Epinephelus Rivulatus
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''Epinephelus rivulatus'', the halfmoon grouper, halfmoon rockcod, Chinaman rockcod, Charlie court cod, green-finned rock cod, or white-dotted grouper, is a species of marine ray-finned fish, a grouper from the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
Epinephelinae which is part of the family Serranidae, which also includes the
anthias Anthias are members of the family Serranidae and make up the subfamily Anthiinae. Anthias make up a sizeable portion of the population of pink, orange, and yellow reef fishes seen swarming in most coral reef photography and film. The name An ...
and sea basses. It is found in the
Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...
region.


Description

''Epinephelus rivulatus'' has a body with a
standard length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish m ...
which is 2.7 to 3.2 times its depth. The dorsal profile of the head is convex and the intraorbital region is flat. The preopercle is angular with 1 or 2 small spines on the anterior side of the angle while the gill cover is convex. The dorsal fin contains 11 spines and 16-18 soft rays while the
anal fin Fins are distinctive anatomical features composed of bony spines or rays protruding from the body of a fish. They are covered with skin and joined together either in a webbed fashion, as seen in most bony fish, or similar to a flipper, as se ...
as 3 spines and 8 soft rays. The membrane between the spines is notably notched. The caudal fin is rounded. The colour of the body is reddish to greenish-brown, with each scale on the body having a small white or pale blue spot. Sometimes 4 irregular dark vertical bars are visible, with a 5th bar on the caudal peduncle. The head is dark brown to pale reddish in colour with pale blue vermiculations and white spots on lower jaw and upper lip. The pectoral fin is dusky with a dark red or reddish-brown coloured blotch at its base. The remaining fins are greenish-brown or greyish-brown marked with dark mottling. Its maximum length is , and weight reaches


Distribution

''Epinephelus rivulatus'' has a wide Indo-West Pacific distribution. It is found from the east African coast from the
Gulf of Aden The Gulf of Aden ( ar, خليج عدن, so, Gacanka Cadmeed 𐒅𐒖𐒐𐒕𐒌 𐒋𐒖𐒆𐒗𐒒) is a deepwater gulf of the Indian Ocean between Yemen to the north, the Arabian Sea to the east, Djibouti to the west, and the Guardafui Channe ...
south to South Africa and then east to
New Caledonia ) , anthem = "" , image_map = New Caledonia on the globe (small islands magnified) (Polynesia centered).svg , map_alt = Location of New Caledonia , map_caption = Location of New Caledonia , mapsize = 290px , subdivision_type = Sovereign st ...
, north to the Ogasawara Islands of southern Japan, south to Australia. Its range includes northern New Zealand, and it was first recorded from the
Kermadec Islands Marine Reserve Kermadec or de Kermadec may refer to: Geography * Kermadec Islands, a subtropical island arc in the South Pacific Ocean northeast of New Zealand * Kermadec Plate, a long narrow tectonic plate located west of the Kermadec Trench * Kermadec Trench, ...
north of New Zealand in 2015, after researchers examined hundreds of hours of unused documentary film footage. It is d=found in the Indian Ocean islands of the Chagos,
Comoros The Comoros,, ' officially the Union of the Comoros,; ar, الاتحاد القمري ' is an independent country made up of three islands in southeastern Africa, located at the northern end of the Mozambique Channel in the Indian Ocean. It ...
, Madagascar, Mauritius and the Seychelles, it does not occur in Fiji, the Red Sea or Persian Gulf.


Habitat and biology

''Epinephelus rivulatus'' occurs on coral reefs, over areas with rocky substrata, algal flats and seagrass beds. It feeds on fishes and crustaceans. It is found at depths of . It is a protogynous hermaphrodite. Sexually mature females were found between fork lengths of with half of them sexually mature at a fork length of . The maless were larger than females and their fork lengths were . These differ from site to site although the sex ratios were the same between sites, with roughly 5.5 females for every male. Off Western Australia the majority of the females were reproductively active between July and December coinciding relatively cool water temperatures and increasing daylight hours. Spawning occurred periodically with each female able to spawn at least twice in periods of two or three consecutive days. It is the only known host of the parasite '' Pseudorhabdosynochus inversus'', which inhabits its gills. They form aggregations to spawn.


Taxonomy

''Epinephelus rivulatus'' was first formally described as ''Serranus rivulatus'' in 1830 by the
French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...
zoologist
Achille Valenciennes Achille Valenciennes (9 August 1794 – 13 April 1865) was a French zoologist. Valenciennes was born in Paris, and studied under Georges Cuvier. His study of parasitic worms in humans made an important contribution to the study of parasitology. ...
(1794-1865) with the
type locality Type locality may refer to: * Type locality (biology) * Type locality (geology) See also * Local (disambiguation) * Locality (disambiguation) {{disambiguation ...
given as
Réunion Réunion (; french: La Réunion, ; previously ''Île Bourbon''; rcf, label= Reunionese Creole, La Rényon) is an island in the Indian Ocean that is an overseas department and region of France. It is located approximately east of the island ...
. It is a member of the ''
Epinephelus fasciatus The blacktip grouper (''Epinephelus fasciatus''), also known as the redbanded grouper, blacktipped cod, black-tipped rockcod, footballer cod, red-barred cod, red-barred rockcod, scarlet rock-cod or weathered rock-cod, is a species of marine ray-fi ...
'' species complex alongside ''E. fasciatus'', '' E. irroratus'', and '' E. retouti''.


Fisheries

''Epinephelus rivulatus'' is of interest to both recreational and commercial fisheries despite its relative small size.


References


External links


Fishes of Australia: Epinephelus rivulatusItis.org

WoRMS

Animaldiversity Web
{{Taxonbar, from=Q2543301 Epinephelus Fish described in 1830