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''Euptoieta claudia'', the variegated fritillary, is a
North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography. Etymology T ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
n
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
in the family
Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species have a red ...
. Even though the variegated fritillary has some very different characteristics from the ''
Speyeria ''Speyeria'', commonly known as greater fritillaries, is a genus of butterflies in the family Nymphalidae commonly found in North America, Europe, and Asia. Some authors used to consider this taxon a subgenus of ''Argynnis'', but it has been re ...
'' fritillaries, it is still closely related to them. Some of the differences are: variegated fritillaries have two or three broods per year vs. one per year in ''Speyeria''; they are nomadic vs. sedentary; and they use a wide range of host plants vs. just
violets Violet identifies various plant taxa, particularly species in the genus ''Viola'', within which the common violet is the best known member in Eurasia and the common blue violet and common purple violet are the best known members in North America ...
. And because of their use of
passionflower ''Passiflora'', known also as the passion flowers or passion vines, is a genus of about 550 species of flowering plants, the type genus of the family Passifloraceae. They are mostly tendril-bearing vines, with some being shrubs or trees. The ...
s as a host plant, variegated fritillaries also have
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. ...
links to the heliconians. Their flight is low and swift, but even when resting or nectaring, this species is extremely difficult to approach, and, because of this, its genus name was taken from the Greek word ''euptoietos'' meaning "easily scared".Rich Cech and Guy Tudor (2005). ''The Butterflies of the East Coast''. Princeton University Press, Princeton, New Jersey.


Description

The upperside of the wings is checkered with orange and black. Both the forewing and hindwing have a row of submarginal black spots and black median lines running across the wings.Bob Stewart, Priscilla Brodkin, and Hank Brodkin (2001). ''Butterflies of Arizona''. West Coast Lady Press.Jim P. Brock and Kenn Kaufman (2003). ''Butterflies of North America''. Houghton Mifflin, New York, NY. The underside of the forewing is orange with a pale orange spot rimmed in black in the forewing cell. The underside of the hindwing is mottled with browns and grays with a pale postmedian band. There is no silvering. The wingspan measures 1.75–2.25 inches (44–57 mm).Ernest M. Shull (1987). ''The Butterflies of Indiana''. Indiana Academy of Science.


Similar species

In the variegated fritillary's range, the only similar species is the Mexican fritillary ('' Euptoieta hegesia''). The Mexican fritillary is brighter orange, the upper side of its hindwing basal area is unmarked, and the underside of its wings is plainer, with no submarginal spots or median black lines.


Flight period

This species may be seen flying from April to October in the south, while in the north it flies from summer to early fall.


Habitat

This butterfly is often found in open, disturbed habitats such as clover and alfalfa fields, pastures, fields, waste areas, roadsides, and mountain meadows.David C. Iftner, John A. Shuey, and John V. Calhoun (1992). ''Butterflies and Skippers of Ohio''. College of Biological Sciences and The Ohio State University.


Nectar plants

Here is a list of some of the flowers that the variegated fritillary uses as nectar plants: *Dogbane, ''
Apocynum ''Apocynum'', commonly known as dogbane or Indian hemp, is a small genus of the flowering plant family Apocynaceae. Its name comes from Ancient Greek , from "away" and "dog", referring to dogbane ('' Cionura erecta''), which was used to pois ...
'' species *Common milkweed, ''
Asclepias syriaca ''Asclepias syriaca'', commonly called common milkweed, butterfly flower, silkweed, silky swallow-wort, and Virginia silkweed, is a species of flowering plant. It is native to southern Canada and much of the United States east of the Rocky Moun ...
'' *Asters, ''
Aster Aster or ASTER may refer to: Biology * ''Aster'' (genus), a genus of flowering plants ** List of ''Aster'' synonyms, other genera formerly included in ''Aster'' and still called asters in English * Aster (cell biology), a cellular structure shap ...
'' sp. *Bearded beggarticks, '' Bidens aristosa'' *Thistles, ''
Cirsium ''Cirsium'' is a genus of perennial and biennial flowering plants in the Asteraceae, one of several genera known commonly as thistles. They are more precisely known as plume thistles. These differ from other thistle genera ('' Carduus'', ''Silyb ...
'' sp. *Coneflowers, ''
Echinacea ''Echinacea'' is a genus of herbaceous flowering plants in the daisy family. It has ten species, which are commonly called coneflowers. They are found only in eastern and central North America, where they grow in moist to dry prairies and ope ...
'' sp. *Fleabane, ''
Erigeron ''Erigeron'' () is a large genus of plants in the family Asteraceae. It is closely related to the genus '' Aster'' and the true daisies in the genus '' Bellis''. The genus has a cosmopolitan distribution in dry, mountainous areas and grassland, ...
'' sp. *Common boneset, ''
Eupatorium perfoliatum ''Eupatorium perfoliatum'', known as common boneset or just boneset, is a North American perennial plant in the family Asteraceae. It is a common native to the Eastern United States and Canada, widespread from Nova Scotia to Florida, west as far ...
'' *Alfalfa, ''
Medicago sativa Alfalfa () (''Medicago sativa''), also called lucerne, is a perennial flowering plant in the legume family Fabaceae. It is cultivated as an important forage crop in many countries around the world. It is used for grazing, hay, and silage, as we ...
'' *Red clover, ''
Trifolium pratense ''Trifolium pratense'', the red clover, is a herbaceous species of flowering plant in the bean family Fabaceae, native to Europe, Western Asia, and northwest Africa, but planted and naturalized in many other regions. Description Red clove ...
'' *Ironweed, ''
Vernonia ''Vernonia'' is a genus of about 350 species of forbs and shrubs in the Daisy family Asteraceae. Some species are known as ironweed. Some species are edible and of economic value. They are known for having intense purple flowers. There have been ...
'' sp.Judy Burris and Wayne Richards (2006). ''The Life Cycle of Butterflies''. Storey Publishing, North Adams, MA. *Sedum, ''
Sedum ''Sedum'' is a large genus of flowering plants in the family Crassulaceae, members of which are commonly known as stonecrops. The genus has been described as containing up to 600 species, subsequently reduced to 400–500. They are leaf succulen ...
''


Life cycle

Image:VariegatedFritillaryMating1.JPG, Preparing to mate Image:VariegatedFritillaryMating2.JPG, Beginning mating Image:VariegatedFritillaryMating3.JPG, Mating Males actively patrol for females. Females lay their pale-green or cream-colored eggs singly on host plant leaves and stems. The larva eats the leaves, flowers, and stems of the food plant.James A. Scott (1986). ''The Butterflies of North America''. Stanford University Press, Stanford, California. The larva is red with black subdorsal and spiracular stripes infused with white spotting. In many individuals, the white is more conspicuous than the black. The red middorsal stripe bears white (sometimes black) oval-shaped spots, one per segment.David L. Wagner (2005). ''Caterpillars of Eastern North America''. Princeton University Press, Princeton, New Jersey. It has six rows of black spines and has a pair of long, clubbed spines on the head.Thomas J. Allen, Jim P. Brock, and Jeffrey Glassberg (2005). ''Caterpillars in the Field and Garden''. Oxford University Press Inc., New York, NY. The chrysalis is mainly shiny white, with small black spots, a variable amount of brown markings, and orange and gold tubercules. Adults overwinter in the south and fly north each spring and summer. It has 2–3 broods per year.


Host plants

This is a list of host plants used by the variegated fritillary: *Five-wing spiderling, '' Boerhaavia intermedia'' *Arizona swallow-wort, '' Melastoma arizonicum'' *Babyslippers, ''
Hybanthus verticillatus ''Hybanthus'' (green-violet) is a genus of flowering plants in the family Violaceae. This genus name is Greek for "humpback flower", referring to the drooping pedicels of plants that are part of this genus. The genus is grossly polyphyletic and m ...
'' *Southern flax, '' Linum australe'' *Stiffstem flax, '' Linum rigidum'' *Grooved yellow flax, '' Linum sulcatum'' *Canadian moonseed, ''
Menispermum canadense ''Menispermum canadense'', the Canadian moonseed, common moonseed, or yellow parilla, is a flowering plant in the family Menispermaceae, native to eastern North America, from southern Canada south to northern Florida, and from the Atlantic coas ...
'' *Mexican moonseed, '' Menispermum mexicanum'' *Blue passionflower, ''
Passiflora caerulea ''Passiflora caerulea'', the blue passionflower, bluecrown passionflower or common passion flower, is a species of flowering plant native to South America. It has been introduced elsewhere. It is a vigorous, deciduous or semi-evergreen tendril vi ...
'' *Foetid passionflower, ''
Passiflora foetida ''Passiflora foetida'' (common names: stinking passionflower, wild maracuja, bush passion fruit, wild water lemon, stoneflower, love-in-a-mist, or running pop) is a species of passion flower that is native to the southwestern United States (south ...
'' *Purple passionflower, ''
Passiflora incarnata ''Passiflora incarnata'', commonly known as maypop, purple passionflower, true passionflower, wild apricot, and wild passion vine, is a fast-growing perennial vine with climbing or trailing stems. A member of the passionflower genus ''Passiflora ...
'' *Plantain, ''
Plantago ''Plantago'' is a genus of about 200 species of flowering plants in the family Plantaginaceae, commonly called plantains or fleaworts. The common name plantain is shared with the unrelated cooking plantain. Most are herbaceous plants, though a ...
'' species *Mayapple, ''
Podophyllum peltatum ''Podophyllum peltatum'' is an herbaceous perennial plant in the family Berberidaceae. Its common names are mayapple, American mandrake, wild mandrake, and ground lemon. It is widespread across most of the eastern United States and southeastern C ...
'' *Common purslane, ''
Portulaca oleracea ''Portulaca oleracea'' (common purslane, also known as little hogweed, or pursley) is an annual (actually tropical perennial in USDA growing zones 10–11) succulent in the family Portulacaceae. Description The plant may reach in height. It ...
'' *Yellow alder, ''
Turnera ulmifolia ''Turnera ulmifolia'', the ramgoat dashalong or yellow alder, is a species of plant of family Passifloraceae, native to Mexico and the West Indies. A recent study found that yellow alder potentiated the antibiotic activity against methicillin-res ...
'' *Lance-leaf stonecrop, ''
Sedum lanceolatum ''Sedum lanceolatum'' is a species of flowering plant in the family Crassulaceae known by the common names spearleaf stonecrop and lanceleaf stonecrop. It is native to western North America and occurs in western Canada and the United States. It ...
'' *American field pansy, '' Viola bicolor'' *Common blue violet, ''
Viola sororia ''Viola sororia'', known commonly as the common blue violet, is a short-stemmed herbaceous perennial plant that is native to eastern North America. It is known by a number of common names, including common meadow violet, purple violet, woolly bl ...
'' *Heartsease, ''
Viola tricolor ''Viola tricolor'' is a common European wild flower, growing as an annual or short-lived perennial. The species is also known as wild pansy, Johnny Jump up (though this name is also applied to similar species such as the yellow pansy), heartsea ...
'' subspecies


References


External links


Variegated Fritillary
Butterflies of Canada {{DEFAULTSORT:Euptoieta Claudia Argynnini Butterflies of North America Butterflies of Central America Nymphalidae of South America Lepidoptera of Chile Butterflies described in 1775 Taxa named by Pieter Cramer