Dromomeron Romeri
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''Dromomeron'' (meaning "running
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates wit ...
") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of lagerpetid
avemetatarsalian Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Dinos ...
which lived around 220 to 211.9 ± 0.7 million years ago. The genus contains species known from
Late Triassic The Late Triassic is the third and final epoch of the Triassic Period in the geologic time scale, spanning the time between Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch and followed by the Early Jurassic Epoch. ...
-age rocks of the Southwestern United States and northwestern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. It is described as most closely related to the earlier ''
Lagerpeton ''Lagerpeton'' is a genus of lagerpetid avemetatarsalian, comprising a single species, ''L. chanarensis''. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, ''Lagerpeton'' anatomy is somewhat incompletely kn ...
'' of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, but was found among remains of true dinosaurs like ''
Chindesaurus ''Chindesaurus'' ( ) is an extinct genus of basal saurischian dinosaur from the Late Triassic (213-210 million years ago) of the southwestern United States. It is known from a single species, ''C. bryansmalli'', based on a partial skeleton recovere ...
'', indicating that the first dinosaurs did not immediately replace related groups. Based on the study of the overlapping material of ''Dromomeron'' and ''
Tawa hallae ''Tawa'' (named after the Hopi word for the Puebloan sun god) is a genus of possible basal theropod dinosaurs from the Late Triassic period. The fossil remains of ''Tawa hallae'', the type and only species were found in the Hayden Quarry of ...
'', Christopher Bennett proposed that the two taxa were conspecific, forming a single growth series of ''Dromomeron''. However, noting prominent differences between their femurs which cannot be attributed to variation with age, Rodrigo Muller rejected this proposal in 2017. He further noted that, while ''D. romeri'' is known from juveniles only, it shares many traits in common with ''D. gigas'', which is known from mature specimens.


Description

It is known from partial remains, largely from the hindlimbs, which indicate an animal with an overall length of .


Classification

The bones of ''Dromomeron'' are most similar to those of the older pterosauromorph ''Lagerpeton'', and the two animals have been classified together in a clade
Lagerpetonidae Lagerpetidae (; originally Lagerpetonidae) is a family of basal avemetatarsalians. Though traditionally considered the earliest-diverging dinosauromorphs (reptiles closer to dinosaurs than to pterosaurs), fossils described in 2020 suggest that ...
.Baron, M.G., Norman, D.B., and Barrett, P.M. (2017). A new hypothesis of dinosaur relationships and early dinosaur evolution. ''Nature'', 543: 501–506.
Cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
simplified after Kammerer, Nesbitt & Shubin (2012):


Discovery and species


''Dromomeron romeri''

The species name ''romeri'' honors influential 20th-century
vertebrate paleontologist Vertebrate paleontology is the subfield of paleontology that seeks to discover, through the study of fossilized remains, the behavior, reproduction and appearance of extinct animals with vertebrae or a notochord. It also tries to connect, by u ...
Alfred Sherwood Romer Alfred Sherwood Romer (December 28, 1894 – November 5, 1973) was an American paleontologist and biologist and a specialist in vertebrate evolution. Biography Alfred Romer was born in White Plains, New York, the son of Harry Houston Romer an ...
. ''Dromomeron'' and
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specime ...
''D. romeri'' are based on GR 218, a complete left
thigh bone The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates wit ...
from the Hayden Quarry at
Ghost Ranch Ghost Ranch is a retreat and education center located close to the village of Abiquiú in Rio Arriba County in north central New Mexico, United States. It was the home and studio of Georgia O'Keeffe, as well as the subject of many of her painti ...
,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ke ...
. The rocks there are in the lower portion of the Petrified Forest Member of the Chinle Formation, and are Norian in age. Additional hindlimb bones, some probably from the same individual, are also known, and a partial skeleton has been recovered from Hayden Quarry, but has not yet been fully prepared. A few other specimens have been recovered from nearby localities, including the Snyder Quarry. Other specimens from the Chinle Formation of
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
and a roughly contemporaneous part of the
Dockum Group The Dockum is a Late Triassic (approximately late Carnian through Rhaetian, or 223–200 Ma) geologic group found primarily on the Llano Estacado of western Texas and eastern New Mexico with minor exposures in southwestern Kansas, eastern Colora ...
of
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
also have been assigned to this genus. These have been assigned to a second species ''D. gregorii'', named in 2009.


''Dromomeron gregorii''

''D. gregorii'', named for Joseph T. Gregory, is based on TMM 31100–1306, a right femur (thigh bone) from the Otis Chalk Quarry,
Colorado City Formation The Colorado City Formation is a geologic formation in Texas, United States. It preserves fossils dating back to the Triassic period. Biochronological significance The Otis Chalk localities that are situated in the Colorado City Formation form ...
, (Dockum Group), near
Otis Chalk Otis may refer to: Arts and entertainment Characters * Otis (Superman), in the films ''Superman'' and ''Superman II'' and related DC Comics media ** Otis Graves, in the TV series ''Supergirl'' * Otis (''The Walking Dead''), in the Image Comics ...
, Texas. Several other limb bones from the quarry, and a partial femur (thigh bone) from the ''
Placerias ''Placerias'' (meaning 'broad body') is an extinct genus of dicynodonts that lived during the Carnian to the Norian age of the Triassic Period (230–220 million years ago). ''Placerias'' belongs to a group of dicynodonts called Kannemeyeriifor ...
'' Quarry of eastern
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
have been assigned to this species. The rocks that ''D. gregorii'' is known from are older than those ''romeri'' has been found in. As with the Hayden Quarry, the Otis Chalk Quarry has at least one specimen of a herrerasaurid.


''Dromomeron gigas''

A third species, ''D. gigas'', was described by Martínez ''et al''. (2016) on the basis of fossils recovered from the Norian
Quebrada del Barro Formation The Quebrada del Barro Formation is a geological formation of the Marayes-El Carrizal Basin in San Juan Province, Argentina. This formation is the most fossiliferous portion of the Triassic Marayes Group, and is also the youngest unit of the gr ...
in northwestern Argentina.


Paleoecology

Also found at the Hayden Quarry are the remains of
phytosaur Phytosaurs (Φυτόσαυροι in greek) are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. Phytosauria and Phytosauridae are often considered to be equivalent g ...
s,
aetosaur Aetosaurs () are heavily armored reptiles belonging to the extinct order (biology), order Aetosauria (; from Ancient Greek, Greek, (aetos, "eagle") and (, "lizard")). They were medium- to large-sized Omnivore, omnivorous or Herbivore, herbivoro ...
s,
rauisuchia "Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilians ...
ns, and several types of dinosaurs and dinosaur relatives, including a '' Silesaurus''-like animal, the
herrerasaur Herrerasauridae is a family of carnivorous dinosaurs, possibly basal to either theropods or even all of saurischians, or even their own branching from dracohors, separate from dinosauria altogether. They are among the oldest known dinosaurs, ...
id ''
Chindesaurus ''Chindesaurus'' ( ) is an extinct genus of basal saurischian dinosaur from the Late Triassic (213-210 million years ago) of the southwestern United States. It is known from a single species, ''C. bryansmalli'', based on a partial skeleton recovere ...
'', and the basal theropod ''Tawa''. Finding the remains of four types of dinosaurs and dinosaur relatives (including ''Dromomeron'' itself) is noteworthy because it shows that dinosaurs did not immediately replace their dinosauromorph predecessors; that some of these groups, like the lagerpetonids, persisted (for longer than previously known) and diversified; and that dinosaurian replacement may have occurred at different times in different areas.


References


External links


Online supplementary material
for the Irmis ''et al.''. article (PDF).

for the Irmis ''et al.''. article. {{Taxonbar, from=Q2566615 Prehistoric avemetatarsalians Late Triassic archosaurs of North America Late Triassic reptiles of South America Fossil taxa described in 2007 Triassic Argentina Lagerpetidae