Ceratosauria
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Ceratosauria
Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with ''Ceratosaurus'' than with birds. The oldest known ceratosaur, '' Saltriovenator'', dates to the earliest part of the Jurassic, around 199 million years ago. According to the majority of the latest research, Ceratosauria includes three major clades: Ceratosauridae, Noasauridae, and Abelisauridae, found primarily (though not exclusively) in the Southern Hemisphere. Originally, Ceratosauria included the above dinosaurs plus the Late Triassic to Early Jurassic Coelophysoidea and Dilophosauridae, implying a much earlier divergence of ceratosaurs from other theropods. However, most recent studies have shown that coelophysoids and dilophosaurids do not form a natural group with other ceratosaurs, and are excluded from this group. Ceratosauria derives its names from the type species, ''Ceratosaurus nasicornis'', described by O.C. Marsh in 1884. A ...
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Noasauridae
Noasauridae is an extinct family of theropod dinosaurs belonging to the group Ceratosauria. They were closely related to the short-armed abelisaurids, although most noasaurids had much more traditional body types generally similar to other theropods. Their heads, on the other hand, had unusual adaptations depending on the subfamily. 'Traditional' noasaurids, sometimes grouped in the subfamily Noasaurinae, had sharp teeth which splayed outwards from a downturned lower jaw. The most complete and well-known example of these kinds of noasaurids was '' Masiakasaurus knopfleri'' from Madagascar. Another group, Elaphrosaurinae, has also been placed within Noasauridae by some studies. Elaphrosaurines developed toothless jaws and herbivorous diets, at least as adults. The most complete and well known elaphrosaurine was ''Limusaurus inextricabilis''. At least some noasaurids had pneumatised cervical vertebrae.Arthur Souza Brum, Elaine Batista Machado, Diogenes de Almeida Campos & Al ...
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Berberosaurus
''Berberosaurus'' (meaning "Berber lizard", in reference to the Berbers of Morocco) is a genus of neotheropod dinosaur, possibly a ceratosaur, from the Early Toarcian-age (Lower Jurassic) Azilal Formation found in the High Atlas of Toundoute, Ouarzazate, Morocco. The type species of the genus ''Berberosaurus'' is ''B. liassicus'', in reference to the Lias epoch. ''Berberosaurus'' might be the oldest known ceratosaur, and is based on partial postcranial remains. Classification Ronan Allain and colleagues, who described ''Berberosaurus'', performed a phylogenetic analysis and found their new genus to be the most basal known abelisauroid, more derived than ''Elaphrosaurus'', ''Ceratosaurus'', and '' Spinostropheus'', but less so than '' Xenotarsosaurus'' and abelisaurians. ''Berberosaurus'' is distinguished from other theropods by anatomical details found in its vertebrae, metacarpals, and hindlimb bones. Its assignment as an abelisauroid pushes back the record of this group and ...
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Ceratosaurus Nasicornis
''Ceratosaurus'' (from Greek κέρας/κέρατος, ' meaning "horn" and σαῦρος ' meaning "lizard") was a carnivorous theropod dinosaur in the Late Jurassic period (Kimmeridgian to Tithonian). The genus was first described in 1884 by American paleontologist Othniel Charles Marsh based on a nearly complete skeleton discovered in Garden Park, Colorado, in rocks belonging to the Morrison Formation. The type species is ''Ceratosaurus nasicornis''. The Garden Park specimen remains the most complete skeleton known from the genus, and only a handful of additional specimens have been described since. Two additional species, ''Ceratosaurus dentisulcatus'' and ''Ceratosaurus magnicornis'', were described in 2000 from two fragmentary skeletons from the Cleveland-Lloyd Quarry of Utah and from the vicinity of Fruita, Colorado. The validity of these additional species has been questioned, however, and all three skeletons possibly represent different growth stages of the same ...
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Abelisauridae
Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are found on the modern continents of Africa and South America, as well as on the Indian subcontinent and the island of Madagascar. Isolated teeth were found in the Late Jurassic of Portugal, and the Late Cretaceous genera ''Tarascosaurus'' and '' Arcovenator'' have been described in France. Abelisaurids first appear in the fossil record of the early middle Jurassic period, and at least two genera (the Moroccan '' Chenanisaurus'' and the Madagascan '' Majungasaurus'') survived until the end of the Mesozoic era 66 million years ago. Like most theropods, abelisaurids were carnivorous bipeds. They were characterized by stocky hind limbs and extensive ornamentation of the skull bones, with grooves and pits. In many abelisaurids, such as '' Carnotaur ...
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Ceratosaurus
''Ceratosaurus'' (from Greek κέρας/κέρατος, ' meaning "horn" and σαῦρος ' meaning "lizard") was a carnivorous theropod dinosaur in the Late Jurassic period ( Kimmeridgian to Tithonian). The genus was first described in 1884 by American paleontologist Othniel Charles Marsh based on a nearly complete skeleton discovered in Garden Park, Colorado, in rocks belonging to the Morrison Formation. The type species is ''Ceratosaurus nasicornis''. The Garden Park specimen remains the most complete skeleton known from the genus, and only a handful of additional specimens have been described since. Two additional species, ''Ceratosaurus dentisulcatus'' and ''Ceratosaurus magnicornis'', were described in 2000 from two fragmentary skeletons from the Cleveland-Lloyd Quarry of Utah and from the vicinity of Fruita, Colorado. The validity of these additional species has been questioned, however, and all three skeletons possibly represent different growth stages of the sa ...
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Fosterovenator
''Fosterovenator'' (meaning "Foster's hunter") is a genus of ceratosaur dinosaur known from the Late Jurassic Morrison Formation of Wyoming. The holotype is YPM VP 058267A, B, and C, a tibia with an articulated astragalus. An additional specimen is known, the paratype YPM VP 058267D, a fibula of a larger individual. The holotype remains were in 1879 discovered by Arthur Lakes at Como Bluff, Wyoming, and consist of a nearly-complete right tibia with a co-ossified astragalus, probably of a juvenile. The paratype consists of a complete right fibula measuring in length and belonging to a much larger individual. The overall shape of the known material is similar to that of ''Elaphrosaurus''. However, ceratosaurian affinities of ''Fosterovenator'' (at least of the paratype) have been questioned. Etymology The generic name ''Fosterovenator'' was in 2014 named by S. G. Dalman for John Russell Foster and the Latin word ''venator'' ("hunter"). The specific name ''churei'' is named for ...
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Saltriovenator
''Saltriovenator'' (meaning "Saltrio hunter") is a genus of ceratosaurian dinosaur that lived during the Sinemurian stage of the Early Jurassic in what is now Italy. The type and only species is ''Saltriovenator zanellai''; in the past, the species had been known under the informal name "saltriosaur". Although a full skeleton has not yet been discovered, ''Saltriovenator'' is thought to have been a large, bipedal carnivore similar to ''Ceratosaurus''. Discovery and naming On 4 August 1996, the first remains of ''Saltriovenator'' were discovered by amateur paleontologist Angelo Zanella, searching for ammonites in the ''Salnova'' marble quarry in Saltrio, northern Italy. Zanella had already been working for the ''Museo Civico di Storia Naturale di Milano'' and this institution after being informed sent out a team to investigate the find. Cristiano Dal Sasso and the volunteers of the Paleontological Group of Besano, under the direction of Giorgio Teruzzi managed to salvage a num ...
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Ceratosauridae
Ceratosauridae is an extinct family of theropod dinosaurs belonging to the infraorder Ceratosauria. The family's type genus, ''Ceratosaurus'', was first found in Jurassic rocks from North America. Ceratosauridae is made up of the genera ''Ceratosaurus'', found in North America, Tanzania, and Portugal, and '' Genyodectes'', from the Early Cretaceous of Argentina. Unnamed probable ceratosaurids are known from limited material in the Middle Jurassic of Madagascar, the Late Jurassic of Switzerland, the Late Jurassic of Tanzania, and the Late Jurassic or possibly Early Cretaceous of Uruguay. Classification Othniel Charles Marsh named the family Ceratosauridae in 1884 to contain the type species, ''Ceratosaurus nasicornis''. Since then, a number of other species have been referred to this family, mainly from the genus ''Ceratosaurus''. Currently recognized ceratosaurid species include ''Genyodectes serus'' and possibly ''Eoabelisaurus mefi,'' ''C. stechowi'' and ''Ostafrikasaurus ...
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Coelophysoidea
Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the coelurosaurs, with which they were formerly classified, and some species had delicate cranial crests. Sizes range from about 1 to 6 m in length. It is unknown what kind of external covering coelophysoids had, and various artists have portrayed them as either scaly or feathered. Some species may have lived in packs, as inferred from sites where numerous individuals have been found together. Examples of coelophysoids include ''Coelophysis'', ''Procompsognathus'' and '' Liliensternus''. Most dinosaurs formerly referred to as being in the dubious taxon "Podokesauridae" are now classified as coelophysoids. Classification Despite their very early occurrence in the fossil record, coelophysoids have a number of derived features that separate ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, ar ...
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Austrocheirus
''Austrocheirus'' is an extinct genus of theropod dinosaur, possibly a neoceratosaurian, which existed during the Late Cretaceous period. It was named and described by Martin Ezcurra, Federico Agnolin and Fernando Novas in 2010. It contains the type species ''Austrocheirus isasii''. The generic name means "southern hand". The specific epithet honours discoverer and preparator Marcelo Pablo Isasi. The fossils were found on 17 March 2002 in the Pari Aike Formation, which was originally dated to the Maastrichtian, around 71 to 66 million years ago, but was more recently dated to the Cenomanian, around 96.2 million years ago. However, this strata is now believed to date to the Campanian. These fossils consist of a partial manus (hand), a tibia, axial bones, and a foot bone. In 2016, its length was estimated to be . In 2012 Holtz estimated it at 6.5 meters (24 feet), while in 2016 Molina-Pérez & Larramendi gave a higher estimation at 9.3 meters (30.5 feet) in length and 1 tonne ...
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Deltadromeus
''Deltadromeus'' (meaning "delta runner") is a genus of theropod dinosaur from Northern Africa. It had long, unusually slender hind limbs for its size, suggesting that it was a swift runner. The skull is not known. One fossil specimen of a single species (''D. agilis'', or "agile delta runner") has been described, found in the Kem Kem Beds, which date to the mid-Cretaceous Period (mid-Cenomanian age), about 95 million years ago. It may be a junior synonym of the contemporary ''Bahariasaurus''.Holtz, Thomas R. Jr. (2008) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages'Supplementary Information/ref> The classification of ''Deltadromeus'' has been in flux since its original description. In 2016, a South American theropod known as '' Gualicho shinyae'' was found to possess many similarities with ''Deltadromeus''. Depending on the phylogenetic position of ''Gualicho'', ''Deltadromeus'' may have been a neovenatorid carnosaur, a tyrannosauroi ...
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