Echium Angustifolium
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''Echium'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of approximately 70
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
and several
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Boraginaceae Boraginaceae, the borage or forget-me-not family, includes about 2,000 species of shrubs, trees and herbs in 146, to 156 genera with a worldwide distribution. The APG IV system from 2016 classifies the Boraginaceae as single family of the order ...
. Species of ''Echium'' are
native Native may refer to: People * Jus soli, citizenship by right of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Native Americans (disambiguation) In arts and entert ...
to North Africa, mainland Europe to Central Asia, and the
Macaronesian islands Macaronesia (Portuguese: ''Macaronésia,'' Spanish: ''Macaronesia'') is a collection of four volcanic archipelagos in the North Atlantic, off the coasts of Africa and Europe. Each archipelago is made up of a number of Atlantic oceanic islands ...
where the genus reaches its maximum diversity. 29 species of ''Echium'' are endemic to the Canary, Madeira, and Cape Verde archipelagos. The continental species are
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
, whereas all but two of the endemic species of the Macaronesian islands are woody perennial shrubs.


Etymology

The Latin genus name ''echium'' comes from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
''echion'' referring to
Echium plantagineum ''Echium plantagineum'', commonly known as purple viper's-bugloss or Patterson's curse, is a species of the genus ''Echium'' native to western and southern Europe (from southern England south to Iberia and east to the Crimea), northern Africa, a ...
and itself deriving from ''echis'' "
viper The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs tha ...
"; the Greek term dates to
Dioscorides Pedanius Dioscorides ( grc-gre, Πεδάνιος Διοσκουρίδης, ; 40–90 AD), “the father of pharmacognosy”, was a Greek physician, pharmacologist, botanist, and author of ''De materia medica'' (, On Medical Material) —a 5-vol ...
who noted a resemblance between the shape of the nutlets to a viper’s head. The genus Echium was published by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in 1753.


Cultivation and uses

Many species are used as ornamental and garden plants and may be found in suitable climates throughout the world. In
Crete Crete ( el, Κρήτη, translit=, Modern: , Ancient: ) is the largest and most populous of the Greek islands, the 88th largest island in the world and the fifth largest island in the Mediterranean Sea, after Sicily, Sardinia, Cyprus, and ...
''Echium italicum'' is called pateroi (πάτεροι) or voidoglosses (βοϊδόγλωσσες) and its tender shoots are eaten boiled or steamed. ''Echium'' species are used as food plants by the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e of some
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
species including ''
Coleophora onosmella ''Coleophora pennella'' is a moth of the family Coleophoridae. It is found in most of Europe. The wingspan is . Adults are on wing from June to July. The larvae feed on alkanet (''Anchusa officinalis''), houndstongue (''Cynoglossum officinale'' ...
'' and
orange swift The orange swift or orange moth (''Triodia sylvina'') is a moth belonging to the family Hepialidae. The species was first described by Carl Linnaeus in 1761 and was previously placed in the genus ''Hepialus''. It is distributed throughout Europ ...
.


Echium seed oil

The seed oil from ''Echium plantagineum'' contains high levels of alpha linolenic acid (ALA), gamma linolenic acid (GLA) and
stearidonic acid Stearidonic acid (SDA: C18 H28 O2; 18: 4, n-3) is an ω-3 fatty acid, sometimes called moroctic acid. It is biosynthesized from alpha-linolenic acid (ALA: C18H30O2; 18:3, n-3) by the enzyme delta-6-desaturase, that removes two hydrogen (H) atoms ...
(SDA), making it valuable in cosmetic and skin care applications, with further potential as a functional food, as an alternative to fish oils. However, despite it's high ALA content, echium seed oil does not increase
docosahexaenoic acid Docosahexaenoic acid (DHA) is an omega-3 fatty acid that is a primary structural component of the human brain, cerebral cortex, skin, and retina. In physiological literature, it is given the name 22:6(n-3). It can be synthesized from alpha-lino ...
(DHA) and
eicosapentaenoic acid Eicosapentaenoic acid (EPA; also icosapentaenoic acid) is an omega-3 fatty acid. In physiological literature, it is given the name 20:5(n-3). It also has the trivial name timnodonic acid. In chemical structure, EPA is a carboxylic acid with a 20-c ...
(EPA) levels.


Invasiveness

Some species have been widely naturalized in Mediterranean climates, including South Africa, Australia, New Zealand and parts of South America and the United States. For example, ''
Echium plantagineum ''Echium plantagineum'', commonly known as purple viper's-bugloss or Patterson's curse, is a species of the genus ''Echium'' native to western and southern Europe (from southern England south to Iberia and east to the Crimea), northern Africa, a ...
'' has become a major
invasive species in Australia Invasive species in Australia are a serious threat to the native biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic ('' genetic variabi ...
.


Species

#'' Echium acanthocarpum'' Svent. #'' Echium aculeatum'' Poir. #'' Echium albicans'' Lag. & Rodr. #'' Echium amoenum'' Fisch. & C.A.Mey #'' Echium anchusoides'' Bacch., Brullo & Selvi #'' Echium angustifolium'' Lam. #'' Echium arenarium'' Guss. #'' Echium asperrimum'' Lam. #'' Echium auberianum'' Webb & Berthel. #'' Echium bethencourtii'' Santos. #'' Echium biebersteinii'' Lacaita. #'' Echium boissieri'' Steud. #'' Echium bonnetii'' Coincy. #'' Echium brevirame'' Sprague & Hutch. #'' Echium callithyrsum'' Webb ex Bolle. #''
Echium candicans ''Echium candicans'', the 'Pride of Madeira', is a species of flowering plant in the family Boraginaceae, native to the island of Madeira. It is a large herbaceous perennial subshrub, growing to . In the first year after germination, the plant p ...
'' L.f. #'' Echium canum'' Emb. & Maire #'' Echium clandestinum'' Pomel #'' Echium creticum'' L. #'' Echium decaisnei'' Webb & Berthel. #''
Echium flavum ''Echium'' is a genus of approximately 70 species and several subspecies of flowering plant in the family Boraginaceae. Species of ''Echium'' are native to North Africa, mainland Europe to Central Asia, and the Macaronesian islands where ...
'' Desf. #'' Echium gaditanum'' Boiss. #'' Echium giganteum'' L.f. #'' Echium glomeratum'' Poir. #'' Echium handiense'' Svent. #'' Echium hierrense'' Webb ex Bolle #'' Echium horridum'' Batt. #'' Echium humile'' Desf. #''
Echium hypertropicum ''Echium hypertropicum'' is a species of flowering plants of the family Boraginaceae. The species is endemic to Cape Verde.Oromí, Martín, Zurita & Cabrera, 2005 : Lista preliminar de especies silvestres de Cabo Verde: Hongos, Plantas y Animales ...
'' Webb. #''
Echium italicum ''Echium italicum'', the Italian viper's bugloss, Lady Campbell weed,John H. Wiersema and Blanca León or pale bugloss, is a species of plant from the family Boraginaceae, found in the Mediterranean Basin (especially in Italy, hence the name 'i ...
'' L. #'' Echium judaeum'' Lacaita. #'' Echium khuzistanicum'' Mozaff. #'' Echium × lemsii'' G.Kunkel. #'' Echium leucophaeum'' Webb ex Sprague & Hutch. #'' Echium × lidii'' G.Kunkel #'' Echium longifolium'' Delile. #'' Echium lusitanicum'' L. #'' Echium modestum'' Ball. #'' Echium nervosum'' W.T. Aiton #'' Echium onosmifolium'' Webb & Berthel. #'' Echium orientale'' L. #'' Echium pabotii'' Mouterde. #'' Echium parviflorum'' Moench #'' Echium petiolatum'' Barratte & Coincy. #''
Echium pininana ''Echium pininana'', commonly known as the tree echium, pine echium, giant viper's-bugloss, or tower of jewels, is a species of flowering plant in the borage family Boraginaceae. It is endemic to the Canary Islands, where it is restricted to t ...
'' Webb et Berthel. #'' Echium pitardii'' A.Chev. #''
Echium plantagineum ''Echium plantagineum'', commonly known as purple viper's-bugloss or Patterson's curse, is a species of the genus ''Echium'' native to western and southern Europe (from southern England south to Iberia and east to the Crimea), northern Africa, a ...
'' L. #'' Echium portosanctense'' J.A.Carvalho, Pontes, Bat.-Marques & R.Jardim #'' Echium rauwolfii'' Delile. #'' Echium rosulatum'' Lange #'' Echium rubrum'' Forssk. #'' Echium sabulicola'' Pomel. #'' Echium salmanticum'' Lag. #'' Echium simplex'' DC. #'' Echium spurium'' Lojac. #''
Echium stenosiphon ''Echium stenosiphon'' is a plant species of the family Boraginaceae. The species is endemic to Cape Verde.Oromí, Martín, Zurita & Cabrera, 2005 : Lista preliminar de especies silvestres de Cabo Verde: Hongos, Plantas y Animales Terrestres.' G ...
'' Webb. #'' Echium strictum'' L.f. #'' Echium suffruticosum'' Barratte. #'' Echium sventenii'' Bramwell. #'' Echium x taibiquense'' P.Wolff & Rosinski. #'' Echium tenue'' Roth. #'' Echium thyrsiflorum'' Masson ex Link. #'' Echium triste'' Svent. #'' Echium trygorrhizum'' Pomel. #'' Echium tuberculatum'' Hoffmanns. & Link. #'' Echium velutinum'' Coincy. #''
Echium virescens ''Echium virescens'' is a flowering plant in the genus ''Echium''. It is endemic to the island of Tenerife, mainly in Macizo de Anaga and the Orotava Valley. It grows in forests and on lower south slopes of the island. Description It is a herb ...
'' DC. #''
Echium vulcanorum ''Echium vulcanorum'' is a species of flowering plants of the family Boraginaceae. The species is endemic to Cape Verde.Oromí, Martín, Zurita & Cabrera, 2005 : Lista preliminar de especies silvestres de Cabo Verde: Hongos, Plantas y Animales Te ...
'' A.Chev. #''
Echium vulgare ''Echium vulgare'', known as viper's bugloss and blueweed,Dickinson, T.; Metsger, D.; Bull, J.; & Dickinson, R. (2004) ROM Field Guide to Wildflowers of Ontario. Toronto:Royal Ontario Museum, p. 203. is a species of flowering plant in the borag ...
'' L. #''
Echium webbii ''Echium webbii'' is a species of flowering plants of the family Boraginaceae. It is endemic to the Canary Islands, where it is restricted to the island of La Palma.Manuel Arechavaleta, S. Rodríguez, Nieves Zurita, A. García (Hrsg.): Lista de e ...
'' Coincy. #''
Echium wildpretii ''Echium wildpretii'' is a species of flowering plant in the family Boraginaceae. It is an herbaceous biennial plant that grows up to in height. The species is endemic to the Canary Islands, and is found mainly on Mount Teide in Tenerife. T ...
'' Pears. ex Hook. fil.


References

{{Taxonbar, from=Q157711 Boraginaceae genera Taxa named by Carl Linnaeus