Euconulidae is a
taxonomic family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of minute, air-breathing
land snails,
terrestrial
Terrestrial refers to things related to land or the planet Earth.
Terrestrial may also refer to:
* Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate gastropod mollusk
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is es ...
s or
micromollusk
A micromollusk is a shelled mollusk which is extremely small, even at full adult size. The word is usually, but not exclusively, applied to marine mollusks, although in addition, numerous species of land snails and freshwater mollusks also ...
s in the superfamily
Trochomorphoidea
Trochomorphoidea is a taxonomic superfamily of small to large terrestrial pulmonate gastropod mollusks, that belong to the infraorder Limacoidei.MolluscaBase eds. (2021). MolluscaBase. Trochomorphoidea Möllendorff, 1890. Accessed through: Worl ...
.
This land snail family is closely allied to the Zonitidae, the glass snails.
Taxonomy
The family Euconulidae was originally placed within the superfamily
Gastrodontoidea
Gastrodontoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the limacoid clade.
Taxonomy
According to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), families in this superfamily ...
according to the
taxonomy of the Gastropoda (Bouchet & Rocroi, 2005)
The taxonomy of the Gastropoda as it was revised in 2005 by Philippe Bouchet and Jean-Pierre Rocroi is a system for the scientific classification of gastropod mollusks. (Gastropods are a taxonomic class of animals which consists of snails and ...
.
Since 2017, its classification has been revised and it now belongs to the superfamily
Trochomorphoidea
Trochomorphoidea is a taxonomic superfamily of small to large terrestrial pulmonate gastropod mollusks, that belong to the infraorder Limacoidei.MolluscaBase eds. (2021). MolluscaBase. Trochomorphoidea Möllendorff, 1890. Accessed through: Worl ...
Distribution
The distribution of the Euconulidae includes the
Nearctic
The Nearctic realm is one of the eight biogeographic realms constituting the Earth's land surface.
The Nearctic realm covers most of North America, including Greenland, Central Florida, and the highlands of Mexico. The parts of North America ...
, the western-
Palearctic
The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa.
The realm consists of several bioregions: the Euro-Si ...
, the eastern-Palearctic, the
Neotropical
The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone.
Definition
In bi ...
zone, the
Ethiopia
Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the Er ...
n zone,
Malagasy, south-eastern Asia, Australia, Polynesia and
Hawaii
Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only ...
.
Humidity, temperature, rainfall, and foliar dripping derived from dew, mist, and rain, affect the behavior and substrate selection of small terrestrial molluscs, such as ''Tikoconus costarricanus'', which inhabit shrubs in humid tropical montane forests. There is also a preference of some parts of a leaf, for example, in some cases the lower side is preferred, possibly because it has better humidity and protects small snails from the impact of raindrops.
Shell description
These minute snails have a
shell which is roundly conical and broad-based, like the shape of an old-fashioned European woven bee hive or
skep
A beehive is an enclosed structure in which some honey bee species of the subgenus ''Apis'' live and raise their young. Though the word ''beehive'' is commonly used to describe the nest of any bee colony, scientific and professional literature ...
. For this reason these snails are sometimes known as "hive snails".
The shells of most Euconulidae are only about 3 mm in size, amber-colored and translucent.
Anatomy
In this family, the number of
haploid chromosome
A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins ar ...
s lies between 26 and 30 (according to the values in this table).
[Barker G. M.: Gastropods on Land: ''Phylogeny, Diversity and Adaptive Morphology''. in Barker G. M. (ed.): ]
The biology of terrestrial molluscs
'. CABI Publishing, Oxon, UK, 2001, . 1-146, cited pages: 139 and 142.
Genera
Subfamilies and genera in the family Euconulidae include:
Euconulinae
* ''
Afroconulus
''Afroconulus'' is a genus of gastropods belonging to the family Euconulidae.
The species of this genus are found in Africa.
Species:
*''Afroconulus concavispira''
*''Afroconulus diaphanus''
*''Afroconulus iredalei''
*''Afroconulus roseus'' ...
'' Van Mol & van Bruggen, 1971
[MolluscaBase (2018). Euconulinae H.B. Baker, 1928. Accessed through: World Register of Marine Species at: http://marinespecies.org/aphia.php?p=taxdetails&id=994926 on 2018-09-14]
* ''
Afroguppya
''Afroguppya'' is a genus of gastropods belonging to the family Euconulidae
Euconulidae is a taxonomic family of minute, air-breathing land snails, terrestrial pulmonate gastropod mollusks or micromollusks in the superfamily Trochomorphoidea.
...
'' de Winter & Bruggen, 1992
* ''
Afropunctum
''Afropunctum'' is a genus of gastropods belonging to the family Euconulidae.
The species of this genus are found in Northern Europe and Africa.
Species:
*''Afropunctum quadrisculptum''
*''Afropunctum seminium
''Afropunctum'' is a genus of g ...
'' F. Haas, 1934
* ''
Cancelloconus'' I. Rensch, 1932
* ''
Coneuplecta'' Möllendorff, 1893
* ''
Diepenheimia'' Preston, 1913
* ''
Discoconulus'' Reinhardt, 1883
* ''
Dryachloa'' F. G. Thompson & H. G. Lee, 1980
* ''
Euconulus
''Euconulus'' is a genus of very small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Euconulidae, the hive snails. MolluscaBase eds. (2020). MolluscaBase. Euconulus Reinhardt, 1883. Accessed through: World Reg ...
'' Reinhardt, 1883 - type genus of the family Euconulidae
* ''
Guppya
''Guppya'' is a genus of gastropods belonging to the family Euconulidae.
The species of this genus are found in America, Africa, Malesia.
Species:
*'' Guppya angasi''
*'' Guppya bauri''
*'' Guppya biolleyi''
*'' Guppya capsula''
*'' Guppya ...
'' Mörch, 1867
* ''
Habroconus'' Crosse & P. Fischer, 1872
* ''
Kororia'' H. B. Baker, 1941
* ''
Louisia'' Godwin-Austen, 1908
* ''
Luchuconulus'' Pilsbry, 1928
* ''
Palaua'' H.B. Baker, 1941
* ''
Papuarion'' Van Mol, 1973
* ''
Parasitala'' Thiele, 1931
* ''
Sabalimax'' Tillier & Bouchet, 1989
* ''
Serostena'' Iredale, 1941
* ''
Tikoconus'' Barrientos, 2019
* ''
Velifera'' W.G. Binney, 1879
Microcystinae
* ''
Allenoconcha''
Preston, 1913
* ''
Aukena'' Baker, 1940
[Bouchet P. & Abdou A. (2001). "Recent Extinct Land Snails (Euconulidae) from the Gambier Islands with Remarkable Apertural Barriers". '' Pacific Science'' 55(2): 121-127. .]
* ''
Buffetia''
Iredale, 1945 (nomen nudum)
* ''
Cookeana''
* ''
Diastole''
Gude, 1913
* ''
Faunulena''
* ''
Greenwoodoconcha''
* ''
Hiona''
Cooke, 1940
* ''
Innesoconcha
''Innesoconcha'' is a genus of four species of tiny glass-snails that are endemic to Australia's Lord Howe Island in the Tasman Sea
The Tasman Sea (Māori: ''Te Tai-o-Rēhua'', ) is a marginal sea of the South Pacific Ocean, situated betwe ...
''
Iredale, 1944
* ''
Iredaleoconcha
''Iredaleoconcha'' is a genus of air-breathing land snails or semislugs, terrestrial pulmonate gastropod
The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca c ...
''
Preston, 1913
* ''
Kaala''
* ''
Kusaiea
''Kusaiea'' is a genus of small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the family Euconulidae, the hive snails.
Species
Species within the genus ''Kusaiea'' include:
* ''Kusaiea frivola
''Kusaiea frivol ...
''
Baker, 1938
* ''
Lamprocystis
''Lamprocystis'' is a genus of small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Microcystinae in the family Euconulidae, the hive snails. MolluscaBase eds. (2021). MolluscaBase. Lamprocystis L. Pfeffer, 1 ...
''
* ''
Liardetia
''Liardetia'' is a genus of land snails in the family Euconulidae, the hive snails.
Species include:
* ''Liardetia boninensis''
* ''Liardetia doliolum''
* ''Liardetia indifferens''
* ''Liardetia samoensis''
* ''Liardetia sculpta''
* ''Liardetia ...
''
* ''
Melloconcha''
Iredale, 1944
* ''
Mendana''
* ''
Microcystis
''Microcystis'' is a genus of freshwater cyanobacteria that includes the harmful algal bloom-forming '' Microcystis aeruginosa''. Many members of a ''Microcystis'' community can produce neurotoxins and hepatotoxins, such as microcystin and cy ...
''
* ''
Nancibella
''Nancibella'' is a genus of air-breathing land snails or semislugs, terrestrial pulmonate gastropod mollusks in the family Euconulidae
Euconulidae is a taxonomic family of minute, air-breathing land snails, terrestrial pulmonate gastropod mo ...
''
* ''
Periclocystis''
* ''
Philonesia
''Philonesia'' is a genus of air-breathing land snails or semislugs, terrestrial pulmonate gastropod mollusks in the family Euconulidae.MolluscaBase eds. (2021). MolluscaBase. Philonesia Sykes, 1900. Accessed through: World Register of Marine ...
''
Sykes, 1900
* ''
Piena''
* ''
Pukaloa''
* ''
Tengchiena''
Cladogram
The following
cladogram
A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
shows the
phylogenic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
relationships of this family with the other families within the
limacoid clade:
[Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". '']Journal of Biogeography
The ''Journal of Biogeography'' is a Peer review, peer-reviewed scientific journal in biogeography that was established in 1974. It covers aspects of spatial, ecological, and historical biogeography. The founding editor-in-chief was David Watts (b ...
'' 27(2): 379-390.
JSTOR
References
External links
*
{{Taxonbar, from=Q120147