Flyingfish
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The Exocoetidae are a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of
marine fish Saltwater fish, also called marine fish or sea fish, are fish that live in seawater. Saltwater fish can swim and live alone or in a large group called a school. Saltwater fish are very commonly kept in aquariums for entertainment. Many saltwater f ...
in the order
Beloniformes Beloniformes is an order composed of six families (and about 264 species) of freshwater and marine ray-finned fish: * Adrianichthyidae (ricefish and medakas) * Belonidae (needlefish) * Exocoetidae (flyingfishes) * Hemiramphidae (halfbeaks) ...
class Class or The Class may refer to: Common uses not otherwise categorized * Class (biology), a taxonomic rank * Class (knowledge representation), a collection of individuals or objects * Class (philosophy), an analytical concept used differentl ...
Actinopterygii, known colloquially as flying fish or flying cod. About 64
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
are grouped in seven to nine genera. While they cannot fly in the same way a bird does, flying fish can make powerful, self-propelled leaps out of the water where their long wing-like fins enable gliding for considerable distances above the water's surface. The main reason for this behavior is thought to be to escape from underwater predators, which include swordfish, mackerel,
tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae ( mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max len ...
, and
marlin Marlins are fish from the family Istiophoridae, which includes about 10 species. A marlin has an elongated body, a spear-like snout or bill, and a long, rigid dorsal fin which extends forward to form a crest. Its common name is thought to deri ...
, among others, though their periods of flight expose them to attack by avian predators such as
frigate bird Frigatebirds are a family of seabirds called Fregatidae which are found across all tropical and subtropical oceans. The five extant species are classified in a single genus, ''Fregata''. All have predominantly black plumage, long, deeply forked ...
s.
Barbados Barbados is an island country in the Lesser Antilles of the West Indies, in the Caribbean region of the Americas, and the most easterly of the Caribbean Islands. It occupies an area of and has a population of about 287,000 (2019 estimate) ...
is known as "the land of the flying fish", and the fish is one of the national symbols of the country. The
Exocet The Exocet () is a French-built anti-ship missile whose various versions can be launched from surface vessels, submarines, helicopters and fixed-wing aircraft. Etymology The missile's name was given by M. Guillot, then the technical director ...
missile is named after them, as variants are launched from underwater, and take a low trajectory, skimming the surface, before striking their targets.


Etymology

The term Exocoetidae is both the scientific name and the general name in
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
for a flying fish. The suffix ''-idae'', common for indicating a family, follows the root of the Latin word ', a transliteration of the
Ancient Greek Ancient Greek includes the forms of the Greek language used in ancient Greece and the ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Dark Ages (), the Archaic p ...
name . This means literally "sleeping outside", from "outside" and "bed", "resting place", verb root κει- "to lie down" (not "untruth"), so named as flying fish were believed to leave the water to sleep ashore, or due to flying fish flying and thus stranding themselves in boats.


Taxonomy

The Exocoetidae is divided into four subfamilies and seven genera: * Subfamily
Exocoetinae ''Exocoetus'' is a genus of flying fishes. It is a bony fish. The body is covered with cycloid scales. The mouth is wide, and the jaws bear teeth. It is a marine fish. The tail has hypobatic fins as the ventral lobe. Species Five species in thi ...
Risso, 1827 ** Genus ''
Exocoetus ''Exocoetus'' is a genus of flying fishes. It is a bony fish. The body is covered with cycloid scales. The mouth is wide, and the jaws bear teeth. It is a marine fish. The tail has hypobatic fins as the ventral lobe. Species Five species in thi ...
''
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, 1758
* Subfamily
Fodiatorinae ''Fodiator'' is a genus of flying fishes. It is the only genus in the subfamily Fodiatorinae. Etymology ''Fodiator'' means "one who stab", likely a reference to the long, thin, sharp snout with its projecting lower jaw. About ''Fodiator'' speci ...
Fowler, 1925 ** Genus ''
Fodiator ''Fodiator'' is a genus of flying fishes. It is the only genus in the subfamily Fodiatorinae. Etymology ''Fodiator'' means "one who stab", likely a reference to the long, thin, sharp snout with its projecting lower jaw. About ''Fodiator'' speci ...
'' D.S. Jordan &
Meek Meekness is an attribute of human nature and behavior that has been defined as an amalgam of righteousness, inner humility, and patience. Meekness has been contrasted with humility alone insomuch as humility simply refers to an attitude towards o ...
, 1885
* Subfamily
Parexocoetinae ''Parexocoetus'' is a genus of flying fishes. They are found in the tropical and subtropical western Atlantic and Indo-Pacific Oceans. It is the only genus in the subfamily Parexocoetinae which is unique among the flying fishes in having a ja ...
Bruun, 1935 ** Genus ''
Parexocoetus ''Parexocoetus'' is a genus of flying fishes. They are found in the tropical and subtropical western Atlantic and Indo-Pacific Oceans. It is the only genus in the subfamily Parexocoetinae which is unique among the flying fishes in having a jaw wh ...
'' Bleeker, 1865 * Subfamily
Cypsellurinae Cypsellurinae is a subfamily of flying fishes, one of four in the family Exocoetidae and the only one which is not monogeneric. Genera The following four genera make of the subfamily: * '' Cheilopogon'' Lowe, 1841 * ''Cypselurus ''Cypselurus ...
Hubbs, 1933 ** Genus '' Cheilopogon'' Lowe, 1841 ** Genus ''
Cypselurus ''Cypselurus'' is a genus of flying fish in the family Exocoetidae. Species Currently, there are twelve recognized species in this genus: * '' Cypselurus angusticeps'' ( Nichols & Breder, 1935) — narrowhead flying fish * '' Cypselurus callop ...
'' Swainson, 1838 ** Genus '' Hirundichthys'' Breder, 1928 ** Genus '' Prognichthys'' Breder, 1928


Distribution and description

Flying fish live in all of the
oceans The ocean (also the sea or the world ocean) is the body of salt water that covers approximately 70.8% of the surface of Earth and contains 97% of Earth's water. An ocean can also refer to any of the large bodies of water into which the worl ...
, particularly in
tropical The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in the Southern Hemisphere at S. The tropics are also referred to ...
and warm subtropical waters. They are commonly found in the
epipelagic zone The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological proc ...
, the top layer of the ocean to a depth of about 200 m (656 ft). Numerous morphological features give flying fish the ability to leap above the surface of the ocean. One such feature is fully broadened
neural arches The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
, which act as insertion sites for connective tissues and ligaments in a fish's skeleton. Fully broadened neural arches act as more stable and sturdier sites for these connections, creating a strong link between the
vertebral column The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordate ...
and cranium. This ultimately allows a rigid and sturdy vertebral column (body) that is beneficial in flight. Having a rigid body during glided flight gives the flying fish
aerodynamic Aerodynamics, from grc, ἀήρ ''aero'' (air) + grc, δυναμική (dynamics), is the study of the motion of air, particularly when affected by a solid object, such as an airplane wing. It involves topics covered in the field of fluid dyn ...
advantages, increasing its speed and improving its aim. Furthermore, flying fish have developed vertebral columns and
ossified Ossification (also called osteogenesis or bone mineralization) in bone remodeling is the process of laying down new bone material by cells named osteoblasts. It is synonymous with bone tissue formation. There are two processes resulting in t ...
caudal complexes. These features provide the majority of strength to the flying fish, allowing them to physically lift their bodies out of water and glide remarkable distances. These additions also reduce the flexibility of the flying fish, allowing them to perform powerful leaps without weakening midair. At the end of a glide, they fold their
pectoral fins Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as see ...
to re-enter the sea, or drop their tails into the water to push against the water to lift for another glide, possibly changing direction. The curved profile of the "wing" is comparable to the aerodynamic shape of a bird wing. The fish is able to increase its time in the air by flying straight into or at an angle to the direction of
updraft In meteorology, an updraft is a small-scale current of rising air, often within a cloud. Overview Localized regions of warm or cool air will exhibit vertical movement. A mass of warm air will typically be less dense than the surrounding regi ...
s created by a combination of
air The atmosphere of Earth is the layer of gases, known collectively as air, retained by Earth's gravity that surrounds the planet and forms its planetary atmosphere. The atmosphere of Earth protects life on Earth by creating pressure allowing f ...
and
ocean current An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth contours ...
s. Species of genus ''
Exocoetus ''Exocoetus'' is a genus of flying fishes. It is a bony fish. The body is covered with cycloid scales. The mouth is wide, and the jaws bear teeth. It is a marine fish. The tail has hypobatic fins as the ventral lobe. Species Five species in thi ...
'' have one pair of fins and streamlined bodies to optimize for speed, while ''
Cypselurus ''Cypselurus'' is a genus of flying fish in the family Exocoetidae. Species Currently, there are twelve recognized species in this genus: * '' Cypselurus angusticeps'' ( Nichols & Breder, 1935) — narrowhead flying fish * '' Cypselurus callop ...
'' spp. have flattened bodies and two pairs of fins, which maximize their time in the air. From 1900 to the 1930s, flying fish were studied as possible models used to develop airplanes. The Exocoetidae feed mainly on
plankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a crucia ...
. Predators include
dolphin A dolphin is an aquatic mammal within the infraorder Cetacea. Dolphin species belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontoporiidae (the ...
s,
tuna A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae ( mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max len ...
,
marlin Marlins are fish from the family Istiophoridae, which includes about 10 species. A marlin has an elongated body, a spear-like snout or bill, and a long, rigid dorsal fin which extends forward to form a crest. Its common name is thought to deri ...
,
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s, squid, and porpoises.


Flight measurements

In May 2008, a Japanese television crew (
NHK , also known as NHK, is a Japanese public broadcaster. NHK, which has always been known by this romanized initialism in Japanese, is a statutory corporation funded by viewers' payments of a television license fee. NHK operates two terrestr ...
) filmed a flying fish (dubbed "Icarfish") off the coast of
Yakushima Island , native_name_link = , image_caption = Landsat image of Yakushima , image_size = , nickname = , location = East China Sea , coordinates = , map = Japan#Japan Kagoshima Prefecture , map_relief ...
, Japan. The fish spent 45 seconds in flight. The previous record was 42 seconds. The flights of flying fish are typically around 50 m (160 ft),
Ross Piper Ross Piper is a British zoologist, entomologist, and explorer. Biography Piper's fascination by animals began at a young age through early encounters with a violet ground beetle and the caterpillar of an elephant hawk moth. This early interes ...
(2007), ''Extraordinary Animals: An Encyclopedia of Curious and Unusual Animals'', Greenwood Press.
though they can use updrafts at the leading edge of waves to cover distances up to . They can travel at speeds of more than . Maximum altitude is above the surface of the sea. Flying fish often accidentally land on the decks of smaller vessels.


Fishery and cuisine

Flying fish are commercially fished in Japan,
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...
, and China by
gillnetting Gillnetting is a fishing method that uses gillnets: vertical panels of netting that hang from a line with regularly spaced floaters that hold the line on the surface of the water. The floats are sometimes called "corks" and the line with corks is ...
, and in
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
and
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
by dipnetting. Often in
Japanese cuisine Japanese cuisine encompasses the regional and traditional foods of Japan, which have developed through centuries of political, economic, and social changes. The traditional cuisine of Japan (Japanese: ) is based on rice with miso soup and oth ...
, the fish is preserved by drying to be used as fish stock for
dashi is a family of stocks used in Japanese cuisine. ''Dashi'' forms the base for miso soup, clear broth soup, noodle broth soup, and many simmering liquids to accentuate the savory flavor known as umami. ''Dashi'' is also mixed into the flour b ...
broth. The
roe Roe ( ) or hard roe is the fully ripe internal egg masses in the ovaries, or the released external egg masses, of fish and certain marine animals such as shrimp, scallop, sea urchins and squid. As a seafood, roe is used both as a cooked in ...
of ''
Cheilopogon agoo ''Cheilopogon'' is a genus of flyingfishes. Species Currently, 29 species in this genus are recognized: * '' Cheilopogon abei'' Parin, 1996 (Abe's flyingfish) * '' Cheilopogon agoo'' (Temminck & Schlegel, 1846) (Japanese flyingfish) * '' Cheilo ...
'', or Japanese flying fish, is used to make some types of
sushi is a Japanese dish of prepared , usually with some sugar and salt, accompanied by a variety of , such as seafood, often raw, and vegetables. Styles of sushi and its presentation vary widely, but the one key ingredient is "sushi rice," also ...
, and is known as ''
tobiko is flying fish roe in Japanese cuisine, known for its use in ''sushi''. The eggs are small, ranging from 0.5 to 0.8 mm. For comparison, ''tobiko'' is larger than ''masago'' (capelin roe), but smaller than ''ikura'' (salmon roe). Natural ...
''. It is also a staple in the diet of the
Tao people The Tao people are an Austronesian ethnic group native to the tiny outlying Orchid Island of Taiwan. They have a maritime culture, with great ritual and spiritual significance placed on boat-building and fishing. Their ways of life have been th ...
of
Orchid Island Orchid Island, also known by other names, is a volcanic island off the southeastern coast of Taiwan Island. The island is part of Taiwan. It is separated from the Batanes of the Philippines by the Bashi Channel of the Luzon Strait. It is g ...
,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...
. Flying fish is part of the national dish of Barbados, ''
cou-cou Cou-cou, coo-coo (as it is known in the Windward Islands), or fungie (as it is known in the Leeward Islands and Dominica) makes up part of the national dishes of Antigua and Barbuda, Barbados, British Virgin Islands and the U.S. Virgin Islands. ...
'' and flying fish. The taste is close to that of a sardine. Flying fish roe is known as cau-cau in southern Peru, and is used to make several local dishes. In the
Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capit ...
, the fish are caught while they are flying, using nets held from
outrigger canoe Outrigger boats are various watercraft featuring one or more lateral support floats known as outriggers, which are fastened to one or both sides of the main hull. They can range from small dugout canoes to large plank-built vessels. Outrigger ...
s. They are attracted to the light of torches. Fishing is done only when no moonlight is available.


Importance


Barbados

Barbados Barbados is an island country in the Lesser Antilles of the West Indies, in the Caribbean region of the Americas, and the most easterly of the Caribbean Islands. It occupies an area of and has a population of about 287,000 (2019 estimate) ...
is known as "the land of the flying fish", and the fish is one of the national symbols of the country. Once abundant, it migrated between the warm,
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and ...
-filled
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
surrounding the island of Barbados and the plankton-rich outflows of the Orinoco River in
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
. Just after the completion of the Bridgetown Harbor / Deep Water Harbor in Bridgetown, Barbados had an increase of ship visits, linking the island to the world. The overall health of the
coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in groups. C ...
s surrounding Barbados suffered due to ship-based
pollution Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the ...
. Additionally, Barbadian overfishing pushed them closer to the Orinoco
delta Delta commonly refers to: * Delta (letter) (Δ or δ), a letter of the Greek alphabet * River delta, at a river mouth * D ( NATO phonetic alphabet: "Delta") * Delta Air Lines, US * Delta variant of SARS-CoV-2 that causes COVID-19 Delta may also ...
, no longer returning to Barbados in large numbers. Today, the flying fish only migrate as far north as
Tobago Tobago () is an List of islands of Trinidad and Tobago, island and Regions and municipalities of Trinidad and Tobago, ward within the Trinidad and Tobago, Republic of Trinidad and Tobago. It is located northeast of the larger island of Trini ...
, around southwest of Barbados. Despite the change, flying fish remain a coveted delicacy. Many aspects of Barbadian culture center around the flying fish; it is depicted on coins, as sculptures in fountains, in artwork, and as part of the official logo of the Barbados Tourism Authority. Additionally, the Barbadian coat of arms features a pelican and
dolphinfish ''Coryphaena'' is a genus of marine ray-finned fishes known as the dolphinfishes, and is currently the only known genus in the family Coryphaenidae. The generic name is from Greek κορυφή (''koryphē'', "crown, top") and -αινα (-''aina' ...
on either side of the shield, but the dolphinfish resembles a flying fish. Furthermore, actual artistic renditions and
hologram Holography is a technique that enables a wavefront to be recorded and later re-constructed. Holography is best known as a method of generating real three-dimensional images, but it also has a wide range of other applications. In principle, i ...
s of the flying fish are also present within the
Barbadian passport A Barbados passport is a travel document issued to citizens of Barbados, in accordance with Citizenship Act (CAP. 186) from 1978, the Immigration Act (CAP. 190) from 1997, and the Barbados Constitution, for the purpose of facilitating internation ...
.


Maritime disputes

Flying fish have also been gaining in popularity in other islands, fueling several maritime disputes. In 2006, the council of the
United Nations Convention on the Law of the Sea The United Nations Convention on the Law of the Sea (UNCLOS), also called the Law of the Sea Convention or the Law of the Sea Treaty, is an international agreement that establishes a legal framework for all marine and maritime activities. , 167 c ...
fixed the
maritime boundaries A maritime boundary is a conceptual division of the Earth's water surface areas using physiographic or geopolitical criteria. As such, it usually bounds areas of exclusive national rights over mineral and biological resources,VLIZ Maritime Bound ...
between
Barbados Barbados is an island country in the Lesser Antilles of the West Indies, in the Caribbean region of the Americas, and the most easterly of the Caribbean Islands. It occupies an area of and has a population of about 287,000 (2019 estimate) ...
and
Trinidad and Tobago Trinidad and Tobago (, ), officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean. Consisting of the main islands Trinidad and Tobago, and numerous much smaller islands, it is situated south of ...
over the flying fish dispute, which gradually raised tensions between the neighbours. The ruling stated both countries must preserve stocks for the future. Barbadian fishers still follow the flying fish southward.


Prehistoric analogues

The oldest known fossil of a flying or gliding fish are those of the extinct family Thoracopteridae, dating back to the
Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and ...
, 235–242 million years ago. However, they are not related to modern flying fish, with the wing-like pectoral fins being convergently evolved in both lineages.


See also

*
Flying and gliding animals A number of animals are capable of aerial locomotion, either by powered flight or by gliding. This trait has appeared by evolution many times, without any single common ancestor. Flight has evolved at least four times in separate animals: inse ...
*
Neuston Neuston, also known as pleuston, are organisms that live at the surface of the ocean or an estuary, or at the surface of a lake, river or pond. Neuston can live on top of the water surface or may be attached to the underside of the water surface. ...
* Tradeoffs for locomotion in air and water


References


External links


Flying Fish, National Geographic Society
{{Authority control Exocoetidae Beloniformes Gliding animals Fish of Oceania Marine fish of Africa Marine fish of Asia Fish of Hawaii Fish of Japan Extant Miocene first appearances Miocene fish Quaternary fish Barbadian cuisine