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''Thymallus thymallus'', the grayling or European grayling, is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...
in the
salmon Salmon () is the common name for several list of commercially important fish species, commercially important species of euryhaline ray-finned fish from the family (biology), family Salmonidae, which are native to tributary, tributaries of the ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Salmonidae Salmonidae is a family of ray-finned fish that constitutes the only currently extant family in the order Salmoniformes . It includes salmon (both Atlantic and Pacific species), trout (both ocean-going and landlocked), chars, freshwater whitefis ...
. It is the only species of the genus ''
Thymallus ''Thymallus'' is a genus of freshwater fish in the salmon family Salmonidae; it is the only genus of subfamily Thymallinae. The type species is ''Thymallus thymallus,'' the grayling. The species in the genus are generically called graylings, but ...
'' (the graylings) native to
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, where it is widespread from the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...
and
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
to the
Ural Mountains The Ural Mountains ( ; rus, Ура́льские го́ры, r=Uralskiye gory, p=ʊˈralʲskʲɪjə ˈɡorɨ; ba, Урал тауҙары) or simply the Urals, are a mountain range that runs approximately from north to south through western ...
in
Russia Russia (, , ), or the Russian Federation, is a List of transcontinental countries, transcontinental country spanning Eastern Europe and North Asia, Northern Asia. It is the List of countries and dependencies by area, largest country in the ...
, and
Balkans The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
on the south-east, but does not occur in the southern parts of the continent. It was introduced to
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
in 1948, but it does not appear to have become established there.


Description

The grayling grows to a maximum recorded length of and a maximum recorded weight of . Of typical ''
Thymallus ''Thymallus'' is a genus of freshwater fish in the salmon family Salmonidae; it is the only genus of subfamily Thymallinae. The type species is ''Thymallus thymallus,'' the grayling. The species in the genus are generically called graylings, but ...
'' appearance, the grayling proper is distinguished from the similar
Arctic grayling The Arctic grayling (''Thymallus arcticus'') is a species of freshwater fish in the salmon family Salmonidae. ''T. arcticus'' is widespread throughout the Arctic and Pacific drainages in Canada, Alaska, and Siberia, as well as the upper Misso ...
(''T. arcticus arcticus'') by the presence of 5–8
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage * Dorsal co ...
and 3–4
anal Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involvin ...
spines, which are absent in the other species; ''T. thymallus'' also has a smaller number of soft rays in these fins. Individuals of the species have been recorded as reaching an age of 14 years. The grayling prefers cold, clean, running riverine waters, but also occurs in lakes and, exceptionally, in brackish waters around the
Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and from ...
.
Omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutri ...
, the fish feeds on vegetable matter, as well as
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, amphipods and mantis shrimp. The crustacean group ...
s,
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s and
spider Spiders ( order Araneae) are air-breathing arthropods that have eight legs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species ...
s,
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is e ...
s,
zooplankton Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
, and smaller fishes, such as
Eurasian minnow ''Phoxinus'' is a genus of freshwater fish in the family Leuciscidae of order Cypriniformes, and the only members of the subfamily Phoxininae, or Eurasian minnows. The type species is ''Phoxinus phoxinus''. The other species in this genus are ...
s. Grayling are also prey for larger fish, including the
huchen The huchen (''Hucho hucho'') (, from German), also known as Danube salmon or redfish (german: Rotfisch), is a large species of freshwater fish in the salmon family of order Salmoniformes. It is the type species of its genus ''Hucho''. Distribut ...
(''Hucho hucho''). With the Arctic grayling, ''T. thymallus'' is one of the economically important ''Thymallus'' species, being raised commercially and fished for sport. The grayling is a protected species listed in Appendix III of the Bern Convention. It has become critically endangered in the Baltic Sea. The term "grayling" is often used to refer generically to the ''Thymallus'' species, and ''T. thymallus'' is sometimes called the European grayling for clarity. There are many obsolete synonyms for the species. The generic name ''Thymallus'' derives from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
θύμαλλος, "
thyme Thyme () is the herb (dried aerial parts) of some members of the genus ''Thymus'' of aromatic perennial evergreen herbs in the mint family Lamiaceae. Thymes are relatives of the oregano genus ''Origanum'', with both plants being mostly indigenou ...
smell", a name derived from the fragrance of wild thyme that freshly caught graylings are believed to smell similar to. ''Thymallus thymallus'' is the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
of its genus.


Angling

The grayling is known as the 'lady of the stream'. In Europe, they used to be persecuted by anglers for the false perception that they stopped
trout Trout are species of freshwater fish belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of the subfamily Salmoninae of the family Salmonidae. The word ''trout'' is also used as part of the name of some non-salmoni ...
colonizing stretches of
river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of wate ...
s and
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a stream ...
s. However, research has shown that grayling and
trout Trout are species of freshwater fish belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of the subfamily Salmoninae of the family Salmonidae. The word ''trout'' is also used as part of the name of some non-salmoni ...
feed on different prey items and generally prefer different microhabitats within
river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of wate ...
s and
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a stream ...
s but do occupy similar niches to smaller, less-predatory trout. In England and Wales, they can be fished for throughout the
coarse fishing In Britain and Ireland, coarse fishing (, ) refers to angling for rough fish, which are fish species traditionally considered undesirable as a food or game fish. Freshwater game fish are all salmonids — most particularly salmon, trout and cha ...
season (16 June to 14 March), providing thrilling sport on the
fly Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwing ...
when the trout season is closed. There is no closed season for grayling in Scotland; where they have been introduced. There are no grayling in Ireland. Well-known grayling flies include the grayling witch, klinkhamers, various nymphs and 'red tags', along with other trout patterns. Flies tied to resemble small pink shrimps have also been found to be useful. A method known as 'Czech-nymphing' has been known to be helpful to anglers where grayling shoal up in colder periods. The method involves moving a series of Czech nymphs under the tip of the fly rod with the flow of the river and the nymphs should entice the grayling to take one. Fly-anglers may wade in the river to perform this method where they can access deeper water. Wading does not necessarily spook the grayling as they are generally less cautious than trout and are not as easily put off by human presence. In
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, too, the season is limited depending upon several factors. The
Allier Allier ( , , ; oc, Alèir) is a department in the Auvergne-Rhône-Alpes region that borders Cher to the west, Nièvre to the north, Saône-et-Loire and Loire to the east, Puy-de-Dôme to the south, and Creuse to the south-west. Named afte ...
is one of the rare places in Southern Europe where the common grayling occurs in a natural habitat.


References


Further reading

* * * * * {{Authority control
thymallus ''Thymallus'' is a genus of freshwater fish in the salmon family Salmonidae; it is the only genus of subfamily Thymallinae. The type species is ''Thymallus thymallus,'' the grayling. The species in the genus are generically called graylings, but ...
Freshwater fish of Europe Taxa named by Carl Linnaeus