Feitianius
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''Feitianius'' is a
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
, belonging to the
Enantiornithes The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and cla ...
, that during the
Early Cretaceous The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Pro ...
lived in the area of modern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
. A single species has been named in the genus, ''Feitianius paradisi''. In 2005, fragmentary enantiornithean fossils were excavated near Changma, in
Gansu Gansu (, ; alternately romanized as Kansu) is a province in Northwest China. Its capital and largest city is Lanzhou, in the southeast part of the province. The seventh-largest administrative district by area at , Gansu lies between the Tibet ...
. Two of the best preserved were reported in the scientific literature in 2009.Lamanna, M., D.-Q. Li, J.D. Harris, J.A. Atterholt, and H.-L. You, 2009, "Exceptionally preserved Enantiornithes (Aves: Ornithothoraces) from the Early Cretaceous of northwestern China", ''Journal of Vertebrate Paleontology'' 29(3, Supplement): 131A In 2015, the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
''Feitianius paradisi'' was named and described by Jingmai Kathleen O’Connor, Li Daqing, Matthew C. Lamanna, Wang Min, Jerald D. Harris, Jessie Atterholt and You Hailou. The generic name has been derived from the feitia, a flying
apsara An apsaras or apsara ( sa, अप्सरा ' lso ' pi, अक्चरा, translit=accharā) is a type of female spirit of the clouds and waters in Hinduism and Buddhist culture. They figure prominently in the sculpture, dance, literat ...
or nymph from
Buddhist mythology The Buddhist traditions have created and maintained a vast body of mythological literature. The central myth of Buddhism is the life of the Buddha. This is told in relatively realistic terms in the earliest texts, and was soon elaborated into ...
. Historical murals featuring such nymphs are common in the caves around Changma. The name was "for fun" made masculine because it was concluded that the type specimen was a male individual. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
means "from paradise" in
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
and refers to the similar
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
in the tail feathers of birds-of-paradise. The species name as a whole thus means "feitia of paradise", the translation of the Chinese 飛天/飞天.O’Connor, J. K., D.-Q. Li, M. C. Lamanna, M. Wang, J. D. Harris, J. Atterholt, and H.-L. You. 2015. "A new Early Cretaceous enantiornithine (Aves, Ornithothoraces) from northwestern China with elaborate tail ornamentation". ''Journal of Vertebrate Paleontology''. DOI: 10.1080/02724634.2015.1054035 https://www.tandfonline.com/doi/abs/10.1080/02724634.2015.1054035?journalCode=ujvp20 The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, GSGM-05-CM-004, was found in a layer of the
Xiagou Formation The Xiagou Formation is the middle strata of the Xinminbao Group. It is named for its type site in Xiagou, in the Changma Basin of Gansu Province, northwestern China and is considered Early Cretaceous in age. It is known outside the specialized ...
dating from the early
Aptian The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), a ...
. In consists of the hind part of a skeleton on a stone plate. The skeleton is visible from the left side. It is articulated but the part in front of the middle back is lacking. It preserves extensive remains of the soft tissues, including the plumage. An indication of the size of ''Feitianius'' is given by the length of the thighbone, twenty-seven millimetres. A single unique feature was established for ''Feitianius'': the tail plumage contains three distinct feather types whereas all known relatives have tails with a single type of feather. Soft tissue remains are preserved around the tail base. These seem to represent a fleshy structure similar to the "parson's nose" of modern birds, in which the tail feathers are embedded.
Collagen Collagen () is the main structural protein in the extracellular matrix found in the body's various connective tissues. As the main component of connective tissue, it is the most abundant protein in mammals, making up from 25% to 35% of the whole ...
bands are visible running perpendicular to the body outline. These bands have a length of seven to eight millimetres and a width of about half a millimetre. On the belly and the front tail base short downy covert feathers are visible. More to behind the feathers of the tail base are gradually changing into longer structures, a dozen of which are clearly visible. Uniquely for the Enantiornithes, among these three
morphotype In biology, polymorphism is the occurrence of two or more clearly different morphs or forms, also referred to as alternative ''phenotypes'', in the population of a species. To be classified as such, morphs must occupy the same habitat at the s ...
s can be distinguished. Morphotype A consists of feathers that are forty to forty-five millimetres long. They have vanes that cover the distal 55% of the rachis. Morphotype B is eighteen millimetres long and half as wide as morphotype A. Here, the vanes cover the full length of the rachis. Both types are pennaceous and somewhat curved downwards. Of morphotype C only a single rachis has been preserved. It is very long and its distal end has been lost on the fossil. The describing authors concluded that it was one of a pair of very long central "streamers" having a display function, probably for a male individual. It was considered unlikely that the tail feathers had an important aerodynamic effect. It is possible that the shorter morphotype A and B feathers formed a differentially coloured tuft on the tail base. Because of a varnish applied to the fossil, it can no longer be checked for colouring melanosomes though. The ornamental structure as a whole might have resembled that of the extant
greater bird-of-paradise The greater bird-of-paradise (''Paradisaea apoda'') is a bird-of-paradise in the genus '' Paradisaea''. Carl Linnaeus named the species ''Paradisaea apoda'', or "legless bird-of-paradise", because early trade-skins to reach Europe were prepared ...
. ''Feitianius'' was in 2016 placed in the Enantiornithes. A cladistic analysis showed it to be part of a polytomy with '' Qiliania'' and other enantiornithean specimens of the Changma Basin.


References

{{Taxonbar, from=Q54500314 Enantiornitheans Fossil taxa described in 2015