Eopteranodon
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''Eopteranodon'' (meaning "dawn '' Pteranodon'' (toothless wing)") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
tapejarid Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primit ...
pterosaur Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 ...
from the Aptian-age
Lower Cretaceous Lower may refer to: *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is situated about five miles south west of Dursley, eig ...
Yixian Formation The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its exq ...
of
Beipiao City Beipiao () is a city in Chaoyang prefecture, Liaoning province, in Northeast China. It has a population of 202,807. The main industry in the area is coal mining. With vertical shafts of almost 1000m, these are some of the deepest coal mines in ...
, Liaoning, China. The genus was named in 2005 by paleontologists
Lü Junchang Lü Junchang (; 1965 – 9 October 2018) was a Chinese palaeontologist and professor at the Institute of Geology, Chinese Academy of Geological Sciences. An expert on Mesozoic reptiles, he described and named dozens of dinosaur and pterosaur taxa ...
and
Zhang Xingliao Zhang may refer to: Chinese culture, etc. * Zhang (surname) (張/张), common Chinese surname ** Zhang (surname 章), a rarer Chinese surname * Zhang County (漳县), of Dingxi, Gansu * Zhang River (漳河), a river flowing mainly in Henan * ''Zha ...
. The type and only species is ''Eopteranodon lii''.


Description and classification

''Eopteranodon'' is based on the
type specimen In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes th ...
or
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of sever ...
BPV-078, an incomplete skeleton and skull. Its skull, including a large crest, was toothless and similar to that of ''Pteranodon''. The skull lacks the point of the snout but it was in life less than 200 millimeters long (7.9 inches), and the animal had a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan o ...
of about 1.1 meters (3.6 feet). A second specimen, D2526, described in 2006, had a larger wingspan. Despite its similarities to ''Pteranodon'', ''Eopteranodon'' was not placed into a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
by its describers, who put it into the clade
Pteranodontia Pteranodontia is an extinct group of ornithocheiroid pterodactyloid pterosaurs that lived during the Late Cretaceous period (Coniacian to Maastrichtian stages) of North America and Africa. They were some of the most advanced pterosaurs, and posse ...
as ''
incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...
'' (uncertain position). Shortly thereafter, a
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
study of all known Yixian pterosaurs by the same scientists found it to be close to the azhdarchoids, noted for the crested genera '' Tapejara'' and '' Tupuxuara'', and the giant, long-necked '' Quetzalcoatlus''. A further analysis of other recently discovered forms, in 2006 still considered basal to (having split off earlier than) azhdarchoids, helped the original authors, along with
David Unwin David (; , "beloved one") (traditional spelling), , ''Dāwūd''; grc-koi, Δαυΐδ, Dauíd; la, Davidus, David; gez , ዳዊት, ''Dawit''; xcl, Դաւիթ, ''Dawitʿ''; cu, Давíдъ, ''Davidŭ''; possibly meaning "beloved one". w ...
, to place these species together with ''Eopteranodon'' in a new clade
Chaoyangopteridae Chaoyangopteridae (or chaoyangopterids) is a family of pterosaurs within the larger group Azhdarchoidea. Chaoyangopterids lived mostly during the Early Cretaceous period, though possible members, ''Microtuban'', ''Xericeps'' and ''Argentinadraco' ...
, the possible sister group of the Azhdarchidae.Lü, J., Unwin, D.M., Xu, L., and Zhang, X. (2008). "A new azhdarchoid pterosaur from the Lower Cretaceous of China and its implications for pterosaur phylogeny and evolution." ''Naturwissenschaften'' However, in 2014, in an analysis by Brian Andres and colleagues, ''Eopteranodon'' was recovered as a basal member of the clade Tapejaromorpha, closely related to the species "''
Sinopterus ''Sinopterus'' (meaning "Chinese wing") was a genus of tapejarid pterodactyloid pterosaur from the Aptian-age Lower Cretaceous Jiufotang Formation of Chaoyang, Liaoning, China. It was first described and named by Wang Xiaolin and Zhou Zhonghe. ...
''" ''
gui The GUI ( "UI" by itself is still usually pronounced . or ), graphical user interface, is a form of user interface that allows users to interact with electronic devices through graphical icons and audio indicator such as primary notation, inste ...
'' and ''
Nemicolopterus crypticus ''Nemicolopterus'' is a genus of tapejaromorph pterosaur, based on a very small specimen described as pterosaur size, the smallest known "adult" pterosaur to date. It lived in the Jehol Biota 120 million years ago. Discovery and naming The Genu ...
''. Their cladogram is shown on the left. Later, in 2017, in another phylogenetic analysis, ''Eopteranodon'' was considered a member of the family
Tapejaridae Tapejaridae (from a Tupi word meaning "the old being") are a family of pterodactyloid pterosaurs from the Cretaceous period. Members are currently known from Brazil, England, Hungary, Morocco, Spain, the United States, and China. The most primi ...
based on proportions of the crest on the lower jaw and the limbs. This concept has been followed by several studies in 2019, including the one by Borja Holgado and colleagues,Borja Holgado, Rodrigo V. Pêgas, José Ignacio Canudo, Josep Fortuny, Taissa Rodrigues, Julio Company & Alexander W.A. Kellner, 2019, "On a new crested pterodactyloid from the Early Cretaceous of the Iberian Peninsula and the radiation of the clade Anhangueria", ''Scientific Reports'' 9: 4940 and the one by
Alexander Kellner Alexander Wilhelm Armin Kellner (born September 26, 1961) is a Brazilian geologist and paleontologist who is a leading expert in the field of studying pterosaurs. His research has focused mainly on fossil reptiles from the Cretaceous Period, i ...
and colleagues. The cladogram on the right is based on the topology made by Kellner and colleagues. Topology 1: Andres ''et al.'' (2014). Topology 2: Kellner ''et al.'' (2019).


See also

*
List of pterosaur genera This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered inval ...
*
Timeline of pterosaur research This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs w ...


References


External links


''Eopteranodon''
in The Pterosauria
at The Grave Yard--> {{Portal bar, Paleontology, Cretaceous, China Early Cretaceous pterosaurs of Asia Prehistoric animals of China Tapejaromorphs Fossil taxa described in 2005 Taxa named by Lü Junchang