Dysdera Spinicrus
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''Dysdera'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of woodlouse hunting spiders that was first described by
Pierre André Latreille Pierre André Latreille (; 29 November 1762 – 6 February 1833) was a French zoologist, specialising in arthropods. Having trained as a Roman Catholic priest before the French Revolution, Latreille was imprisoned, and only regained his freedom ...
in 1804. They originated from Central Asia to Central Europe. The family has gained many common names from their individual species, including the "European garden spider", the "slater-eating spider", the "sow-bug killer", the "woodlouse hunter", and the "woodlouse spider". A bite from one of these spiders can be painful due to their large fangs and wide jaw. It may leave an itchy, swollen, or red bump, but the
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
from one of their bites is not harmful to humans.


Description

Adults have a reddish-brown body and legs, and can grow up to long. Females are generally larger growing from , while males are about . Their six eyes are close together in an oval shape, and they have eight reddish legs, the second pair facing backward. ''Dysdera'' live in natural shelters, which they wrap by totally white silk. The inhabitants of hot and humid forest will take any potential shelter on or close to the ground. The shelters are used to hide from predators as well as for keeping the spider warm. During the day, they are commonly found taking shelter under objects like gravel with organic material covering it, stones, bark, and occasionally in suburban gardens.


Diet

''Dysdera'' are one of the few known arthropods to hunt and prey on
woodlice A woodlouse (plural woodlice) is an isopod crustacean from the polyphyleticThe current consensus is that Oniscidea is actually triphyletic suborder Oniscidea within the order Isopoda. They get their name from often being found in old wood. ...
, one of their main food sources. These spiders have wide jaws and large fangs to help to overcome the solid armor-like shells of
woodlice A woodlouse (plural woodlice) is an isopod crustacean from the polyphyleticThe current consensus is that Oniscidea is actually triphyletic suborder Oniscidea within the order Isopoda. They get their name from often being found in old wood. ...
. It makes them powerful predators for their size, allowing them to dominate or kill competitors, such as
centipedes Centipedes (from New Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', lip, and New Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, an a ...
or other spiders. '' D. crocata'' is the only species from the ''Dysdera'' family known to prey on other spiders. They can also excrete certain enzymes that neutralize the chemical defenses of potential prey, allowing them to subsist on other common ground-dwelling invertebrates, including
silverfish The silverfish (''Lepisma saccharinum'') is a species of small, primitive, wingless insect in the order Zygentoma (formerly Thysanura). Its common name derives from the insect's silvery light grey colour, combined with the fish-like appearance ...
,
earwigs Earwigs make up the insect order Dermaptera. With about 2,000 species in 12 families, they are one of the smaller insect orders. Earwigs have characteristic cerci, a pair of forcep-like pincers on their abdomen, and membranous wings folded ...
,
millipedes Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a resu ...
, and small burying beetles. '' D. unguimannis'' is considered the most remarkable case of
troglomorphism Troglomorphism is the morphological adaptation of an animal to living in the constant darkness of caves, characterised by features such as loss of pigment, reduced eyesight or blindness, and frequently with attenuated bodies and/or appendages. The ...
(adaptation to cave life) in the genus ''Dysdera''.


Mating

Mating is mainly done during the month of April. The female is the main caregiver for the young. After mating, the male has minimal to no role in the child rearing process. Before laying the eggs, females will make a silk pouch to protect and give them shelter. She can lay up to seventy eggs at once, and will stay in the silk pouch with the eggs, protecting them and waiting for them to hatch.


Distribution

The ''D. crocata'', '' D. ninnii'', '' D. dubrovninnii'', '' D. hungarica'', and '' D. longirostris'' are the five species still found in Central
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
after the last glacial period. They are also abundantly found in North African countries like
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
and
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...
, but also in
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
, the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, defi ...
, and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
. In the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
, ''Dysdera crocata'' is found from
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
down to
Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to the ...
, and all the way across the country in
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
. At least two species inhabit South America: '' D. solers'' in
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
- possibly a relict species from the post-miocene era- and '' D. magna'' in
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
,
Uruguay Uruguay (; ), officially the Oriental Republic of Uruguay ( es, República Oriental del Uruguay), is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast; while bordering ...
, and the central area of
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the east a ...
.


Canary Islands

''Dysdera'' inhabits all of the
Macaronesia Macaronesia (Portuguese: ''Macaronésia,'' Spanish: ''Macaronesia'') is a collection of four volcanic archipelagos in the North Atlantic, off the coasts of Africa and Europe. Each archipelago is made up of a number of List of islands in the At ...
n archipelagos, but the most drastic variety is in the
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
, a 22 million year old volcanic archipelago nearly off the northwestern coast of
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
. These islands house over forty endemic species of ''Dysdera'', thirty-six of which likely descended from a single ancestor, and six of which are associated with the oldest eastern island. On
Lanzarote Lanzarote (, , ) is a Spanish island, the easternmost of the Canary Islands in the Atlantic Ocean. It is located approximately off the north coast of Africa and from the Iberian Peninsula. Covering , Lanzarote is the fourth-largest of the i ...
and
Fuerteventura Fuerteventura () is one of the Canary Islands, in the Atlantic Ocean, part of the North Africa region, and politically part of Spain. It is located away from the northwestern coast of Africa. The island was declared a biosphere reserve by UNES ...
, the spider populations are limited to the highest elevation. The most likely reason that these spiders are so abundant on the Canary Islands is due to the abundance of species on the nearby Iberian Peninsula and North Africas. Groups like ''Dysdera crocata'' and ''Dysdera erythrina'', found on two neighboring lands, are found more often than '' D. lata'' and ''Dysdera longirostris'', found also in North Africa and Iberia. Over time, these spiders either made their way to the islands, then began evolving separately, or were separated when the islands broke off from the greater land masses. In total, two to four colonization events are assumed. This probably happened by
rafting Rafting and whitewater rafting are recreational outdoor activities which use an inflatable raft to navigate a river or other body of water. This is often done on whitewater or different degrees of rough water. Dealing with risk is often a ...
, or even more likely by transport on
floating island A floating island is a mass of floating aquatic plants, mud, and peat ranging in thickness from several centimeters to a few meters. Floating islands are a common natural phenomenon that are found in many parts of the world. They exist less co ...
s, for ''Dysdera'' is not known to use
ballooning Ballooning may refer to: * Hot air ballooning * Balloon (aeronautics) * Ballooning (spider) * Ballooning degeneration, a disease * Memory ballooning See also * Balloon (disambiguation) A balloon is a flexible container for (partially or fully) co ...
. ''Dydera lancerotensis'' is the only species where an independent origin from continental ancestors is unquestionable; it was originally described as a subspecies of ''Dysdera crocata''. While some of the remaining
Macaronesia Macaronesia (Portuguese: ''Macaronésia,'' Spanish: ''Macaronesia'') is a collection of four volcanic archipelagos in the North Atlantic, off the coasts of Africa and Europe. Each archipelago is made up of a number of List of islands in the At ...
n archipelagos have been colonized from the Canaries, the
Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...
have been independently colonized from the continent. The radiation of ''Dysdera'' is surpassed on the Canary Islands only by the
snail A snail is, in loose terms, a shelled gastropod. The name is most often applied to land snails, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gastro ...
genus ''
Napaeus ''Napaeus'' is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Eninae of the family Enidae.MolluscaBase eds. (2021). MolluscaBase. Napaeus Albers, 1850. Accessed through: World Register of Marine Sp ...
'', the
millipede Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a resu ...
genus '' Dolichoiulus'', and the
beetle Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 describ ...
genera '' Attalus'' and '' Laparocerus''.


Species

it contains 285 species. A study published in 2021 used an integrative approach combining morphological and molecular evidence to describe 8 species new to science as well as re-describing and synonymising some existing species. *'' D. aberrans'' Gasparo, 2010 – Italy *'' D. aciculata''
Simon Simon may refer to: People * Simon (given name), including a list of people and fictional characters with the given name Simon * Simon (surname), including a list of people with the surname Simon * Eugène Simon, French naturalist and the genus ...
, 1882
– Algeria *'' D. aculeata'' Kroneberg, 1875 – Central Asia, Iran? Introduced to Croatia *'' D. adriatica'' Kulczyński, 1897 – Austria, Balkans *'' D. affinis'' Ferrández, 1996 – Spain *'' D. afghana'' Denis, 1958 – Afghanistan *'' D. akpinarae'' Varol, 2016 – Turkey *'' D. alegranzaensis'' Wunderlich, 1992 – Canary Is. *'' D. alentejana'' Ferrández, 1996 – Portugal *'' D. ambulotenta'' Ribera, Ferrández & Blasco, 1986 – Canary Is. *'' D. anatoliae'' Deeleman-Reinhold, 1988 – Turkey *'' D. ancora'' Grasshoff, 1959 – Italy *'' D. andamanae'' Arnedo & Ribera, 1997 – Canary Is. *'' D. andreini'' Caporiacco, 1928 – Italy, Albania *'' D. aneris'' Macías-Hernández & Arnedo, 2010 – Selvagens Is. *'' D. anonyma'' Ferrández, 1984 – Spain *'' D. apenninica'' Alicata, 1964 – Italy **'' Dysdera a. aprutiana'' Alicata, 1964 – Italy *'' D. arabiafelix'' Gasparo & van Harten, 2006 – Yemen *'' D. arabica'' Deeleman-Reinhold, 1988 – Oman *'' D. arabisenen'' Arnedo & Ribera, 1997 – Canary Is. *'' D. argaeica'' Nosek, 1905 – Turkey *'' D. arganoi'' Gasparo, 2004 – Italy *'' D. armenica'' Charitonov, 1956 – Armenia, Georgia *'' D. arnedoi'' Lissner, 2017 – Spain (Majorca) *'' D. arnoldii'' Charitonov, 1956 – Central Asia *'' D. asiatica'' Nosek, 1905 – Turkey, Iran (?) *'' D. atlantea'' Denis, 1954 – Morocco *'' D. atlantica'' Simon, 1909 – Morocco *'' D. aurgitana'' Ferrández, 1996 – Spain *'' D. azerbajdzhanica'' Charitonov, 1956 – Caucasus (Russia, Georgia, Azerbaijan) *'' D. baetica'' Ferrández, 1984 – Spain *'' D. balearica'' Thorell, 1873 – Spain (Majorca) *'' D. bandamae''
Schmidt Schmidt may refer to: * Schmidt (surname), including list of people with the surname * Schmidt (singer) (born 1990), German pop and jazz singer * Schmidt (lunar crater), a small lunar impact crater * Schmidt (Martian crater), a List of craters on ...
, 1973
– Canary Is. *'' D. baratellii'' Pesarini, 2001 – Italy *'' D. beieri'' Deeleman-Reinhold, 1988 – Greece *'' D. bellimundi'' Deeleman-Reinhold, 1988 – Montenegro, Albania *'' D. bernardi'' Denis, 1966 – Libya *'' D. bicolor''
Taczanowski Taczanowski (Polish feminine: Taczanowska; plural: Taczanowscy) is the surname of a Polish szlachta (nobility) family from Poznań bearing the Jastrzębiec coat of arms and the motto: ''Plus penser que dire''. They took their name from th ...
, 1874
– French Guiana *'' D. bicornis'' Fage, 1931 – Spain *'' D. bidentata'' Dunin, 1990 – Azerbaijan *'' D. bogatschevi'' Dunin, 1990 – Georgia, Azerbaijan *'' D. borealicaucasica'' Dunin, 1991 – Russia (Caucasus) *'' D. bottazziae'' Caporiacco, 1951 – Italy, Croatia *'' D. breviseta'' Wunderlich, 1992 – Canary Is. *'' D. brevispina'' Wunderlich, 1992 – Canary Is. *'' D. brignoliana'' Gasparo, 2000 – Italy *'' D. brignolii'' Dunin, 1989 – Turkmenistan *'' D. caeca'' Ribera, 1993 – Morocco *'' D. calderensis'' Wunderlich, 1987 – Canary Is. *'' D. castillonensis'' Ferrández, 1996 – Spain *'' D. catalonica'' Řezáč, 2018 – Spain *'' D. cechica'' Řezáč, 2018 – Austria, Czechia, Slovakia, Hungary, Serbia? *'' D. centroitalica'' Gasparo, 1997 – Italy *'' D. cephalonica'' Deeleman-Reinhold, 1988 – Greece *'' D. charitonowi'' Mcheidze, 1979 – Georgia *'' D. chioensis'' Wunderlich, 1992 – Canary Is. *'' D. circularis'' Deeleman-Reinhold, 1988 – Greece *'' D. coiffaiti'' Denis, 1962 – Madeira *'' D. collucata'' Dunin, 1991 – Armenia *'' D. concinna''
L. Koch Ludwig Carl Christian Koch (8 November 1825 – 1 November 1908) was a German entomologist and arachnologist. He was born in Regensburg, Germany, and died in Nuremberg, Germany. He studied in Nuremberg, initially law, but then turned to medicin ...
, 1878
– Azerbaijan, Iran (?) *'' D. corallina'' Risso, 1826 – Spain, France *'' D. corfuensis'' Deeleman-Reinhold, 1988 – Albania, Greece (Corfu) *'' D. cornipes'' Karsch, 1881 – Libya *'' D. cribellata'' Simon, 1883 – Canary Is. *'' D. cribrata'' Simon, 1882 – France, Italy, Andorra *'' D. cristata'' Deeleman-Reinhold, 1988 – Syria, Lebanon *'' D. crocata''
C. L. Koch Carl Ludwig Koch (21 September 1778 – 23 August 1857) was a German entomologist and arachnologist. He was responsible for classifying a great number of spiders, including the Brazilian whiteknee tarantula and common house spider. He was born ...
, 1838
– Europe, Caucasus, Iraq, Central Asia. Introduced to North America, Chile, Brazil, Australia, New Zealand, Hawaii **'' Dysdera c. mutica'' Simon, 1911 – Algeria **'' Dysdera c. parvula'' Simon, 1911 – Algeria *'' D. crocolita'' Simon, 1911 – Algeria *'' D. curviseta'' Wunderlich, 1987 – Canary Is. *'' D. cylindrica''
O. Pickard-Cambridge Octavius Pickard-Cambridge FRS (3 November 1828 – 9 March 1917) was an English clergyman and zoologist. He was a keen arachnologist who described and named more than 900 species of spider. Life and work Pickard-Cambridge was born in Blox ...
, 1885
– Pakistan *'' D. daghestanica'' Dunin, 1991 – Russia (Caucasus) *'' D. dentichelis'' Simon, 1882 – Lebanon *'' D. deserticola'' Simon, 1911 – Algeria *'' D. dissimilis'' sp. nov. *'' D. diversa'' Blackwall, 1862 – Madeira *'' D. dolanskyi'' Řezáč, 2018 – Spain *'' D. drescoi'' Ribera, 1983 – Morocco *'' D. dubrovninnii'' Deeleman-Reinhold, 1988 – SE Europe (Balkans), Romania, Slovakia *'' D. dunini'' Deeleman-Reinhold, 1988 – Greece, Turkey, Ukraine, Caucasus (Russia, Georgia, Azerbaijan) *'' D. dysderoides'' (Caporiacco, 1947) – Ethiopia *'' D. edumifera'' Ferrández, 1983 – Spain *'' D. enghoffi'' Arnedo, Oromí & Ribera, 1997 – Canary Is. *'' D. enguriensis'' Deeleman-Reinhold, 1988 – Turkey *'' D. erythrina'' (Walckenaer, 1802) ( type) – Southwestern and Western to Central Europe *'' D. espanoli'' Ribera & Ferrández, 1986 – Spain *'' D. esquiveli'' Ribera & Blasco, 1986 – Canary Is. *'' D. exigua'' sp. nov. *'' D. fabrorum'' Řezáč, 2018 – Spain *'' D. falciformis'' Barrientos & Ferrández, 1982 – Spain *'' D. fedtschenkoi'' Dunin, 1992 – Tajikistan *'' D. ferghanica'' Dunin, 1985 – Kyrgyzstan *'' D. fervida'' Simon, 1882 – France (Corsica), Spain (Balearic Is.)? *'' D. festai'' Caporiacco, 1929 – Greece (Rhodes) *'' D. flagellata'' Grasshoff, 1959 – Italy *'' D. flagellifera'' Caporiacco, 1947 – Italy **'' Dysdera f. aeoliensis'' Alicata, 1973 – Italy *'' D. flavitarsis'' Simon, 1882 – Spain *'' D. fragaria'' Deeleman-Reinhold, 1988 – Greece (Rhodes) *'' D. furcata'' Varol & Danışman, 2018 – Turkey *'' D. fuscipes'' Simon, 1882 – Portugal, Spain, France *'' D. fustigans'' Alicata, 1966 – Italy *'' D. galinae'' Dimitrov, 2018 – Turkey *'' D. gamarrae'' Ferrández, 1984 – Spain *'' D. garrafensis'' Řezáč, 2018 – Spain *'' D. gemina'' Deeleman-Reinhold, 1988 – Israel *'' D. ghilarovi'' Dunin, 1987 – Azerbaijan *'' D. gibbifera'' Wunderlich, 1992 – Canary Is. *'' D. gigas'' Roewer, 1928 – Greece (Crete) *'' D. gmelini'' Dunin, 1991 – Georgia *'' D. gollumi'' Ribera & Arnedo, 1994 – Canary Is. *'' D. gomerensis''
Strand Strand may refer to: Topography *The flat area of land bordering a body of water, a: ** Beach ** Shoreline *Strand swamp, a type of swamp habitat in Florida Places Africa *Strand, Western Cape, a seaside town in South Africa * Strand Street, ...
, 1911
– Canary Is. *'' D. graia'' Řezáč, 2018 – France *'' D. granulata'' Kulczyński, 1897 – Italy, Balkans, Albania *'' D. gruberi'' Deeleman-Reinhold, 1988 – Turkey *'' D. guayota'' Arnedo & Ribera, 1999 – Canary Is. *'' D. halkidikii'' Deeleman-Reinhold, 1988 – Macedonia, Greece *'' D. hamifera'' Simon, 1911 – Algeria **'' Dysdera h. macellina'' Simon, 1911 – Algeria *'' D. hattusas'' Deeleman-Reinhold, 1988 – Turkey *'' D. helenae'' Ferrández, 1996 – Spain *'' D. hernandezi'' Arnedo & Ribera, 1999 – Canary Is. *'' D. hiemalis'' Deeleman-Reinhold, 1988 – Greece (Crete) *'' D. hirguan'' Arnedo, Oromí & Ribera, 1997 – Canary Is. *'' D. hirsti'' Denis, 1945 – Algeria *'' D. hungarica'' Kulczyński, 1897 – Central Europe to Azerbaijan **'' Dysdera h. atra'' Mcheidze, 1979 – Georgia, Azerbaijan **'' Dysdera h. subalpina'' Dunin, 1992 – Russia (Caucasus) *'' D. iguanensis'' Wunderlich, 1987 – Canary Is. *'' D. imeretiensis'' Mcheidze, 1979 – Georgia *'' D. incertissima'' Denis, 1961 – Morocco *'' D. incognita'' Dunin, 1991 – Russia (Europe, Caucasus) *'' D. inermis'' Ferrández, 1984 – Spain *'' D. inopinata'' Dunin, 1991 – Georgia *'' D. insulana'' Simon, 1883 – Canary Is. *'' D. isambertoi'' sp. nov. *'' D. jana'' Gasparo & Arnedo, 2009 – Italy (Sardinia) *'' D. karabachica'' Dunin, 1990 – Azerbaijan *'' D. kati'' Komnenov & Chatzaki, 2016 – Greece *'' D. kollari'' Doblika, 1853 – Italy, Balkans, Greece, Turkey *'' D. krisis'' Komnenov & Chatzaki, 2016 – Greece, Turkey *'' D. kronebergi'' Dunin, 1992 – Tajikistan *'' D. kropfi'' Řezáč, 2018 – Switzerland *'' D. kugitangica'' Dunin, 1992 – Turkmenistan *'' D. kulczynskii'' Simon, 1914 – France, Italy *'' D. kusnetsovi'' Dunin, 1989 – Turkmenistan *'' D. labradaensis'' Wunderlich, 1992 – Canary Is. *'' D. lagrecai'' Alicata, 1964 – Italy *'' D. lancerotensis'' Simon, 1907 – Canary Is. *'' D. lantosquensis'' Simon, 1882 – France, Italy *'' D. lata'' Reuss, 1834 – Mediterranean to Georgia *'' D. laterispina'' Pesarini, 2001 – Greece *'' D. leprieuri'' Simon, 1882 – Algeria *'' D. levipes'' Wunderlich, 1987 – Canary Is. *'' D. ligustica'' Gasparo, 1997 – Italy *'' D. limitanea'' Dunin, 1985 – Turkmenistan *'' D. limnos'' Deeleman-Reinhold, 1988 – Greece *'' D. liostetha'' Simon, 1907 – Canary Is. *'' D. littoralis'' Denis, 1962 – Morocco *'' D. longa'' Wunderlich, 1992 – Canary Is. *'' D. longibulbis'' Denis, 1962 – Madeira *'' D. longimandibularis'' Nosek, 1905 – Turkey, Cyprus *'' D. longirostris'' Doblika, 1853 – Central to south-eastern and eastern Europe, Turkey, Caucasus *'' D. lubrica'' Simon, 1907 – Egypt *'' D. lucidipes'' Simon, 1882 – Algeria **'' Dysdera l. melillensis'' Simon, 1911 – Morocco *'' D. lusitanica'' Kulczyński, 1915 – Portugal, Spain *'' D. machadoi'' Ferrández, 1996 – Portugal, Spain *'' D. macra'' Simon, 1883 – Canary Is. *'' D. madai'' Arnedo, 2007 – Canary Is. *'' D. mahan'' Macías-Hernández & Arnedo, 2010 – Canary Is. *'' D. maronita'' Gasparo, 2003 – Lebanon *'' D. martensi'' Dunin, 1991 – Caucasus (Russia, Georgia) *'' D. mauritanica'' Simon, 1909 – Morocco **'' Dysdera m. aurantiaca'' Simon, 1909 – Morocco *'' D. maurusia'' Thorell, 1873 – Algeria, Hungary?, Slovakia?, USA? *'' D. mazini'' Dunin, 1991 – Armenia, Azerbaijan *'' D. meschetiensis'' Mcheidze, 1979 – Georgia *'' D. microdonta'' Gasparo, 2014 – Italy, Austria, Slovenia, Serbia *'' D. minairo'' Řezáč, 2018 – Spain *'' D. minuta'' Deeleman-Reinhold, 1988 – Greece (Rhodes) *'' D. minutissima'' Wunderlich, 1992 – Canary Is. *'' D. mixta'' Deeleman-Reinhold, 1988 – Turkey *'' D. montanetensis'' Wunderlich, 1992 – Canary Is. *'' D. monterossoi'' Alicata, 1964 – Italy *'' D. moravica'' Řezáč, 2014 – Germany to Romania *'' D. mucronata'' Simon, 1911 – Morocco, Spain *'' D. murphyorum'' Deeleman-Reinhold, 1988 – Albania, Greece (Corfu) *'' D. nakhchivanica'' Beydizade, Shafaie & Guseinov, 2018 – Azerbaijan *'' D. nenilini'' Dunin, 1989 – Turkmenistan *'' D. neocretica'' Deeleman-Reinhold, 1988 – Greece (Crete), Turkey *'' D. nesiotes'' Simon, 1907 – Selvagens Is., Canary Is. *'' D. nicaeensis'' Thorell, 1873 – France, Italy *'' D. ninnii''
Canestrini Giovanni Canestrini (26 December 1835 – 14 February 1900) was an Italian naturalist and biologist and translator who was a native of Revò. Career He initially studied in Gorizia and Meran, then furthered his education in natural sciences at ...
, 1868
– Switzerland, Italy, Slovenia, Croatia *'' D. nomada'' Simon, 1911 – Tunisia *'' D. nubila'' Simon, 1882 – France (Corsica), Italy *'' D. orahan'' Arnedo, Oromí & Ribera, 1997 – Canary Is. *'' D. ortunoi'' Ferrández, 1996 – Spain *'' D. osellai'' Alicata, 1973 – Italy *'' D. paganettii'' Deeleman-Reinhold, 1988 – Italy *'' D. pamirica'' Dunin, 1992 – Tajikistan *'' D. pandazisi'' Hadjissarantos, 1940 – Greece *'' D. paucispinosa'' Wunderlich, 1992 – Canary Is. *'' D. pavani'' Caporiacco, 1941 – Italy *'' D. pectinata'' Deeleman-Reinhold, 1988 – Bulgaria, Macedonia, Greece *'' D. pharaonis'' Simon, 1907 – Egypt *'' D. pococki'' Dunin, 1985 – Turkmenistan *'' D. pominii'' Caporiacco, 1947 – Italy *'' D. portisancti'' Wunderlich, 1995 – Madeira *'' D. portsensis'' Řezáč, 2018 – Spain *'' D. pradesensis'' Řezáč, 2018 – Spain *'' D. praepostera'' Denis, 1961 – Morocco *'' D. precaria'' sp. nov. *'' D. presai'' Ferrández, 1984 – Spain *'' D. pretneri'' Deeleman-Reinhold, 1988 – Croatia, Montenegro, Greece *'' D. pristiphora'' Pesarini, 2001 – Italy *'' D. punctata'' C. L. Koch, 1838 – Southern Europe, Slovakia?, Georgia? *'' D. punctocretica'' Deeleman-Reinhold, 1988 – Greece (Corfu) *'' D. pyrenaica'' Řezáč, 2018 – Spain *'' D. quindecima'' Řezáč, 2018 – Spain *'' D. raddei'' Dunin, 1990 – Azerbaijan *'' D. ramblae'' Arnedo, Oromí & Ribera, 1997 – Canary Is. *'' D. ratonensis'' Wunderlich, 1992 – Canary Is. *'' D. ravida'' Simon, 1909 – Morocco *'' D. recondita'' sp. nov. *'' D. richteri'' Charitonov, 1956 – Azerbaijan, Armenia, Georgia *'' D. roemeri'' Strand, 1906 – Ethiopia *'' D. romana'' Gasparo & Di Franco, 2008 – Italy *'' D. romantica'' Deeleman-Reinhold, 1988 – Greece *'' D. rostrata'' Denis, 1961 – Morocco *'' D. rubus'' Deeleman-Reinhold, 1988 – Turkey, Greece *'' D. rudis'' Simon, 1882 – France *'' D. rugichelis'' Simon, 1907 – Canary Is. *'' D. rullii'' Pesarini, 2001 – Italy *'' D. sanborondon'' Arnedo, Oromí & Ribera, 2000 – Canary Is. *'' D. sandrae'' sp. nov. *'' D. satunini'' Dunin, 1990 – Azerbaijan *'' D. scabricula'' Simon, 1882 – France, Spain *'' D. sciakyi'' Pesarini, 2001 – Greece *'' D. seclusa'' Denis, 1961 – Morocco *'' D. sefrensis'' Simon, 1911 – Morocco *'' D. septima'' Řezáč, 2018 – Spain *'' D. shardana'' Opatova & Arnedo, 2009 – Italy (Sardinia) *'' D. sibyllina'' Arnedo, 2007 – Canary Is. *'' D. sibyllinica'' Kritscher, 1956 – Italy *'' D. silana'' Alicata, 1965 – Italy *'' D. silvatica'' Schmidt, 1981 – Canary Is. *'' D. simbeque'' Macías-Hernández & Arnedo, 2010 – Canary Is. *'' D. simoni'' Deeleman-Reinhold, 1988 – Syria, Israel, Lebanon *'' D. snassenica'' Simon, 1911 – Morocco, Algeria **'' Dysdera s. collina'' Simon, 1911 – Morocco *'' D. soleata'' Karsch, 1881 – Libya *'' D. solers'' Walckenaer, 1837 – Colombia *'' D. spasskyi'' Charitonov, 1956 – Georgia *'' D. spinicrus'' Simon, 1882 – Balkans, Greece, Syria *'' D. spinidorsa'' Wunderlich, 1992 – Canary Is. *'' D. stahlavskyi'' Řezáč, 2018 – France *'' D. subcylindrica'' Charitonov, 1956 – Central Asia *'' D. subnubila'' Simon, 1907 – Italy, Tunisia, Egypt *'' D. subsquarrosa'' Simon, 1914 – France, Italy *'' D. sultani'' Deeleman-Reinhold, 1988 – Greece, Turkey *'' D. sutoria'' Denis, 1945 – Morocco *'' D. tartarica'' Kroneberg, 1875 – Central Asia *'' D. tbilisiensis'' Mcheidze, 1979 – Georgia *'' D. tenuistyla'' Denis, 1961 – Morocco *'' D. teixeirai'' sp. nov. *'' D. tezcani'' Varol & Akpınar, 2016 – Turkey *'' D. tilosensis'' Wunderlich, 1992 – Canary Is. *'' D. titanica'' sp. nov. *'' D. topcui'' Gasparo, 2008 – Turkey *'' D. tredecima'' Řezáč, 2018 – Spain *'' D. turcica'' Varol, 2016 – Turkey *'' D. tystshenkoi'' Dunin, 1989 – Turkmenistan *'' D. ukrainensis'' Charitonov, 1956 – Ukraine, Russia (Europe), Georgia *'' D. undecima'' Řezáč, 2018 – Spain *'' D. unguimmanis'' Ribera, Ferrández & Blasco, 1986 – Canary Is. *'' D. valentina'' Ribera, 2004 – Spain *'' D. vandeli'' Denis, 1962 – Madeira *'' D. veigai'' Ferrández, 1984 – Spain *'' D. ventricosa'' Grasshoff, 1959 – Italy *'' D. vermicularis''
Berland Berland is a surname. Notable people with the surname include: * Bjørn Berland (born 1977), Norwegian footballer *Eliezer Berland (born 1937), Israeli Breslov rabbi * François Berland (born 1958), French actor *Gary Berland (1950–1988), Americ ...
, 1936
– Cape Verde Is. *'' D. verneaui'' Simon, 1883 – Canary Is. *'' D. vesiculifera'' Simon, 1882 – Algeria *'' D. vignai'' Gasparo, 2003 – Lebanon *'' D. vivesi'' Ribera & Ferrández, 1986 – Spain *'' D. volcania'' Ribera, Ferrández & Blasco, 1986 – Canary Is. *'' D. werneri'' Deeleman-Reinhold, 1988 – Greece *'' D. westringi'' O. Pickard-Cambridge, 1872 – Eastern Mediterranean, Iraq *'' D. yguanirae'' Arnedo & Ribera, 1997 – Canary Is. *'' D. yozgat'' Deeleman-Reinhold, 1988 – Turkey *'' D. zarudnyi'' Charitonov, 1956 – Central Asia, Afghanistan


References


Further reading

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External links


Dysdera
at
BugGuide BugGuide (or BugGuide.net) is a website and online community of naturalists, both amateur and professional, who share observations of arthropods such as insects, spiders, and other related creatures. The website consists of informational guide p ...
{{Taxonbar, from=Q2083007 Araneomorphae genera Dysderidae Invertebrates of North Africa Spiders of Africa Spiders of Asia Taxa named by Pierre André Latreille