about 66 Ma. In the Jurassic, birds evolved from small specialized coelurosaurian theropods, and are today represented by about 10,500 living species. Biology Diet and teeth Theropods exhibit a wide range of diets, from insectivores to herbivores and carnivores. Strict carnivory has always been considered the ancestral diet for theropods as a group, and a wider variety of diets was historically considered a ch ...
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Noasauridae
Noasauridae is an extinct family of theropod dinosaurs belonging to the group Ceratosauria. They were closely related to the short-armed abelisaurids, although most noasaurids had much more traditional body types generally similar to other theropods. Their heads, on the other hand, had unusual adaptations depending on the subfamily. 'Traditional' noasaurids, sometimes grouped in the subfamily Noasaurinae, had sharp teeth which splayed outwards from a downturned lower jaw. The most complete and well-known example of these kinds of noasaurids was '' Masiakasaurus knopfleri'' from Madagascar. Another group, Elaphrosaurinae, has also been placed within Noasauridae by some studies. Elaphrosaurines developed toothless jaws and herbivorous diets, at least as adults. The most complete and well known elaphrosaurine was '' Limusaurus inextricabilis''. At least some noasaurids had pneumatised cervical vertebrae.Arthur Souza Brum, Elaine Batista Machado, Diogenes de Almeida Campos & Al ...
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Carnotaurus
''Carnotaurus'' (; ) is a genus of theropod dinosaur that lived in South America during the Late Cretaceous period, probably sometime between 71 and 69 million years ago. The only species is ''Carnotaurus sastrei''. Known from a single well-preserved skeleton, it is one of the best-understood theropods from the Southern Hemisphere. The skeleton, found in 1984, was uncovered in the Chubut Province of Argentina from rocks of the La Colonia Formation. ''Carnotaurus'' is a derived member of the Abelisauridae, a group of large theropods that occupied the large predatorial niche in the southern landmasses of Gondwana during the late Cretaceous. Within the Abelisauridae, the genus is often considered a member of the Brachyrostra, a clade of short-snouted forms restricted to South America. ''Carnotaurus'' was a lightly built, bipedal predator, measuring in length and weighing . As a theropod, ''Carnotaurus'' was highly specialized and distinctive. It had thick horns above the ey ...
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Deltadromeus
''Deltadromeus'' (meaning "delta runner") is a genus of theropod dinosaur from Northern Africa. It had long, unusually slender hind limbs for its size, suggesting that it was a swift runner. The skull is not known. One fossil specimen of a single species (''D. agilis'', or "agile delta runner") has been described, found in the Kem Kem Beds, which date to the mid-Cretaceous Period (mid- Cenomanian age), about 95 million years ago. It may be a junior synonym of the contemporary '' Bahariasaurus''.Holtz, Thomas R. Jr. (2008) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages'Supplementary Information/ref> The classification of ''Deltadromeus'' has been in flux since its original description. In 2016, a South American theropod known as '' Gualicho shinyae'' was found to possess many similarities with ''Deltadromeus''. Depending on the phylogenetic position of ''Gualicho'', ''Deltadromeus'' may have been a neovenatorid carnosaur, a tyrannosauro ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species ...
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Coelophysoidea
Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivore, carnivorous forms with a superficial similarity to the coelurosaurs, with which they were formerly classified, and some species had delicate cranium, cranial crests. Sizes range from about 1 to 6 m in length. It is unknown what kind of external covering coelophysoids had, and various artists have portrayed them as either scaly or feathered dinosaurs, feathered. Some species may have lived in packs, as inferred from sites where numerous individuals have been found together. Examples of coelophysoids include ''Coelophysis'', ''Procompsognathus'' and ''Liliensternus''. Most dinosaurs formerly referred to as being in the dubious taxon "Podokesauridae" are now classified as coelophysoids. Classification Despite their very early occurrence in the fossil record, coelophysoids have a numb ...
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Jacques Gauthier
Jacques Armand Gauthier (born June 7, 1948 in New York City) is an American vertebrate paleontologist, comparative morphologist, and systematist, and one of the founders of the use of cladistics in biology. Life and career Gauthier is the son of Edward Paul Gauthier and Patricia Marie Grogan. He received a B.S. degree in Zoology at San Diego State University in 1973, a Masters of Biological Science at the same institute in 1980, and a PhD in Paleontology from the University of California, Berkeley, in 1984. Currently he is a Professor of Geology and Geophysics and Ecology and Evolutionary Biology and Curator of Vertebrate Paleontology and Vertebrate Zoology at Yale University. His master's thesis, the content of which was published in 1982, is a classic work on the paleontology and phylogeny of the lizard clade Anguimorpha that remains a core reference for morphological research on Xenosauridae and Anguidae in particular. His PhD thesis constituted the first major cladist ...
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