Ficus Disticha
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''Ficus'' ( or ) is a genus of about 850 species of woody trees,
shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
s, vines,
epiphyte An epiphyte is an organism that grows on the surface of a plant and derives its moisture and nutrients from the air, rain, water (in marine environments) or from debris accumulating around it. The plants on which epiphytes grow are called phoroph ...
s and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species extending into the semi-warm temperate zone. The
common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the world ...
(''F. carica'') is a temperate species native to southwest Asia and the Mediterranean region (from Afghanistan to Portugal), which has been widely cultivated from ancient times for its fruit, also referred to as figs. The fruit of most other species are also edible though they are usually of only local economic importance or eaten as bushfood. However, they are extremely important food resources for wildlife. Figs are also of considerable cultural importance throughout the tropics, both as objects of worship and for their many practical uses.


Description

''Ficus'' is a pantropical genus of trees, shrubs, and vines occupying a wide variety of ecological niches; most are evergreen, but some deciduous species are found in areas outside of the tropics and to higher elevations. Fig species are characterized by their unique inflorescence and distinctive pollination syndrome, which uses wasp species belonging to the family Agaonidae for pollination. Specific identification of many of the species can be difficult, but members of the genus ''Ficus'' are relatively easy to recognize. Many have aerial roots and a distinctive shape or habit, and their fruits distinguish them from other plants. The fruit of ''Ficus'' is an inflorescence enclosed in an urn-like structure called a
syconium Syconium (plural ''syconia'') is the type of inflorescence borne by figs (genus ''Ficus''), formed by an enlarged, fleshy, hollow receptacle with multiple ovaries on the inside surface. In essence, it is really a fleshy stem with a number of flow ...
, which is lined on the inside with the fig's tiny flowers that develop into multiple ovaries on the inside surface. In essence, the fig fruit is a fleshy stem with multiple tiny flowers that fruit and coalesce. The unique fig pollination system, involving tiny, highly specific wasps, known as fig wasps that enter via
ostiole An ''ostiole'' is a small hole or opening through which algae or fungi release their mature spores. The word is a diminutive of "ostium", "opening". The term is also used in higher plants, for example to denote the opening of the involuted ...
these subclosed inflorescences to both pollinate and lay their own eggs, has been a constant source of inspiration and wonder to biologists. Notably, three vegetative traits together are unique to figs. All figs present a white to yellowish latex, some in copious quantities; the twig shows paired
stipules In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). Stipules are considered part of the anatomy of the leaf of a typical flowering plant, although in many speci ...
—or circular scars if the stipules have fallen off; the lateral veins at the base of the leaf are steep, forming a tighter angle with the midrib than the other lateral veins, a feature referred to as "triveined". No unambiguous older fossils of ''Ficus'' are known. However, current molecular clock estimates indicate that ''Ficus'' is a relatively ancient genus being at least 60 million years old, and possibly as old as 80 million years. The main radiation of
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
species, however, may have taken place more recently, between 20 and 40 million years ago. Some better-known species that represent the diversity of the genus include the
common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the world ...
, a small, temperate deciduous tree whose fingered fig leaf is well known in art and
iconography Iconography, as a branch of art history, studies the identification, description and interpretation of the content of images: the subjects depicted, the particular compositions and details used to do so, and other elements that are distinct fro ...
; the weeping fig (''F. benjamina''), a hemiepiphyte with thin, tough leaves on pendulous stalks adapted to its rain forest habitat; the rough-leaved sandpaper figs from Australia; and the creeping fig (''F. pumila''), a vine whose small, hard leaves form a dense carpet of foliage over rocks or garden walls. Moreover, figs with different plant habits have undergone adaptive radiation in different biogeographic regions, leading to very high levels of
alpha diversity In ecology, alpha diversity (α-diversity) is the mean species diversity in a site at a local scale. The term was introduced by R. H. WhittakerWhittaker, R. H. (1960) Vegetation of the Siskiyou Mountains, Oregon and California. Ecological Monograp ...
. In the tropics, ''Ficus'' commonly is the most species-rich plant genus in a particular forest. In Asia, as many as 70 or more species can co-exist. ''Ficus'' species richness declines with an increase in latitude in both hemispheres. A description of fig tree cultivation is set out in Ibn al-'Awwam's 12th-century agricultural work entitled, ''Book on Agriculture''.


Ecology

Figs are
keystone species A keystone species is a species which has a disproportionately large effect on its natural environment relative to its abundance, a concept introduced in 1969 by the zoologist Robert T. Paine. Keystone species play a critical role in maintaini ...
in many tropical forest ecosystems. Their fruit are a key resource for some frugivores including fruit bats, and primates including: capuchin monkeys, langurs,
gibbons Gibbons may refer to: * The plural of gibbon, an ape in the family Hylobatidae * Gibbons (surname) * Gibbons, Alberta * Gibbons (automobile), a British light car of the 1920s * Gibbons P.C., a leading American law firm headquartered in New Jersey ...
and mangabeys. They are even more important for birds such as Asian barbets, pigeons,
hornbill Hornbills (Bucerotidae) are a family (biology), family of bird found in tropical and subtropical Africa, Asia and Melanesia. They are characterized by a long, down-curved bill which is frequently brightly coloured and sometimes has a Casque (an ...
s, fig-parrots and
bulbul The bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropical As ...
s, which may almost entirely subsist on figs when these are in plenty. Many
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Sym ...
s feed on fig leaves, for example several '' Euploea'' species (crow butterflies), the plain tiger (''Danaus chrysippus''), the giant swallowtail (''Papilio cresphontes''), the
brown awl ''Badamia exclamationis'', commonly known as the brown awl or narrow-winged awl,Markku Savela's website on Lepidoptera. Page on genuTOL web page on genu''Badamia''/ref> is a butterfly belonging to the family Hesperiidae. It is found in south an ...
(''Badamia exclamationis''), and ''
Chrysodeixis eriosoma ''Chrysodeixis eriosoma'', the green garden looper, is a moth of the family Noctuidae. Mostly cosmopolitan in distribution, it is a pest in Japan, China, India, Sri Lanka, the Malay Peninsula and Australasia. It is present in Hawaii and recorded ...
'', Choreutidae and
Copromorphidae Copromorphidae, the "tropical fruitworm moths", is a family of insects in the lepidopteran order. These moths have broad, rounded forewings, and well-camouflaged scale patterns. Unlike Carposinidae the mouthparts include "labial palps" with the ...
moths. The
citrus long-horned beetle The citrus long-horned beetle (''Anoplophora chinensis'', also appearing in many sources as ''Anoplophora malasiaca'') is a long-horned beetle native to Japan, China, Korea, and Southeast Asia where it is considered a serious pest. Several cou ...
(''Anoplophora chinensis''), for example, has larvae that feed on wood, including that of fig trees; it can become a
pest Pest or The Pest may refer to: Science and medicine * Pest (organism), an animal or plant deemed to be detrimental to humans or human concerns ** Weed, a plant considered undesirable * Infectious disease, an illness resulting from an infection ** ...
in fig plantations. Similarly, the sweet potato whitefly (''Bemisia tabaci'') is frequently found as a pest on figs grown as potted plants and is spread through the export of these plants to other localities. For a list of other diseases common to fig trees, see List of foliage plant diseases (Moraceae).


Fig fruit and reproduction system

Many fig species are grown for their fruits, though only ''
Ficus carica The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the w ...
'' is cultivated to any extent for this purpose. A fig "fruit" is a type of
multiple fruit Multi-fruits, also called collective fruits, are fruiting bodies formed from a cluster of flowers, the ''inflorescence''. Each flower in the inflorescence produces a fruit, but these mature into a single mass. After flowering the mass is called a ...
known as a
syconium Syconium (plural ''syconia'') is the type of inflorescence borne by figs (genus ''Ficus''), formed by an enlarged, fleshy, hollow receptacle with multiple ovaries on the inside surface. In essence, it is really a fleshy stem with a number of flow ...
, derived from an arrangement of many small flowers on an inverted, nearly closed receptacle. The many small flowers are unseen unless the fig is cut open. The fruit typically has a bulbous shape with a small opening (the
ostiole An ''ostiole'' is a small hole or opening through which algae or fungi release their mature spores. The word is a diminutive of "ostium", "opening". The term is also used in higher plants, for example to denote the opening of the involuted ...
) at the outward end that allows access to
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are the maj ...
s. The flowers are pollinated by very small wasps that crawl through the opening in search of a suitable place to lay eggs. Without this pollinator service fig trees could not reproduce by seed. In turn, the flowers provide a safe haven and nourishment for the next generation of wasps. This accounts for the frequent presence of wasp larvae in the fruit, and has led to a
coevolution In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well ...
ary relationship. Technically, a fig fruit proper would be only one of the many tiny matured, seed-bearing
gynoecia Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
found inside one fig – if you cut open a fresh fig, individual fruit will appear as fleshy "threads", each bearing a single seed inside. The genus '' Dorstenia'', also in the fig family (Moraceae), exhibits similar tiny flowers arranged on a receptacle but in this case the receptacle is a more or less flat, open surface. Fig plants can be
monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...
(
hermaphrodite In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes. Many Taxonomy (biology), taxonomic groups of animals (mostly invertebrate ...
) or gynodioecious (hermaphrodite and female). Nearly half of fig species are gynodioecious, and therefore have some plants with inflorescences (syconium) with long styled pistillate flowers, and other plants with staminate flowers mixed with short styled pistillate flowers. The long-styled flowers tend to prevent wasps from laying their eggs within the ovules, while the short-styled flowers are accessible for egg laying. All the native fig trees of the American continent are hermaphrodites, as well as species like Indian banyan (''F. benghalensis''), weeping fig (''F. benjamina''), Indian rubber plant (''F. elastica''),
fiddle-leaved fig ''Ficus lyrata'', commonly known as the fiddle-leaf fig, is a species of flowering plant in the mulberry and fig family Moraceae. It is native to western Africa, from Cameroon west to Sierra Leone, where it grows in lowland tropical rainforest. ...
(''F. lyrata''), Moreton Bay fig (''F. macrophylla''), Chinese banyan (''F. microcarpa''), sacred fig (''F. religiosa'') and sycamore fig (''F. sycomorus''). The common fig (''Ficus carica'') is a gynodioecious plant, as well as
lofty fig Lofty may refer to: Places * Mount Lofty (disambiguation), several places and associated subjects in Australia People * Lofty Blomfield, (1908–1971), New Zealand professional wrestler * Lofty Drews (born 1940), World Rally Championship co-dri ...
or clown fig ('' F. aspera''),
Roxburgh fig ''Ficus auriculata'', the Roxburgh fig, is a type of fig tree, native to Asia, noted for its big and round leaves. Description This plant is a small tree of high with numerous bristle-covered branches. The leaves are big and round, and are up t ...
('' F. auriculata''), mistletoe fig ('' F. deltoidea''), '' F. pseudopalma'', creeping fig (''F. pumila'') and related species. The hermaphrodite common figs are called "inedible figs" or "caprifigs"; in traditional culture in the Mediterranean region they were considered food for goats (''Capra aegagrus''). In the female fig trees, the male flower parts fail to develop; they produce the "'edible figs". Fig wasps grow in common fig caprifigs but not in the female syconiums because the female flower is too long for the wasp to successfully lay her eggs in them. Nonetheless, the wasp pollinates the flower with pollen from the caprifig it grew up in. When the wasp dies, it is broken down by enzymes ( Ficain) inside the fig. Fig wasps are not known to transmit any diseases harmful to humans. When a caprifig ripens, another caprifig must be ready to be pollinated. In temperate climes, wasps hibernate in figs, and there are distinct crops. Caprifigs have three crops per year; common figs have two. The first crop ( breba) is larger and juicier, and usually eaten fresh. In cold climates the breba crop is often destroyed by spring frosts. Some parthenocarpic cultivars of common figs do not require pollination at all, and will produce a crop of figs (albeit
sterile Sterile or sterility may refer to: *Asepsis, a state of being free from biological contaminants * Sterile (archaeology), a sediment deposit which contains no evidence of human activity *Sterilization (microbiology), any process that eliminates or ...
) in the absence of caprifigs and fig wasps. Depending on the species, each fruit can contain hundreds or even thousand of seeds. Figs can be propagated by seeds, cuttings, air-layering or grafting. However, as with any plant, figs grown from seed are not necessarily genetically identical to the parent and are only propagated this way for breeding purposes.


Mutualism with the pollinating fig wasps

Each species of fig is pollinated by one or a few specialised wasp species, and therefore plantings of fig species outside of their native range results in effectively sterile individuals. For example, in Hawaii, some 60 species of figs have been introduced, but only four of the wasps that fertilize them, so only those species of figs produce viable seeds there and can become
invasive species An invasive species otherwise known as an alien is an introduced organism that becomes overpopulated and harms its new environment. Although most introduced species are neutral or beneficial with respect to other species, invasive species ad ...
. This is an example of mutualism, in which each organism (fig plant and fig wasp) benefit each other, in this case reproductively. The intimate association between fig species and their wasp pollinators, along with the high incidence of a one-to-one plant-pollinator ratio have long led scientists to believe that figs and wasps are a clear example of
coevolution In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well ...
. Morphological and reproductive behavior evidence, such as the correspondence between fig and wasp larvae maturation rates, have been cited as support for this hypothesis for many years. Additionally, recent genetic and molecular dating analyses have shown a very close correspondence in the character evolution and
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...
phylogenies of these two clades. According to meta-analysis of molecular data for 119 fig species 35% (41) have multiple pollinator wasp species. The real proportion is higher because not all wasp species were detected. On the other hand, species of wasps pollinate multiple host fig species. Molecular techniques, like
microsatellite A microsatellite is a tract of repetitive DNA in which certain DNA motifs (ranging in length from one to six or more base pairs) are repeated, typically 5–50 times. Microsatellites occur at thousands of locations within an organism's genome. ...
markers and mitochondrial sequence analysis, allowed a discovery of multiple genetically distinct,
cryptic Cryptic may refer to: In science: * Cryptic species complex, a group of species that are very difficult to distinguish from one another * Crypsis, the ability of animals to blend in to avoid observation * Cryptic era, earliest period of the Earth ...
wasp species. Not all these cryptic species are sister taxa and thus must have experienced a host fig shift at some point. These cryptic species lacked evidence of genetic
introgression Introgression, also known as introgressive hybridization, in genetics is the transfer of genetic material from one species into the gene pool of another by the repeated backcrossing of an interspecific hybrid with one of its parent species. Intr ...
or backcrosses indicating limited fitness for hybrids and effective
reproductive isolation The mechanisms of reproductive isolation are a collection of evolutionary mechanisms, behaviors and physiological processes critical for speciation. They prevent members of different species from producing offspring, or ensure that any offspring ...
and
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...
. The existence of cryptic species suggests that neither the number of symbionts nor their evolutionary relationships are necessarily fixed ecologically. While the morphological characteristics that facilitate the fig-wasp mutualisms are likely to be shared more fully in closer relatives, the absence of unique pairings would make it impossible to do a one-to-one tree comparison and difficult to determine cospeciation.


Systematics

With 800 species, ''Ficus'' is by far the largest genus in the Moraceae, and is one of the largest genera of flowering plants currently described. The species currently classified within ''Ficus'' were originally split into several genera in the mid-1800s, providing the basis for a subgeneric classification when reunited into one genus in 1867. This classification put functionally
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...
species into four subgenera based on floral characters. In 1965, E. J. H. Corner reorganized the genus on the basis of breeding system, uniting these four dioecious subgenera into a single dioecious subgenus ''Ficus''.
Monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system alongside gynomonoecy, andromonoecy and trimonoecy. Monoecy is conne ...
figs were classified within the subgenera ''
Urostigma A banyan, also spelled "banian", is a fig that develops accessory trunks from adventitious prop roots, allowing the tree to spread outwards indefinitely. This distinguishes banyans from other trees with a strangler habit that begin life as a ...
'', ''
Pharmacosycea ''Pharmacosycea'' is one of six subgenera currently recognised in the genus ''Ficus''. It was proposed by E. J. H. Corner in 1967 to unite section ''Pharmacosycea'' with ''Oreosycea''. Recent molecular phylogenies has shown that the subgenus i ...
'' and '' Sycomorus''. This traditional classification has been called into question by recent phylogenetic studies employing genetic methods to investigate the relationships between representative members of the various sections of each subgenus. Of Corner's original subgeneric divisions of the genus, only ''Sycomorus'' is supported as monophyletic in the majority of phylogenetic studies. Notably, there is no clear split between dioecious and monoecious lineages. One of the two sections of ''Pharmacosycea'', a monoecious group, form a monophyletic clade basal to the rest of the genus, which includes the other section of ''Pharmacosycea'', the rest of the monoecious species, and all of the dioecious species. These remaining species are divided into two main monophyletic lineages (though the statistical support for these lineages isn't as strong as for the monophyly of the more derived clades within them). One consists of all sections of ''Urostigma'' except for section ''Urostigma s. s.''. The other includes section ''Urostigma s. s.'', subgenus ''Sycomorus'', and the species of subgenus ''Ficus'', though the relationships of the sections of these groups to one another are not well resolved.


Selected species

There are 875 accepted ''Ficus'' species, as of March 2021, according to
Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...
.


Subgenus ''Ficus''

*'' Ficus amplissima'' Sm. – bat fig *''
Ficus carica The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the w ...
'' L. – common fig *''
Ficus daimingshanensis ''Ficus daimingshanensis'' is a plant species native to the Chinese provinces of Guangxi and Hunan. It grows on limestone soils at elevations of approximately . Type locality iDaming Shan a mountain in Guangxi Province near Dafeng Dafeng () ...
'' Chang *''
Ficus deltoidea ''Ficus deltoidea'', commonly known as mistletoe fig (''mas cotek'' in Malaysian, ''tabat barito'' in Indonesia, or ''สาลิกาลิ้นทอง'' in Thai) is a species of flowering plant in the family Moraceae, native to Southeast ...
'' Jack – mistletoe fig *''
Ficus erecta ''Ficus erecta'' (syn. ''Ficus beecheyana''), the Japanese fig, is a species of flowering plant in the family Moraceae. It is found in the eastern Himalayas, Assam, Bangladesh, Vietnam, southern China, Taiwan, Jeju Island of South Korea, the Ryu ...
'' Thunb. – Japanese fig *'' Ficus fulva'' Reinw. ex Blume *'' Ficus grossularioides'' Burman f. – white-leaved fig *'' Ficus neriifolia'' Sm. *'' Ficus palmata'' Forssk. *''
Ficus pandurata ''Ficus pandurata''Hance HF (1862) ''Ann. Sci. Nat., Bot., sér. 4'', 18: 229. is a fig species in the family Moraceae The Moraceae — often called the mulberry family or fig family — are a family of flowering plants comprising about 38 gen ...
'' Hance *'' Ficus simplicissima'' Lour. (synonym '' Ficus hirta'' Vahl) *''
Ficus triloba ''Ficus triloba''Voigt JO (1845) In: ''Hort. Suburb. Calcutt.'': 284. is an Asian species of fig tree in the family Moraceae. It is dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct indivi ...
'' Buch.-Ham. ex Voigt


Subgenus ''Pharmacosycea''

*''
Ficus crassiuscula ''Ficus crassiuscula'' is a species of flowering plant in the family Moraceae, native to Central America (Guatemala, Honduras, Belize, Nicaragua, Costa Rica, Panama) and north-western parts of South America (Colombia, Ecuador, Peru and Venezuela) ...
'' Standl. *''
Ficus gigantosyce ''Ficus gigantosyce'' is a species of tree in the family Moraceae. It is found in South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in ...
'' Dugand *''
Ficus insipida ''Ficus insipida'' is a common tropical tree in the fig genus of the family Moraceae growing in forest habitats along rivers. It ranges from Mexico to northern South America. Taxonomy The tree was described in 1806 under the scientific na ...
'' Willd. *''
Ficus lacunata ''Ficus lacunata'' is a species of plant in the family Moraceae which is endemic to Ecuador. ''F. lacunata'' is a free-standing tree which grows up to 25 m (82 ft) tall in wet forests in the Andes. ''Ficus lacunata'' is member of the ...
'' Kvitvik *'' Ficus maxima'' Mill. *'' Ficus mutabilis'' Bureau *'' Ficus nervosa'' Heyne ex Roth *'' Ficus pulchella'' Schott *''
Ficus yoponensis ''Ficus yoponensis'' is a species of fig tree found in Central and South America. It can grow to heights of tall, having a trunk diameter of . The trunk is buttressed, light grey in colour and reasonably smooth. Its petioles are long, the stip ...
'' Desv.


Subgenus ''Sycidium''

*'' Ficus andamanica'' Corner *'' Ficus aspera'' G.Forst. *''
Ficus assamica ''Ficus assamica'' is a species of flowering plant in the genus ''Ficus'', native to the foothills of the Himalayas, Assam, south-central China, and mainland Southeast Asia. It is a recumbent or creeping shrub reaching at most 60cm. Some authori ...
'' Miq. *'' Ficus bojeri'' Baker *''
Ficus capreifolia The river sandpaper fig (''Ficus capreifolia'') is a fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated sin ...
'' Delile *''
Ficus coronata ''Ficus coronata'', commonly known as the sandpaper fig or creek sandpaper fig, is a species of fig tree, native to Australia. It is found along the east coast from Mackay in Central Queensland, through New South Wales and just into Victoria ne ...
'' Spin – creek sandpaper fig *'' Ficus fraseri'' Miq. – shiny sandpaper fig *'' Ficus heterophylla'' L.f. *''
Ficus lateriflora ''Ficus lateriflora'' is a species of fig tree in the family Moraceae. It is endemic to Mauritius and Réunion, islands off the coast of East Africa in the Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic div ...
'' Vahl *'' Ficus montana'' Burm.f. – oakleaf fig *''
Ficus opposita ''Ficus opposita'' is one of several fig species commonly known as sandpaper figs. It is native to the Northern Territory and Queensland in Australia.Other common names include sweet sandpaper fig, sweet fig and the ambiguous "figwood" and "wate ...
'' Miq. – sweet sandpaper fig *'' Ficus phaeosyce'' K.Schum. & Lauterb. *'' Ficus tinctoria'' G.Forst. – dye fig *''
Ficus ulmifolia ''Ficus ulmifolia'' is a species of plant in the family Moraceae. It is endemic to the Philippines. It is threatened by habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural hab ...
'' Lam. *'' Ficus wassa'' Roxb. *'' Ficus parietalis'' *'' Ficus sinuata'' *''
Ficus hampelas ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species extending int ...
''


Subgenus ''Sycomorus''

*''
Ficus auriculata ''Ficus auriculata'', the Roxburgh fig, is a type of fig tree, native to Asia, noted for its big and round leaves. Description This plant is a small tree of high with numerous bristle-covered branches. The leaves are big and round, and are up t ...
'' Lour. – Roxburgh fig *'' Ficus bernaysii'' King *'' Ficus dammaropsis'' Diels – highland breadfruit, ''kapiak'' *''
Ficus fistulosa ''Ficus fistulosa'' is an Asian species of fig tree in the family Moraceae. No subspecies are listed in the Catalogue of Life; its native range is Assam to Taiwan, Indo-China, Malesia and New Guinea. The species can be found in Vietnam: where it ...
'' Blume *''
Ficus hispida ''Ficus hispida'' also known as the opposite leaf Fig is a small but well distributed species of tropical Ficus, fig tree. It is dioecious, with male and female flowers on separate individuals. It occurs in many parts of Asia and as far south e ...
'' L. *'' Ficus nota'' Merr. – ''tibig'' *'' Ficus pseudopalma'' Blanco *'' Ficus racemosa'' L. – cluster fig *'' Ficus septica'' Burm.f. – hauli tree *'' Ficus sycomorus'' L., 1753 – sycamore fig (Africa) *'' Ficus variegata'' Blume


Subgenus ''Synoecia''

The following species are typically spreading or climbing
liana A liana is a long- stemmed, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the canopy in search of direct sunlight. The word ''liana'' does not refer to a ta ...
s: *'' Ficus hederacea'' Roxb. *''
Ficus pantoniana ''Ficus pantoniana'', commonly known as the climbing fig, is a species of fig tree, native to Australia. It is found in lowland rainforests on the Cape York Peninsula in northern Queensland. It is a woody climber which may reach 3–6 m in hei ...
'' King – climbing fig *'' Ficus pumila'' L. – creeping fig ** ''Ficus pumila'' var. ''awkeotsang'' (Makino) Corner – jelly fig *''
Ficus punctata ''Ficus punctata'' is a lesser-known climbing liana reported from Myanmar, the Andaman-Nicobar Archipelago, Thailand, Indochina, Malaysia, and Indonesia with remarkable ornamental value. It is a handsome woody liana with spectacular spotted ...
'' Thunb. *''
Ficus sagittata ''Ficus sagittata''König J, ex Vahl M (1805) In: ''Symb. Bot.'' 1: 83. is a trailing fig species, in the family Moraceae, which can be found in southern China, Indo-China and Malesia Malesia is a biogeographical region straddling the Equa ...
'' J. König ex Vahl *'' Ficus sarmentosa'' Buch.-Ham. ex Sm. *'' Ficus trichocarpa'' Blume *''
Ficus villosa ''Ficus villosa'', known as the shaggy-leaf fig or villous fig, is a species of ''Ficus'' native to South East Asia. Etymology The species epithet "villosa" is derived from Latin ''villōsa'' "hairy", from ''villus'' "hair" and the adjective ...
'' Blume


Subgenus ''

Urostigma A banyan, also spelled "banian", is a fig that develops accessory trunks from adventitious prop roots, allowing the tree to spread outwards indefinitely. This distinguishes banyans from other trees with a strangler habit that begin life as a ...
''

*'' Ficus abutilifolia'' Miq. *'' Ficus albert-smithii'' Standl. *'' Ficus altissima'' Blume *''
Ficus amazonica ''Ficus amazonica'' is a species of flowering plant in the family Moraceae. It is found in Brazil, Guyana, Trinidad and Tobago, and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Boliv ...
'' Miq. *'' Ficus americana'' Aubl. *''
Ficus aripuanensis ''Ficus aripuanensis'' is a species of plant in the family Moraceae. It is endemic to Mato Grosso and Pará states in Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in bo ...
'' Berg & Kooy *'' Ficus arpazusa'' Carauta and Diaz – Brazil *'' Ficus aurea'' Nutt. – Florida
strangler fig Strangler fig is the common name for a number of tropical and subtropical plant species in the genus ''Ficus'', including those that are commonly known as banyans. Some of the more well-known species are: * ''Ficus altissima'' * ''Ficus aurea'', ...
*''
Ficus beddomei ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species exten ...
'' King – ''thavital'' *'' Ficus benghalensis'' L. – Indian banyan *'' Ficus benjamina'' L. – weeping fig *'' Ficus binnendijkii'' Miq. *'' Ficus bizanae'' Hutch. & Burtt-Davy *''
Ficus blepharophylla ''Ficus blepharophylla'' is a species of fig tree in the family Moraceae. The tree is endemic to Roraima state in northern Brazil. It is an IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Spe ...
'' Vázquez Avila *''
Ficus broadwayi ''Ficus broadwayi'' is a species of plant in the family Moraceae. It is found in Brazil, Colombia, Suriname, and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuel ...
'' Urb. *''
Ficus burtt-davyi ''Ficus burtt-davyi'' is a fig species endemic to Southern Africa, belonging to the Mulberry family of Moraceae. It grows in coastal and inland forests up to 1500m, from the vicinity of Mossel Bay in the Southern Cape to southern Mozambique ...
'' Hutch. *'' Ficus calyptroceras'' Miq. *'' Ficus castellviana'' Dugand *''
Ficus catappifolia ''Ficus catappifolia'' is a species of plant in the family Moraceae. It is found in Brazil and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on ...
'' Kunth & Bouché *'' Ficus citrifolia'' Mill. – short-leaved fig *''
Ficus consociata ''Ficus consociata'' is a banyan fig species in the family Moraceae. No subspecies are listed in the Catalogue of Life. The species can be found in Indo-China and western Malesia. In Vietnam it may be called ''đa đồng hành''. Its common ...
'' Bl. *'' Ficus cordata'' Thunb. *''
Ficus costata ''Ficus costata'' is an Asian species of fig tree in the family Moraceae. No subspecies are listed in the Catalogue of Life The Catalogue of Life is an online database that provides an index of known species of animals, plants, fungi, and m ...
'' Ait. *''
Ficus crassipes ''Ficus crassipes'', commonly known as the round-leaved banana fig is a fig that is endemic to the wet tropical rainforests of northeastern Queensland, Australia. It has large brownish cylindrical syconia. Description ''Ficus crassipes'' is a ...
'' F.M.Bailey – round-leaved banana fig *''
Ficus craterostoma ''Ficus craterostoma'', a species of strangler fig, is a fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated ...
'' Mildbr. & Burret *''
Ficus cyathistipula 'Ficus cyathistipula', the African fig tree, is a species of fig that is native to the tropical forest regions of Africa. They may be small trees, shrubs or hemi-epiphytic lianas, and are widespread in the moist tropics, where they may be found ...
'' Warb. *'' Ficus cyclophylla'' (Miq.) Miq. *'' Ficus dendrocida'' Kunth *'' Ficus depressa'' Bl. *''
Ficus destruens ''Ficus destruens'' is a hemiepiphyte, hemiepiphytic ficus, fig that is endemic to the wet tropical rainforests of northeastern Queensland, Australia. Description ''Ficus destruens'' is a monoecious tree which grows up to tall. Its leaves are ...
'' F.White *'' Ficus drupacea'' Thunb. *'' Ficus elastica'' Hornem. – rubber plant *''
Ficus exasperata ''Ficus exasperata'', also called the sandpaper tree, forest sandpaper fig, white fig, or sandpaper leaf tree, is a deciduous, and dioecious species of plant in the mulberry family Moraceae, native to tropical Africa (an area from Senegal east t ...
'' Vahl. *''
Ficus faulkneriana ''Ficus faulkneriana'' is a species of strangler fig in the family Moraceae native to Africa. Distribution The tree is endemic to coastal Kenya and Tanzania, and in the Usambara Mountains of Tanzania, in tropical East Africa. It is found in c ...
'' Berg *''
Ficus fergusonii ''Ficus fergusonii'', is a species of plant in the fig genus, endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that ar ...
'' (King) T.B.Worth. ex Corner *'' Ficus glaberrima'' Blume *''
Ficus glumosa ''Ficus glumosa'' also known as the Mountain or Hairy Rock Fig is an Afrotropical fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asi ...
'' Delile *''
Ficus greiffiana ''Ficus greiffiana'' is a species of plant in the family Moraceae. It is found in Argentina, Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Lat ...
'' Dugand *'' Ficus hirsuta'' Schott *'' Ficus ilicina'' Miq. *''
Ficus kerkhovenii ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few species extending int ...
'' Valeton – Johore fig *''
Ficus kurzii ''Ficus kurzii'' may be called the Burmese banyan: it is an Asian species of fig tree in the family Moraceae. No subspecies are listed in the Catalogue of Life; its native range is China (Yunnan) Indo-China and Malesia. The species can be foun ...
'' King *''
Ficus luschnathiana ''Ficus luschnathiana'' is a species of plant in the family Moraceae. It is found in Argentina, Brazil, Bolivia, Paraguay, and Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a co ...
'' Miq. *''
Ficus ingens ''Ficus ingens'', the red-leaved fig, is a fig species with an extensive range in the subtropical to dry tropical regions of Africa and southern Arabia. Despite its specific name, which means "huge", or "vast", it is usually a shrub or tree of mo ...
'' Miq. *''
Ficus krukovii ''Ficus krukovii'' is a species of plant in the family Moraceae. It is found in Bolivia, Brazil, and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a coun ...
'' Standl. *'' Ficus lacor'' Buch.-Ham. *''
Ficus lapathifolia ''Ficus lapathifolia'' is a species of plant in the family Moraceae. It is endemic to Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the nor ...
'' Miq. *'' Ficus lauretana'' Vázquez Avila *'' Ficus lutea'' Vahl *''
Ficus lyrata ''Ficus lyrata'', commonly known as the fiddle-leaf fig, is a species of flowering plant in the mulberry and fig family Moraceae. It is native to western Africa, from Cameroon west to Sierra Leone, where it grows in lowland tropical rainforest ...
'' Warb. – fiddle-leaved fig *'' Ficus maclellandii'' King – Alii fig *'' Ficus macrophylla'' Desf. ex Pers. – Moreton Bay fig *''
Ficus malacocarpa ''Ficus popenoei'' is a species of fig found in Latin America, from Brazil and Peru up to Guatemala and Belize. Subspecies There are two subspecies: * ''Ficus popenoei'' subsp. ''malacocarpa'' Standl. – sometimes considered a separate specie ...
'' Standl. *''
Ficus mariae ''Ficus mariae'' is a species of tree in the family Moraceae. It is native to South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the ...
'' Berg, Emygdio & Carauta *''
Ficus mathewsii ''Ficus mathewsii'' is a species of plant in the family Moraceae. It is found in Brazil, Colombia, Guyana, Peru, and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Ven ...
'' Miq. *''
Ficus matiziana ''Ficus matiziana'' is a species of plant in the family Moraceae. It is found in Brazil, Colombia, Guyana, and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela ...
'' Dugand *''
Ficus mexiae ''Ficus mexiae'' is a species of fig in the family Moraceae, native to Brazil. The tree is endemic to the Atlantic Forest ecoregion of Southeast Brazil, within the states of Bahia and Minas Gerais. It is an IUCN Red List Vulnerable species ...
'' Standl. *'' Ficus microcarpa'' L. – Chinese banyan *'' Ficus muelleriana'' Berg *'' Ficus natalensis'' Hochst. – Natal fig *'' Ficus obliqua'' G.Forst. – small-leaved fig *'' Ficus obtusifolia'' Kunth *''
Ficus pakkensis ''Ficus pakkensis'' is a species of plant in the family Moraceae, native to tropical northern South America. It is found in Guyana, and in the states of Pará and Maranhão in northern Brazil. It is an IUCN Red List Vulnerable species A vul ...
'' Standl. *'' Ficus pallida'' Vahl *''
Ficus panurensis ''Ficus panurensis'' is a species of plant in the family Moraceae. It is found in Brazil, Guyana, Suriname, and Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), ...
'' Standl. *'' Ficus pertusa'' L.f. *''
Ficus petiolaris ''Ficus petiolaris'', commonly known as the petiolate fig and rock fig, is a fig that is endemic to Mexico from Baja California and Sonora south to Oaxaca. It grows from 10–20 feet high. It grows best with moderate water and partial shade. A uni ...
'' Kunth *''
Ficus pisocarpa ''Ficus pisocarpa''von Blume CL (1825) In: ''Bijdragen tot de Flora van Nederlandsch Indie'' 454. is a banyan fig species in the family Moraceae. No subspecies are listed in the Catalogue of Life. The species can be found in southern China, Ind ...
'' Bl. *'' Ficus platypoda'' Cunn. – desert fig *''
Ficus pleurocarpa ''Ficus pleurocarpa'', commonly known as the banana fig, karpe fig or gabi fig, is a fig that is endemic to the wet tropical rainforests of northeastern Queensland, Australia. It has characteristic ribbed orange and red cylindrical syconia. It ...
'' DC. – banana fig *'' Ficus polita'' Vahl *''
Ficus religiosa ''Ficus religiosa'' or sacred fig is a species of Ficus, fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipa ...
'' L. – sacred fig *''
Ficus roraimensis ''Ficus roraimensis'' is a species of fig tree in the family Moraceae. The tree is endemic to Roraima state in western Brazil. It is an IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species ...
'' Berg *'' Ficus rubiginosa'' Desf. – Port Jackson fig *''
Ficus rumphii ''Ficus rumphii''von Blume CL (1825) In: ''Bijdragen tot de Flora van Nederlandsch Indie'' 437. is a banyan fig species in the family Moraceae. No subspecies are listed in the Catalogue of Life. The species can be found in: India, southern Chin ...
'' Blume *'' Ficus salicifolia'' Vahl – willow-leaved fig *'' Ficus sansibarica'' Warb. *'' Ficus schippii'' Standl. *''
Ficus schultesii ''Ficus gomelleira'' is a species of flowering plant, a tree in the family Moraceae. This species is monoecious. Through the 1950s to the 1990s this species was widely viewed as the iroko tree. The tree can have a size up to 40 m, with a dia ...
'' Dugand *'' Ficus schumacheri'' Griseb. *''
Ficus sphenophylla ''Ficus sphenophylla'' is a species of fig tree in the family Moraceae. It is found in Bolivia, Brazil, Colombia, and Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_sym ...
'' Standl. *''
Ficus stuhlmannii ''Ficus stuhlmannii'' is a tree in the family Moraceae. It is commonly known as the lowveld fig. These trees are distributed from KwaZulu-Natal in South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern ...
'' Warb. *'' Ficus subcordata'' Bl. *'' Ficus subpisocarpa'' Gagnep. *''
Ficus subpuberula ''Ficus subpuberula'' is a lithophytic fig that is endemic to Australia. It ranges from extreme western Queensland, through the Northern Territory, into Western Australia Western Australia (commonly abbreviated as WA) is a state of Australi ...
'' Corner *''
Ficus sumatrana ''Ficus sumatrana'' is an Asian species of fig tree in the family Moraceae. No subspecies are listed in the Catalogue of Life; its native range is Indo-China to Malesia. Description The species can be found in Vietnam: where it may be called ' ...
'' Miq. *'' Ficus superba'' Miq. ** ''Ficus superba'' var. ''henneana'' (Miq.) Corner *'' Ficus thonningii'' Blume *'' Ficus trichopoda'' Baker *''
Ficus trigona Ficus trigona is a species of tree in the family Moraceae. It is native to South America. Characteristics ''Ficus trigona'' typically starts life as an epiphyte on another tree. Eventually the plant will send roots to the ground in order to see ...
'' L.f. *''
Ficus trigonata ''Ficus trigonata'' is a species of tree in the family Moraceae The Moraceae — often called the mulberry family or fig family — are a family of flowering plants comprising about 38 genera and over 1100 species. Most are widespread in tropi ...
'' L. *'' Ficus triradiata'' Corner – red-stipule fig *'' Ficus ursina'' Standl. *''
Ficus velutina ''Ficus velutina'' is a species of plant in the family Moraceae. It is found in Brazil and Costa Rica Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the ...
'' Willd. *'' Ficus verruculosa'' Warb. *'' Ficus virens'' Aiton – white fig ** ''Ficus virens'' var. ''sublanceolata'' (Miq.) Corner – sour fig *'' Ficus watkinsiana'' F.M.Bailey – Watkins's fig


Unknown subgenus

*''
Ficus bibracteata ''Ficus'' ( or ) is a genus of about 850 species of woody trees, shrubs, vines, epiphytes and hemiepiphytes in the family (biology), family Moraceae. Collectively known as fig trees or figs, they are native throughout the tropics with a few spe ...
'' *'' Ficus callosa'' Willd. *'' Ficus cristobalensis'' *'' Ficus hebetifolia'' *''
Ficus tsjahela ''Ficus tsjahela'' is a fig tree from the family Moraceae which is found in peninsular India and Sri Lanka. It is commonly known as the ''kaaral'' in Malayalam, ''kal-aal'' in Tamil and ''boviyamara'' in Kannada. State wise distribution ' ...
'' Burm.f. * ''
Ficus nymphaeifolia ''Ficus nymphaeifolia'' is a strangler fig found in central and South America. Growing up to 35 metres tall, the habitat is coastal plains, savannah and rainforest. This plant first appeared in scientific literature in 1768. Published in The ...
'' Mill.


Uses

The wood of fig trees is often soft and the latex precludes its use for many purposes. It was used to make mummy caskets in Ancient Egypt. Certain fig species (mainly ''F. cotinifolia'', '' F. insipida'' and '' F. padifolia'') are traditionally used in Mesoamerica to produce ''
papel amate Amate ( es, amate from nah, āmatl ) is a type of bark paper that has been manufactured in Mexico since the precontact times. It was used primarily to create codices. Amate paper was extensively produced and used for both communication, record ...
'' (
Nahuatl Nahuatl (; ), Aztec, or Mexicano is a language or, by some definitions, a group of languages of the Uto-Aztecan language family. Varieties of Nahuatl are spoken by about Nahua peoples, most of whom live mainly in Central Mexico and have smaller ...
: ''āmatl''). ''Mutuba'' ('' F. natalensis'') is used to produce barkcloth in Uganda. ''Pou'' ('' F. religiosa'') leaves' shape inspired one of the standard '' kbach rachana'', decorative elements in Cambodian architecture. Indian banyan (''F. benghalensis'') and the Indian rubber plant, as well as other species, have use in herbalism. The inner bark of an unknown type of wild fig, locally known as ''urú'', was once used by the of Bolivia to produce a fibrous cloth used for clothing. Figs have figured prominently in some human cultures. There is evidence that figs, specifically the
common fig The fig is the edible fruit of ''Ficus carica'', a species of small tree in the flowering plant family Moraceae. Native to the Mediterranean and western Asia, it has been cultivated since ancient times and is now widely grown throughout the world ...
(''F. carica'') and sycamore fig ('' Ficus sycomorus''), were among the first – if not the very first – plant species that were deliberately bred for agriculture in the Middle East, starting more than 11,000 years ago. Nine
subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in ...
''F. carica'' figs dated to about 9400–9200 BCE were found in the early Neolithic village Gilgal I (in the Jordan Valley, 13 km, or 8.1 mi, north of
Jericho Jericho ( ; ar, أريحا ; he, יְרִיחוֹ ) is a Palestinian city in the West Bank. It is located in the Jordan Valley, with the Jordan River to the east and Jerusalem to the west. It is the administrative seat of the Jericho Gove ...
). These were a parthenogenetic type and thus apparently an early cultivar. This find predates the first known cultivation of
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legum ...
in the Middle East by many hundreds of years. Fig is a popular species in the practice of bonsai. It is particularly popular with beginners, as it is widely available and survives well as an indoor plant.


Cultivation

Numerous species of fig are found in cultivation in domestic and office environments, including: *'' F. carica'', common fig – hardy to . Shrub or small tree which can be grown outdoors in mild temperate regions, producing substantial harvests of fruit. Many cultivars are available. *'' F. benjamina'', weeping fig, ficus – hardy to . Widely used as an indoor plant for the home or the office. It benefits from the dry, warm atmosphere of centrally-heated interiors, and can grow to substantial heights in a favoured position. Several variegated cultivars are available. *'' F. elastica'', rubber plant – hardy to : widely cultivated as a houseplant; several cultivars with variegated leaves *'' F. lyrata'', fiddle-leaf fig – hardy to *'' F. maclellandii'' – hardy to *'' F. microcarpa'', Indian laurel – hardy to *'' F. pumila'', creeping fig – hardy to *'' F. rubiginosa'', Port Jackson fig – hardy to


Cultural and spiritual significance

Fig trees have profoundly influenced culture through several religious traditions. Among the more famous species are the sacred fig tree (Pipal, bodhi, bo, or po, ''Ficus religiosa'') and other banyan figs such as ''Ficus benghalensis''. The oldest living plant of known planting date is a ''Ficus religiosa'' tree known as the Sri Maha Bodhi planted in the temple at Anuradhapura, Sri Lanka by King Tissa in 288 BCE. The common fig is one of two significant trees in
Islam Islam (; ar, ۘالِإسلَام, , ) is an Abrahamic religions, Abrahamic Monotheism#Islam, monotheistic religion centred primarily around the Quran, a religious text considered by Muslims to be the direct word of God in Islam, God (or ...
, and there is a
sura A ''surah'' (; ar, سورة, sūrah, , ), is the equivalent of "chapter" in the Qur'an. There are 114 ''surahs'' in the Quran, each divided into '' ayats'' (verses). The chapters or ''surahs'' are of unequal length; the shortest surah ('' Al-K ...
in Quran named "The Fig" or At-Tin (سوره تین). In Asia, figs are important in Buddhism and Hinduism. In Jainism, the consumption of any fruit belonging to this genus is prohibited. The Buddha is traditionally held to have found ''
bodhi The English term enlightenment is the Western translation of various Buddhist terms, most notably bodhi and vimutti. The abstract noun ''bodhi'' (; Sanskrit: बोधि; Pali: ''bodhi''), means the knowledge or wisdom, or awakened intellect ...
'' (enlightenment) while meditating for 49 days under a sacred fig. The same species was '' Ashvattha'', the "
world tree The world tree is a motif present in several religions and mythologies, particularly Indo-European religions, Siberian religions, and Native American religions. The world tree is represented as a colossal tree which supports the heavens, thereb ...
" of Hinduism. The '' Plaksa Pra-sravana'' was said to be a fig tree between the roots of which the Sarasvati River sprang forth; it is usually held to be a sacred fig but more probably is '' Ficus virens''. According to the
Kikuyu people The Kikuyu (also ''Agĩkũyũ/Gĩkũyũ'') are a Bantu ethnic group native to Central Kenya. At a population of 8,148,668 as of 2019, they account for 17.13% of the total population of Kenya, making them Kenya's largest ethnic group. The te ...
, sacrifices to Ngai were performed under a sycomore tree (Mũkũyũ) and if one was not available, a fig tree (Mũgumo) would be used. The common fig tree is cited in the Bible, where in Genesis 3:7, Adam and Eve cover their nakedness with fig leaves. The fig fruit is also one of the traditional crops of Israel, and is included in the list of food found in the Promised Land, according to the Torah ( Deut. 8). Jesus cursed a fig tree for bearing no fruit (). The fig tree was sacred in ancient Greece and Cyprus, where it was a symbol of fertility. File:Ficus religiosa Bo.jpg, Leaves of the sacred fig (''
Ficus religiosa ''Ficus religiosa'' or sacred fig is a species of Ficus, fig native to the Indian subcontinent and Indochina that belongs to Moraceae, the fig or mulberry family. It is also known as the bodhi tree, pippala tree, peepul tree, peepal tree, pipa ...
'') File:เศียรพระพุทธรูปในรากโพธิ์.jpg, Fig tree roots overgrowing a sandstone Buddha statue, near
Wat Maha That A wat ( km, វត្ត, ; lo, ວັດ, ; th, วัด, ; khb, 「ᩅᨯ᩠ᨰ」(waD+Dha); nod, 「ᩅ᩠ᨯ᩶」 (w+Da2)) is a type of Buddhist temple and Hindu temple in Cambodia, Laos, East Shan State, Yunnan, the Southern Provi ...
in Ayutthaya province, Thailand Image:Sarkaradevi Temple Ficus Tree.jpg, Ficus tree in front of Sarkaradevi Temple, Kerala, India


List of famous fig trees

* '' Ashvattha'' – the
world tree The world tree is a motif present in several religions and mythologies, particularly Indo-European religions, Siberian religions, and Native American religions. The world tree is represented as a colossal tree which supports the heavens, thereb ...
of Hinduism, held to be a supernatural ''F. religiosa'' * Bodhi tree – a ''F. religiosa'' *
Charybdis Fig Tree Charybdis (; grc, Χάρυβδις, Khárybdis, ; la, Charybdis, ) is a sea monster in Greek mythology. She, with the sea monster Scylla, appears as a challenge to epic characters such as Odysseus, Jason, and Aeneas. Scholarship locates her in t ...
of Homer's '' Odyssey'', presumably a ''F. carica'' *
Curtain Fig Tree Curtain Fig Tree is a heritage-listed tree at Curtain Fig Tree Road, Yungaburra, Tablelands Region, Queensland, Australia. It is one of the largest trees in Tropical North Queensland, Australia, and one of the best known attractions on t ...
– a ''F. virens'' * Ficus Ruminalis – a ''F. carica'' * '' Plaksa'' – another supernatural fig in Hinduism; usually identified as ''F. religiosa'' but is probably ''F. virens'' * Santa Barbara's Moreton Bay Fig Tree – a ''F. macrophylla'' * Sri Maha Bodhi – another ''F. religiosa'', planted in 288 BCE, the oldest human-planted tree on record * '' The Barren Fig Tree'' – Matthew 21:19 of The Holy Bible, Jesus put a curse on the tree and used this as an example for believers of the promise of the power faith in the only true God. '' * The Great Banyan – a ''F. benghalensis'', a
clonal colony A clonal colony or genet is a group of genetically identical individuals, such as plants, fungi, or bacteria, that have grown in a given location, all originating vegetatively, not sexually, from a single ancestor. In plants, an individual in s ...
and once the largest organism known * Vidurashwatha – " Vidura's Sacred Fig Tree", a village in India named after a famous ''F. religiosa'' that until recently stood there * Wonderboom – the largest fig tree in Pretoria, South Africa


Citations


General references

* * * * * * * *
Supporting Online Material
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*


External links



Major reference site for the genus ''Ficus''
World checklist of Ficus species from the Catalogue of Life
845 species supplied by M. Hassler's World Plants.

Multi-award-winning documentary


BBC: Fig fossil clue to early farming
Video
How the fig tree strangles other plants for survival in the rainforest
{{Authority control Moraceae genera Taxa named by Carl Linnaeus