:''The

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...

genus ''Dryopteris'' is now considered a

junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...

of ''

Oreta ''Oreta'' is a genus of moths belonging to the subfamily Drepaninae. The genus was erected by Francis Walker (entomologist), Francis Walker in 1855. Species *The ''rosea'' species group **''Oreta ancora'' Chu and Wang, 1987 **''Oreta andrema'' Wi ...

. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''

Dryopteris filix-mas ''Dryopteris filix-mas'', the male fern, is a common fern of the temperate Northern Hemisphere, native to much of Europe, Asia, and North America. It favours damp shaded areas in the understory of woodlands, but also shady places on hedge-banks, ...

''), or buckler ferns, is a

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...

genus in the family Dryopteridaceae, subfamily Dryopteridoideae, according to the

Pteridophyte Phylogeny Group The Pteridophyte Phylogeny Group, or PPG, is an informal international group of systematic botanists who collaborate to establish a consensus on the classification of pteridophytes (lycophytes and ferns) that reflects knowledge about plant relation ...

classification of 2016 (PPG I). There are about 300-400 species in the genus. The species are distributed in Asia, the Americas, Europe, Africa, and the Pacific islands, with the highest diversity in eastern Asia. It is placed in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Dryopteridaceae, subfamily Dryopteridoideae, according to the

classification of 2016 (PPG I). Many of the species have stout, slowly creeping rootstocks that form a crown, with a vase-like ring of fronds. The sori are round, with a peltate

indusium A sorus (pl. sori) is a cluster of sporangia (structures producing and containing spores) in ferns and fungi. A coenosorus (plural coenosori) is a compound sorus composed of multiple, fused sori. Etymology This New Latin word is from Ancient ...

. The stipes have prominent scales.

Hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...

and polyploidy are well-known phenomena in this group, with many species formed via these processes. The North American ''Dryopteris'' hybrid complex is a well-known example of

speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution within ...

via allopolyploid hybridization.

Selected species

The genus has a large number of species. The PPG I classification suggested there were about 400 species; , the ''Checklist of Ferns and Lycophytes of the World'' listed 328 species and 83 hybrids. Some genera sunk into ''Dryopteris'', such as ''Dryopsis'', ''Stenolepia'' and ''Nothoperanema'', are distinguished by other sources. *''

Dryopteris abbreviata :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *'' Dryopteris aemula'' – hay-scented buckler fern *'' Dryopteris affinis'' – scaly male fern *'' Dryopteris alpestris'' *'' Dryopteris amurensis'' *'' Dryopteris apicisora'' *'' Dryopteris arguta'' – coastal wood fern *'' Dryopteris azorica'' *'' Dryopteris backeri'' *'' Dryopteris barbigera'' *'' Dryopteris bissetiana'' *'' Dryopteris blandfordii'' *'' Dryopteris bodinieri'' *''

Dryopteris borreri :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *''

Dryopteris cambrensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' – narrow male fern *'' Dryopteris campyloptera'' – mountain wood fern *''

Dryopteris canaliculata :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *'' Dryopteris carthusiana'' – narrow buckler fern *'' Dryopteris caucasica'' *''

Dryopteris caudifrons :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris celsa'' – log fern *'' Dryopteris championii'' *'' Dryopteris changii'' *''

Dryopteris chapaensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris chinensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris chrysocoma :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris cinnamomea :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' – cinnamon wood fern *''

Dryopteris clintoniana ''Dryopteris clintoniana'', commonly known as Clinton's wood fern, is a fern of hybrid origin native to the northern hemisphere. It is a fertile hexaploid, arising as a species by doubling of its chromosome number from a hybrid between ''Dryopte ...

'' – Clinton's wood fern *''

Dryopteris confertipinna :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris cordipinna :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris corleyi :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris costalisora :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris crassirhizoma ''Dryopteris crassirhizoma'' is a fern species in the wood fern family Dryopteridaceae. This semi-evergreen fern grows to tall and broad, with narrowly-divided fronds growing in a vase-like shape from a central crown, which is brown in colour. ...

'' *''

Dryopteris crinalis ''Dryopteris crinalis'' is a species of fern known as the serpent woodfern. It is endemic to Hawaii, where it is known from the main islands.Dryopteris crispifolia'' *'' Dryopteris cristata'' – crested buckler fern *''

Dryopteris cyatheoides :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *'' Dryopteris cycadina'' *'' Dryopteris cyclopeltiformis'' *'' Dryopteris decipiens'' *''

Dryopteris dehuaensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris dickinsii :''The moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of mo ...

'' *'' Dryopteris dilatata'' – broad buckler fern *''

Dryopteris discrita :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to '' Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *''

Dryopteris enneaphylla :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris erythrosa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris erythrosora ''Dryopteris erythrosora'', the autumn fern or Japanese shield fern, is a species of fern in the family (biology), family Dryopteridaceae, native plant, native to east Asia from China and Japan south to the Philippines, growing in light woodland ...

'' – autumn fern *''

Dryopteris expansa ''Dryopteris expansa'', the alpine buckler fern, northern buckler-fern or spreading wood fern, is a species of fern native to cool temperate and subarctic regions of the Northern Hemisphere, south at high altitudes in mountains to Spain and Greec ...

'' – northern buckler fern *''

Dryopteris fibrilosa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris fibrilosissima :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to '' Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *''

'' – male fern *''

Dryopteris floridana :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *'' Dryopteris formosana'' *''

Dryopteris fragrans ''Dryopteris fragrans'', commonly known as the fragrant woodfern, a circumboreal fern, is the smallest of the ''Dryopteris'' species. It can resemble ''Woodsia ilvensis'' in the wild, with which it shares the same habitat of rocky areas, shady cl ...

'' – fragrant buckler fern, fragrant wood fern *''

Dryopteris fructuosa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris fuscipes :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris gamblei'' *''

Dryopteris goerigiana :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris goldieana'' – Goldie's wood fern *''

Dryopteris gongboensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *'' Dryopteris guanchica'' *''

Dryopteris gushanica :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris gushiangensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris gymnophylla :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris gymnosora :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris hasseltii ''Dryopteris hasseltii'' is a species of fern in the family Dryopteridaceae. It occurs from India, through southeast Asia, as far as north-east Queensland in Australia. Its natural habitat is subtropical or tropical moist lowland forests. It is ...

'' *''

Dryopteris hendersoni :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris hirtipes ''Dryopteris hirtipes'' is fern species native to South China and Papua New Guinea Papua New Guinea (abbreviated PNG; , ; tpi, Papua Niugini; ho, Papua Niu Gini), officially the Independent State of Papua New Guinea ( tpi, Independen ...

'' *'' Dryopteris huanganshanensis'' *''

Dryopteris hupehensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris hwangii :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris hypophlebia :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris immixta :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *'' Dryopteris inaequalis'' *''

Dryopteris incisolobata :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris integriloba :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris integripinnula'' *'' Dryopteris intermedia'' – intermediate wood fern *''

Dryopteris junlianensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *''

Dryopteris juxtaposita :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris kinkiensis'' *''

Dryopteris labordei :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris lacera :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to '' Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *''

Dryopteris lancipinnula :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris lepidopoda :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris lepidorachis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris liyangensis'' *'' Dryopteris ludoviciana'' – southern wood fern *''

Dryopteris macropholis ''Dryopteris macropholis'' is a species of fern. It is distributed on the Marquesas Islands. Description ''Dryopteris macropholis'' is a terrestrial fern; rhizomes suberect, 20–25 cm long, 5–7 cm in diameter (to 15 cm includi ...

'' *'' Dryopteris manshurica'' *''

Dryopteris marginalis ''Dryopteris marginalis'', vernacularly known as the marginal shield fern or marginal wood fern, is a perennial species of fern found in damp shady areas throughout eastern North America, from Texas to Minnesota and Newfoundland. It favors modera ...

'' – marginal wood fern *''

Dryopteris matsumurae :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris minjiangensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to '' Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *''

Dryopteris monticola :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris montigena :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris nanpingensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *'' Dryopteris neolepidopoda'' *'' Dryopteris nigropaleacea'' *''

Dryopteris nigrosquamosa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris nyalamensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris nyingchiensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris odontoloma :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris oreades'' – mountain male fern *''

Dryopteris pacifica :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris pallida ''Dryopteris pallida'' is a species of fern. Description ''Dryopteris pallida'' is a winter-green ''Dryopteris'' fern of small to moderate size with very pale broad chaffy scales on the long somewhat pale stem and leaves that are often rather a ...

'' *''

Dryopteris paludicda :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *''

Dryopteris panda :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris parasparsa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris pedata :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris peninsulae :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris podophylla :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris polita :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *''

Dryopteris prosa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *'' Dryopteris pseudo-sikkimensis'' *''

Dryopteris pseudoatrata :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris pseudodontoloma :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris pseudofibrillosa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris pseudomarginata :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris pseudouniformis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris pulcherrima :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris qandoensis'' *''

Dryopteris quatanensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris reflexosquamata :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris remota :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' – scaly buckler fern *''

Dryopteris rigida :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris rosthornii :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris sacrosancta :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris saxifraga :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris scottii :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris semipinnata :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris sericea :''The moth genus ''Dryopteris'' is now considered a junior synonym of '' Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family D ...

'' *''

Dryopteris serrato-dentata :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris shensicola :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris shibipedis'' *''

Dryopteris sichotensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris sieboldii ''Dryopteris sieboldii'' is a species of fern in the family Dryopteridaceae, native to Japan and south-central and southeast China. It has gained the Royal Horticultural Society's Award of Garden Merit The Award of Garden Merit (AGM) is a lon ...

'' *'' Dryopteris sikkimensis'' *''

Dryopteris silaensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris sino-sparsa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris sinoerythrosora :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris sinofibrillosa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris sordidipes :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris sparsa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris squamifera'' *''

Dryopteris squamiseta :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris stenolepis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris subatrata :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris subbarbigera :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris subexaltata'' *''

Dryopteris sublacera :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris sublaeta :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris submarginata'' *'' Dryopteris submontana'' – rigid buckler fern *'' Dryopteris subtriangularis'' *''

Dryopteris tarningensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris tenuicula'' *''

Dryopteris tenuissima :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris thibetica'' *''

Dryopteris tieluensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris tokyoensis ''Dryopteris tokyoensis'' (Tokyo wood fern, 东京鳞毛蕨) is an erect, vase-shaped, deciduous fern native to Japan, as well as Fujian, Hubei, Hunan, Jiangxi, and Zhejiang provinces in China. These ferns grow to 90–110 cm in height, with ...

'' – Tokyo wood fern *''

Dryopteris tsangpoensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris tyrrhena :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris uniformis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris varia ''Dryopteris varia'' is a species of fern in the family Dryopteridaceae. The species is native to China, Japan, Korea, Myanmar, Philippines, Taiwan, Vietnam, Sulawesi, and the Ryukyu Islands. References

Dryopteris, varia Ferns of Asia ...

'' *''

Dryopteris venosa :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris villarii'' *''

Dryopteris wallichiana ''Dryopteris wallichiana'', the alpine wood fern, is a robust species of deciduous or semi-evergreen fern in the family (biology), family Dryopteridaceae, native plant, native to the Himalayas, Hawaii, Mexico and Jamaica. It grows to tall, occas ...

'' – alpine wood fern *''

Dryopteris wenchuanensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *''

Dryopteris wuyishanensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dr ...

'' *''

Dryopteris yigongensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' *'' Dryopteris yungtzeensis'' *''

Dryopteris zayuensis :''The moth genus ''Dryopteris'' is now considered a junior synonym of ''Oreta. ''Dryopteris'' , commonly called the wood ferns, male ferns (referring in particular to ''Dryopteris filix-mas''), or buckler ferns, is a fern genus in the family Dry ...

'' File:Dryopteris goldiana.jpg, '' Dryopteris goldieana'', Goldie's fern File:Dryopteris sieboldii - Atlanta Botanical Garden.JPG, ''

Ecology

''Dryopteris'' species are used as food plants by the

larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...

e of some

Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...

species including '' Batrachedra sophroniella'' (which feeds exclusively on ''D. cyatheoides'') and '' Sthenopiseauratus''.

Cultivation and uses

Many ''Dryopteris'' species are widely used as garden

ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...

s, especially ''D. affinis'', ''D. erythrosora'', and ''D. filix-mas'', with numerous

cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...

s. ''Dryopteris filix-mas'' was throughout much of recent human history widely used as a

vermifuge Anthelmintics or antihelminthics are a group of antiparasitic drugs that expel parasitic worms (helminths) and other internal parasites from the body by either stunning or killing them and without causing significant damage to the host. They may a ...

, and was the only fern listed in the U.S.

Pharmacopoeia A pharmacopoeia, pharmacopeia, or pharmacopoea (from the obsolete typography ''pharmacopœia'', meaning "drug-making"), in its modern technical sense, is a book containing directions for the identification of compound medicines, and published by ...

. Traditional use in

Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion#Europe, subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, ...

against red mite ( Dermanyssus gallinae) infestation is to place fronds in nesting boxes under nesting material and under floor covering material.

References

External links

Flora Europaea: ''Dryopteris''

Flora of North America: ''Dryopteris''

Flora of China: ''Dryopteris'' species list
{{Taxonbar, from=Q1335106 Fern genera

Selected species

Ecology

Cultivation and uses

See also

References

External links