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''Ferocactus viridescens'' is a species of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the cactus family Cactaceae. This rare
barrel cactus Barrel cacti are various members of the two genera ''Echinocactus'' and ''Ferocactus'', endemic to the deserts of Southwestern North America southward to north central Mexico. Some of the largest specimens are found in the Sonoran Desert in So ...
is known by several common names, including coast barrel cactus, keg cactus and San Diego barrel cactus. Most of its native range in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
is in San Diego County, California, where it is threatened by development, agriculture, and other alterations in its habitat. It is also found in northern
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
, Mexico.


Description

''Ferocactus viridescens'' is a solitary barrel cactus, typically spherical, oblate, or nearly cylindrical, reaching heights of 10-30(-45)cm and diameters of 10-20(-35)cm. It has bright green flesh arranged into several 13 to 34 blunt ribs covered in arrays of long spines. The
areole In botany, areoles are small light- to dark-colored bumps on cacti out of which grow clusters of spines. Areoles are important diagnostic features of cacti, and identify them as a family distinct from other succulent plants. Gordon Rowley - W ...
s are narrow elliptic to oval, 10-20 mm long, with brownish tomentum, bearing 10-19 spines per areole, including central and radial spines. The central spines are initially pink or yellowish, becoming duller with age, with the principal central spine measuring 30-50 cm long and 2-3(-5) mm broad. The 10-20 radial spines are similar to the centrals but smaller, 1-2 cm long. Blooming from spring to early summer, the species produces infundibuliform flowers that are 2.5-5 cm long and 3-6 cm in diameter, with greenish-yellow or greenish perianth segments sometimes tinged with pink. The flower tube bears stamens almost to the top of the ovary, with yellow stigma lobes. The fruits are barrel-like, bright yellow (rarely reddish), with a pleasant acid taste, measuring 20-35 mm long and 15-25 mm across. They have a leathery skin and about 30 fimbriate-denticulate scales, dehiscent through a basal pore. The seeds are elongate-obovoid, minutely reticulate, and measure 1.6-1.8 mm long..


Taxonomy

John Torrey and Asa Gray first described ''Ferocactus viridescens'' as ''Echinocactus viridescens'' in 1840. The Latin specific epithet ''viridescens'' means "turning green".
Nathaniel Lord Britton Nathaniel Lord Britton (January 15, 1859 – June 25, 1934) was an American botanist and taxonomist who co-founded the New York Botanical Garden in the Bronx, New York. Early life Britton was born in New Dorp in Staten Island, New York to Jasp ...
and
Joseph Nelson Rose Joseph Nelson Rose (January 11, 1862 – May 4, 1928) was an American botanist. He was born in Union County, Indiana. His father died serving during the Civil War when Joseph Rose was a young boy. He later graduated from high school in Libert ...
later reclassified the species under the genus ''Ferocactus'' in 1922. ''Ferocactus viridescens'' ssp. ''littoralis'' G.E.Linds., also known as the Santo Tomas coast barrel cactus, is found in a small area between Ensenada and
Vicente Guerrero Vicente Ramón Guerrero (; baptized August 10, 1782 – February 14, 1831) was one of the leading revolutionary generals of the Mexican War of Independence. He fought against Spain for independence in the early 19th century, and later served as ...
,
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
. It has also been sighted slightly further south, along the coastline, near El Socorrito and Rancho Los Pinos.


Distribution

San Diego barrel cacti, as their name suggests, are found growing in the extreme south of
Southern California Southern California (commonly shortened to SoCal) is a geographic and cultural region that generally comprises the southern portion of the U.S. state of California. It includes the Los Angeles metropolitan area, the second most populous urban a ...
—primarily in
San Diego County San Diego County (), officially the County of San Diego, is a county in the southwestern corner of the U.S. state of California. As of the 2020 census, the population was 3,298,634, making it California's second-most populous county and the f ...
—as well as across the U.S.-Mexico border into
Baja California state Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
, where it is found mostly along the coast, ranging south towards El Rosario and San Quintín. Beyond
San Diego San Diego ( , ; ) is a city on the Pacific Ocean coast of Southern California located immediately adjacent to the Mexico–United States border. With a 2020 population of 1,386,932, it is the eighth most populous city in the United State ...
,
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
, the species has been observed as far north as
San Clemente San Clemente (; Spanish for " St. Clement") is a city in Orange County, California. Located in the Orange Coast region of the South Coast of California, San Clemente's population was 64,293 in at the 2020 census. Situated roughly midway between ...
(
Orange County Orange County most commonly refers to: *Orange County, California, part of the Los Angeles metropolitan area Orange County may also refer to: U.S. counties *Orange County, Florida, containing Orlando *Orange County, Indiana *Orange County, New ...
) and Perris (
Riverside County Riverside County is a county located in the southern portion of the U.S. state of California. As of the 2020 census, the population was 2,418,185, making it the fourth-most populous county in California and the 10th-most populous in the Uni ...
). Its preferred habitat varies, as it inhabits elevations ranging from sea level to 200 meters (656 feet), in coastal, mild-winter
microclimates A microclimate (or micro-climate) is a local set of atmospheric conditions that differ from those in the surrounding areas, often with a slight difference but sometimes with a substantial one. The term may refer to areas as small as a few squa ...
as well as warmer, inland areas that may experience autumn or winter frosts. It grows in sandy and gravelly soils with a generally high mineral content (but comparatively low amounts of organic matter). Habitats where ''Ferocactus viridescens'' may be found include chapparal, arid hills, dry and grassy
dunes A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, fl ...
, or rocky coastal bluffs and sun-baked cliffs; cacti especially thrive along the Californian and Mexican west coasts, as they experience significant fog cover (
marine layer A marine layer is an air mass that develops over the surface of a large body of water, such as an ocean or large lake, in the presence of a temperature inversion. The inversion itself is usually initiated by the cooling effect of the water on th ...
) throughout the year, notably in the mornings and evenings, allowing the plants to absorb sufficient moisture before the heat of the day. This habitat also gives ''Ferocactus viridescens'' an innate tolerance for salts and minerals that many botanical species simply cannot handle. The San Diego barrel cactus can also be found on shrub-covered hillsides, often at the crests of slopes, and in cobbles. Occasionally, it is found on the edges of
vernal pools Vernal pools, also called vernal ponds or ephemeral pools, are seasonal pools of water that provide habitat for distinctive plants and animals. They are considered to be a distinctive type of wetland usually devoid of fish, and thus allow the safe ...
, sometimes in large numbers. This habitat, which is more moist compared to its usual
xeric Deserts and xeric shrublands are a biome defined by the World Wide Fund for Nature. Deserts and xeric (ancient Greek xērós, “dry") shrublands form the largest terrestrial biome, covering 19% of Earth's land surface area. Ecoregions in this h ...
environment, is unusual for the plant, and may occur sporadically during years of excessive precipitation in the area.


Cultivation

This plant, which is hardy down to , must be grown under glass in temperate regions, though it may be placed outside in a sheltered spot in the warm summer months. It has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's Award of Garden Merit.


References


External links

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Jepson Manual TreatmentUSDA Plants ProfilePhoto gallery
{{Taxonbar, from=Q147469, from2=Q24691865, from3=Q96108196 viridescens Cacti of the United States var. viridescens Flora of California Natural history of the California chaparral and woodlands Natural history of San Diego County, California