Echium Horridum
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''Echium'' is a genus of approximately 70 species and several
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of flowering plant in the family Boraginaceae. Species of ''Echium'' are
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to North Africa, mainland Europe to Central Asia, and the
Macaronesian islands Macaronesia (Portuguese: ''Macaronésia,'' Spanish: ''Macaronesia'') is a collection of four volcanic archipelagos in the North Atlantic, off the coasts of Africa and Europe. Each archipelago is made up of a number of Atlantic oceanic islands ...
where the genus reaches its maximum diversity. 29 species of ''Echium'' are endemic to the Canary, Madeira, and Cape Verde archipelagos. The continental species are
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
, whereas all but two of the endemic species of the Macaronesian islands are woody perennial shrubs.


Etymology

The Latin genus name ''echium'' comes from the Greek ''echion'' referring to
Echium plantagineum ''Echium plantagineum'', commonly known as purple viper's-bugloss or Patterson's curse, is a species of the genus ''Echium'' native to western and southern Europe (from southern England south to Iberia and east to the Crimea), northern Africa, a ...
and itself deriving from ''echis'' "
viper The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
"; the Greek term dates to Dioscorides who noted a resemblance between the shape of the nutlets to a viper’s head. The genus Echium was published by Carl Linnaeus in 1753.


Cultivation and uses

Many species are used as ornamental and garden plants and may be found in suitable climates throughout the world. In Crete ''Echium italicum'' is called pateroi (πάτεροι) or voidoglosses (βοϊδόγλωσσες) and its tender shoots are eaten boiled or steamed. ''Echium'' species are used as food plants by the larvae of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species including '' Coleophora onosmella'' and orange swift.


Echium seed oil

The seed oil from ''Echium plantagineum'' contains high levels of alpha linolenic acid (ALA), gamma linolenic acid (GLA) and stearidonic acid (SDA), making it valuable in cosmetic and skin care applications, with further potential as a functional food, as an alternative to fish oils. However, despite it's high ALA content, echium seed oil does not increase docosahexaenoic acid (DHA) and eicosapentaenoic acid (EPA) levels.


Invasiveness

Some species have been widely naturalized in Mediterranean climates, including South Africa, Australia, New Zealand and parts of South America and the United States. For example, ''
Echium plantagineum ''Echium plantagineum'', commonly known as purple viper's-bugloss or Patterson's curse, is a species of the genus ''Echium'' native to western and southern Europe (from southern England south to Iberia and east to the Crimea), northern Africa, a ...
'' has become a major invasive species in Australia.


Species

#'' Echium acanthocarpum'' Svent. #''
Echium aculeatum ''Echium aculeatum'' is a species of flowering plant of the family Boraginaceae. It is endemic to the Canary Islands, where it occurs on the islands El Hierro, La Palma, La Gomera and Tenerife.Manuel Arechavaleta, S. Rodríguez, Nieves Zu ...
'' Poir. #''
Echium albicans ''Echium albicans'', commonly known as white-leaved bugloss, is a species of purple coloured perennial plants from family Boraginaceae, found in Andalucían mountains and in Marbella Marbella ( , , ) is a city and municipality in southern Spa ...
'' Lag. & Rodr. #''
Echium amoenum ''Echium amoenum'' (In Persian: گل گاوزبان) is one of the important medicinal herbs in Iranian traditional medicine. It is a biennial or perennial herb indigenous to the narrow zone of northern part of Iran, Caucasus and Russia, wh ...
'' Fisch. & C.A.Mey #'' Echium anchusoides'' Bacch., Brullo & Selvi #''
Echium angustifolium ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands where ...
'' Lam. #'' Echium arenarium'' Guss. #''
Echium asperrimum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Lam. #''
Echium auberianum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family (biology), family Boraginaceae. Species of ''Echium'' are native plant, native to North Africa, mainland Europe to Central Asia, and ...
'' Webb & Berthel. #''
Echium bethencourtii ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Santos. #''
Echium biebersteinii ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Lacaita. #''
Echium boissieri ''Echium boissieri'' is a species of flowering plant in the borage family Boraginaceae. It is native to the western Mediterranean Basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Medit ...
'' Steud. #''
Echium bonnetii ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Coincy. #''
Echium brevirame ''Echium brevirame'' is a species of flowering plants of the family Boraginaceae. It is endemic to the Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and a ...
'' Sprague & Hutch. #'' Echium callithyrsum'' Webb ex Bolle. #''
Echium candicans ''Echium candicans'', the 'Pride of Madeira', is a species of flowering plant in the family Boraginaceae, native to the island of Madeira. It is a large herbaceous perennial subshrub, growing to . In the first year after germination, the plant p ...
'' L.f. #'' Echium canum'' Emb. & Maire #'' Echium clandestinum'' Pomel #''
Echium creticum ''Echium creticum'', the Cretan viper's bugloss, (syn. ''Echium calycinum'' Viv., ''Echium rubrum'' Moench, ''Echium sericeum'' Vahl, ''Echium violaceum'' L.) is a species of flowering plant in the family Boraginaceae. It is native to the ...
'' L. #''
Echium decaisnei ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands where ...
'' Webb & Berthel. #''
Echium flavum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands where ...
'' Desf. #''
Echium gaditanum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Boiss. #''
Echium giganteum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' L.f. #''
Echium glomeratum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Poir. #''
Echium handiense ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Svent. #''
Echium hierrense ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Webb ex Bolle #'' Echium horridum'' Batt. #''
Echium humile ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands where ...
'' Desf. #''
Echium hypertropicum ''Echium hypertropicum'' is a species of flowering plants of the family Boraginaceae. The species is endemic to Cape Verde.Oromí, Martín, Zurita & Cabrera, 2005 : Lista preliminar de especies silvestres de Cabo Verde: Hongos, Plantas y Animales ...
'' Webb. #''
Echium italicum ''Echium italicum'', the Italian viper's bugloss, Lady Campbell weed,John H. Wiersema and Blanca León or pale bugloss, is a species of plant from the family Boraginaceae, found in the Mediterranean Basin (especially in Italy, hence the name 'i ...
'' L. #''
Echium judaeum ''Echium judaeum'', commonly known as the Judean viper's bugloss, is an annual plant endemic to southern Lebanon, southern Syria and Israel, of the Boraginaceae family, and which, like other herbaceous flowering plants of the same genus, deriv ...
'' Lacaita. #''
Echium khuzistanicum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family (biology), family Boraginaceae. Species of ''Echium'' are native plant, native to North Africa, mainland Europe to Central Asia, and ...
'' Mozaff. #''
Echium × lemsii ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' G.Kunkel. #''
Echium leucophaeum ''Echium leucophaeum'' is a species of flowering plants of the family Boraginaceae. It is endemic to the Canary Islands, where it is restricted to the island of Tenerife.Manuel Arechavaleta, S. Rodríguez, Nieves Zurita, A. García (Hrsg.): Lis ...
'' Webb ex Sprague & Hutch. #''
Echium × lidii ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' G.Kunkel #''
Echium longifolium ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Delile. #''
Echium lusitanicum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands where ...
'' L. #''
Echium modestum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Ball. #''
Echium nervosum ''Echium nervosum'' is a species of flowering plants of the family Boraginaceae. It is endemic to Madeira. The specific name ''nervosum'' is from Latin and means "veined". Description It is a compact woody shrub, average height , maximum height ...
'' W.T. Aiton #'' Echium onosmifolium'' Webb & Berthel. #''
Echium orientale ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' L. #''
Echium pabotii ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family (biology), family Boraginaceae. Species of ''Echium'' are native plant, native to North Africa, mainland Europe to Central Asia, and ...
'' Mouterde. #''
Echium parviflorum ''Echium parviflorum'' is a species of plant in the family Boraginaceae native to the Mediterranean Basin In biogeography, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands ...
'' Moench #''
Echium petiolatum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Barratte & Coincy. #''
Echium pininana ''Echium pininana'', commonly known as the tree echium, pine echium, giant viper's-bugloss, or tower of jewels, is a species of flowering plant in the borage family Boraginaceae. It is endemic to the Canary Islands, where it is restricted to t ...
'' Webb et Berthel. #'' Echium pitardii'' A.Chev. #''
Echium plantagineum ''Echium plantagineum'', commonly known as purple viper's-bugloss or Patterson's curse, is a species of the genus ''Echium'' native to western and southern Europe (from southern England south to Iberia and east to the Crimea), northern Africa, a ...
'' L. #'' Echium portosanctense'' J.A.Carvalho, Pontes, Bat.-Marques & R.Jardim #''
Echium rauwolfii ''Echium rauwolfii'' is a species of flowering plant in the family Boraginaceae, whose native range is North Africa to the Arabian Peninsula. Description ''Echium rauwolfii'' is an annual. It is not a succulent. The leaves are alternate, and ...
'' Delile. #''
Echium rosulatum ''Echium rosulatum'' is a species of flowering plant in the family Boraginaceae. It is endemic to the western Iberian Peninsula in Portugal, Galicia (Spain) and the Sierra Morena. It favors acidic and somewhat nitrified soil. It has two subs ...
'' Lange #'' Echium rubrum'' Forssk. #''
Echium sabulicola ''Echium sabulicola'' (syn. ''Echium confusum'' Coincy, ''Echium maritimum'' Willd.) is a plant in the genus '' Echium''. It is a herbaceous biennial plant and grows up to in height. It is native to the sandy areas of the coasts of the western ...
'' Pomel. #''
Echium salmanticum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Lag. #'' Echium simplex'' DC. #'' Echium spurium'' Lojac. #''
Echium stenosiphon ''Echium stenosiphon'' is a plant species of the family Boraginaceae. The species is endemic to Cape Verde.Oromí, Martín, Zurita & Cabrera, 2005 : Lista preliminar de especies silvestres de Cabo Verde: Hongos, Plantas y Animales Terrestres.' G ...
'' Webb. #''
Echium strictum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands where ...
'' L.f. #''
Echium suffruticosum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Barratte. #''
Echium sventenii ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Bramwell. #''
Echium x taibiquense ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' P.Wolff & Rosinski. #''
Echium tenue ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Roth. #''
Echium thyrsiflorum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Masson ex Link. #'' Echium triste'' Svent. #'' Echium trygorrhizum'' Pomel. #''
Echium tuberculatum ''Echium tuberculatum'' is a species of flowering plant native to Portugal, Morocco, Algeria and Libya in the Iberian Peninsula and North Africa respectively. This plant is a hardy garden perennial and it forms a sprawling mass of red flowers ...
'' Hoffmanns. & Link. #''
Echium velutinum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands w ...
'' Coincy. #''
Echium virescens ''Echium virescens'' is a flowering plant in the genus ''Echium''. It is endemic to the island of Tenerife, mainly in Macizo de Anaga and the Orotava Valley. It grows in forests and on lower south slopes of the island. Description It is a herb ...
'' DC. #''
Echium vulcanorum ''Echium vulcanorum'' is a species of flowering plants of the family Boraginaceae. The species is endemic to Cape Verde.Oromí, Martín, Zurita & Cabrera, 2005 : Lista preliminar de especies silvestres de Cabo Verde: Hongos, Plantas y Animales Te ...
'' A.Chev. #'' Echium vulgare'' L. #''
Echium webbii ''Echium webbii'' is a species of flowering plants of the family Boraginaceae. It is endemic to the Canary Islands, where it is restricted to the island of La Palma.Manuel Arechavaleta, S. Rodríguez, Nieves Zurita, A. García (Hrsg.): Lista de e ...
'' Coincy. #''
Echium wildpretii ''Echium wildpretii'' is a species of flowering plant in the family Boraginaceae. It is an herbaceous biennial plant that grows up to in height. The species is endemic to the Canary Islands, and is found mainly on Mount Teide in Tenerife. T ...
'' Pears. ex Hook. fil.


References

{{Taxonbar, from=Q157711 Boraginaceae genera Taxa named by Carl Linnaeus