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''Etheostoma osburni'', the candy darter or finescale saddled darter, is a species of fish in the family
Percidae The Percidae are a family of ray-finned fish, part of the order Perciformes, which are found in fresh and brackish waters of the Northern Hemisphere. The majority are Nearctic, but there are also Palearctic species. The family contains more than ...
, a member of the group known as
darters The darters, anhingas, or snakebirds are mainly tropical waterbirds in the family Anhingidae, which contains a single genus, ''Anhinga''. There are four living species, three of which are very common and widespread while the fourth is rarer and ...
. This species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...
to the eastern
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where it is known only from the
Kanawha River The Kanawha River ( ) is a tributary of the Ohio River, approximately 97 mi (156 km) long, in the U.S. state of West Virginia. The largest inland waterway in West Virginia, its valley has been a significant industrial region of the stat ...
system in the states of
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth ar ...
and
West Virginia West Virginia is a state in the Appalachian, Mid-Atlantic and Southeastern regions of the United States.The Census Bureau and the Association of American Geographers classify West Virginia as part of the Southern United States while the Bur ...
.


Description

''Etheostoma osburni'' can reach a length of , though most only reach about . This species has a lifespan of up to three years. It spawns in April and May.NatureServe. 2017
''Etheostoma osburni''.
NatureServe Explorer V.7.1 Accessed 8 August 2017.
It is an invertivore, feeding on aquatic insect larvae and water mites.


Habitat and distribution

''Etheostoma osburni'' lives in a system of rivers, streams, and creeks in the central
Appalachian Mountains The Appalachian Mountains, often called the Appalachians, (french: Appalaches), are a system of mountains in eastern to northeastern North America. The Appalachians first formed roughly 480 million years ago during the Ordovician Period. They ...
. It can be found in rapid riffles in rocky riverbed habitat. It occurs in cold, cool, and warm waters, as long as the substrate is rocky and the water is clear. It tolerates fast currents.


Conservation

''Etheostoma osburni'' has a limited range, it has been recorded in more than 10 locations and does not have a severely fragmented distribution, so it has been designated a
near-threatened species A near-threatened species is a species which has been categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature as that may be vulnerable to endangerment in the near future, but it does not currently qualify fo ...
on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
. In 2018, US Fish and Wildlife Service designated it as a federally protected endangered species.< It is probably declining, however, due to threats from human activity. It prefers clear, unsilted waters, and increases in
silt Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension with water. Silt usually has a floury feel when ...
and
sediment Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sand an ...
may reduce populations by reducing tolerable habitat.


Taxonomy and etymology

''Etheostoma osburni'' was first formally described as ''Poecilichthys osburni'' in 1932 by the
American American(s) may refer to: * American, something of, from, or related to the United States of America, commonly known as the "United States" or "America" ** Americans, citizens and nationals of the United States of America ** American ancestry, pe ...
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish ( Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 33,400 species of fish had been described as of Octobe ...
s Carl Leavitt Hubbs and Milton Bernhard Trautman With the type locality given as Stony Creek which is a
tributary A tributary, or affluent, is a stream or river that flows into a larger stream or main stem (or parent) river or a lake. A tributary does not flow directly into a sea or ocean. Tributaries and the main stem river drain the surrounding drainage ...
of the
Greenbrier River The Greenbrier River is a tributary of the New River, long,McNeel, William P. "Greenbrier River." ''The West Virginia Encyclopedia''. Ken Sullivan, editor. Charleston, WV: West Virginia Humanities Council. 2006. . in southeastern West Virginia ...
in
Pocahontas County, West Virginia Pocahontas County is a county located in the U.S. state of West Virginia. As of the 2020 census, the population was 7,869. Its county seat is Marlinton. The county was established in 1821. It is named after the daughter of the Powhatan chief ...
. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honors the American
zoologist Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the Animal, animal kingdom, including the anatomy, structure, embryology, evolution, Biological clas ...
Raymond Carroll Osburn Raymond Carroll Osburn (January 4, 1872 – August 6, 1955) was an American zoologist. Biography Osburn was born on January 4, 1872, in Newark, Ohio. In 1898, he received his bachelor's degree from the Ohio State University, and continued there, ...
(1872-1955). The candy darter is considered to be closely related to the variegated darter (''E. variatum'').


See also

Etheostoma osburni


References


Further reading

*Kessinger, Brin E., "Utilizing conservation genetics as a strategy for recovering the endangered Candy Darter (Etheostoma osburni) in West Virginia" (2020). Graduate Theses, Dissertations, and Problem Reports. 7670. https://researchrepository.wvu.edu/etd/7670 *Leftwich, K. N., et al. (1996)
The candy darter (''Etheostoma osburni'') in Stony Creek, George Washington–Jefferson National Forest, Virginia. Trout Predation, Distribution, and Habitat.
Center for Aquatic Technology Transfer, USDA Forest Service. *Dunn, C. G. (2017) ''Habitat and Imperilment of the Candy Darter Etheostoma osburni in the New River Drainage, USA'', Master of Science Thesis, Virginia Tech. {{Taxonbar, from=Q305806 Fish of the United States osburni Taxa named by Carl Leavitt Hubbs Fish described in 1932 Taxonomy articles created by Polbot