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''Dromomeron'' (meaning "running
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...
") is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
lagerpetid Lagerpetidae (; originally Lagerpetonidae) is a family of basal avemetatarsalians. Though traditionally considered the earliest-diverging dinosauromorphs (reptiles closer to dinosaurs than to pterosaurs), fossils described in 2020 suggest that ...
avemetatarsalian Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Dinos ...
which lived around 220 to 211.9 ± 0.7 million years ago. The genus contains species known from
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
-age rocks of the
Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, Ne ...
and northwestern
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
. It is described as most closely related to the earlier ''
Lagerpeton ''Lagerpeton'' is a genus of lagerpetid avemetatarsalian, comprising a single species, ''L. chanarensis''. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, ''Lagerpeton'' anatomy is somewhat incompletely kn ...
'' of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, but was found among remains of true dinosaurs like '' Chindesaurus'', indicating that the first dinosaurs did not immediately replace related groups. Based on the study of the overlapping material of ''Dromomeron'' and '' Tawa hallae'', Christopher Bennett proposed that the two taxa were conspecific, forming a single growth series of ''Dromomeron''. However, noting prominent differences between their femurs which cannot be attributed to variation with age, Rodrigo Muller rejected this proposal in 2017. He further noted that, while ''D. romeri'' is known from juveniles only, it shares many traits in common with ''D. gigas'', which is known from mature specimens.


Description

It is known from partial remains, largely from the hindlimbs, which indicate an animal with an overall length of .


Classification

The bones of ''Dromomeron'' are most similar to those of the older pterosauromorph ''Lagerpeton'', and the two animals have been classified together in a
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
Lagerpetonidae Lagerpetidae (; originally Lagerpetonidae) is a family of basal avemetatarsalians. Though traditionally considered the earliest-diverging dinosauromorphs (reptiles closer to dinosaurs than to pterosaurs), fossils described in 2020 suggest that l ...
.Baron, M.G., Norman, D.B., and Barrett, P.M. (2017). A new hypothesis of dinosaur relationships and early dinosaur evolution. ''Nature'', 543: 501–506. Cladogram simplified after Kammerer, Nesbitt & Shubin (2012):


Discovery and species


''Dromomeron romeri''

The species name ''romeri'' honors influential 20th-century
vertebrate paleontologist Vertebrate paleontology is the subfield of paleontology that seeks to discover, through the study of fossilized remains, the behavior, reproduction and appearance of extinct animals with vertebrae or a notochord. It also tries to connect, by u ...
Alfred Sherwood Romer. ''Dromomeron'' and
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
''D. romeri'' are based on GR 218, a complete left
thigh bone The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with t ...
from the
Hayden Quarry Hayden may refer to: Places Inhabited places in the United States *Hayden, Alabama *Hayden, Arizona *Hayden's Ferry, former name of Tempe, Arizona *Hayden, California, former name of Hayden Hill, California *Hayden, Colorado *Hayden, Idaho *Hayden ...
at Ghost Ranch,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
. The rocks there are in the lower portion of the Petrified Forest Member of the
Chinle Formation The Chinle Formation is an Upper Triassic continental geological formation of fluvial, lacustrine, and palustrine to eolian deposits spread across the U.S. states of Nevada, Utah, northern Arizona, western New Mexico, and western Colorado. In Ne ...
, and are
Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...
in age. Additional hindlimb bones, some probably from the same individual, are also known, and a partial skeleton has been recovered from Hayden Quarry, but has not yet been fully prepared. A few other specimens have been recovered from nearby localities, including the Snyder Quarry. Other specimens from the Chinle Formation of
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
and a roughly contemporaneous part of the Dockum Group of
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
also have been assigned to this genus. These have been assigned to a second species ''D. gregorii'', named in 2009.


''Dromomeron gregorii''

''D. gregorii'', named for
Joseph T. Gregory Dr. Joseph Tracy Gregory (July 28, 1914 – November 18, 2007) was an American paleontology, paleontologist and professor. Joseph Tracy Gregory was born in Eureka, California, the only child of Frank Gregory, a civil engineer, and Edith Tracy, ...
, is based on TMM 31100–1306, a right femur (thigh bone) from the Otis Chalk Quarry, Colorado City Formation, (Dockum Group), near Otis Chalk, Texas. Several other limb bones from the quarry, and a partial femur (thigh bone) from the '' Placerias'' Quarry of eastern
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
have been assigned to this species. The rocks that ''D. gregorii'' is known from are older than those ''romeri'' has been found in. As with the Hayden Quarry, the Otis Chalk Quarry has at least one specimen of a
herrerasaurid Herrerasauridae is a family of carnivorous dinosaurs, possibly basal to either theropods or even all of saurischians, or even their own branching from dracohors, separate from dinosauria altogether. They are among the oldest known dinosaurs, fi ...
.


''Dromomeron gigas''

A third species, ''D. gigas'', was described by Martínez ''et al''. (2016) on the basis of fossils recovered from the Norian
Quebrada del Barro Formation The Quebrada del Barro Formation is a geological formation of the Marayes-El Carrizal Basin in San Juan Province, Argentina. This formation is the most fossiliferous portion of the Triassic Marayes Group, and is also the youngest unit of the gro ...
in northwestern Argentina.


Paleoecology

Also found at the
Hayden Quarry Hayden may refer to: Places Inhabited places in the United States *Hayden, Alabama *Hayden, Arizona *Hayden's Ferry, former name of Tempe, Arizona *Hayden, California, former name of Hayden Hill, California *Hayden, Colorado *Hayden, Idaho *Hayden ...
are the remains of phytosaurs, aetosaurs, rauisuchians, and several types of dinosaurs and dinosaur relatives, including a ''
Silesaurus ''Silesaurus'' is a genus of silesaurid dinosauriform from the Late Triassic, of what is now Poland. Discovery Fossilized remains of ''Silesaurus'' have been found in the Keuper Claystone in Krasiejów near Opole, Silesia, Poland, which is al ...
''-like animal, the herrerasaurid '' Chindesaurus'', and the basal theropod ''Tawa''. Finding the remains of four types of dinosaurs and dinosaur relatives (including ''Dromomeron'' itself) is noteworthy because it shows that dinosaurs did not immediately replace their dinosauromorph predecessors; that some of these groups, like the lagerpetonids, persisted (for longer than previously known) and diversified; and that dinosaurian replacement may have occurred at different times in different areas.


References


External links


Online supplementary material
for the Irmis ''et al.''. article (PDF).

for the Irmis ''et al.''. article. {{Taxonbar, from=Q2566615 Prehistoric avemetatarsalians Late Triassic archosaurs of North America Late Triassic reptiles of South America Fossil taxa described in 2007 Triassic Argentina Lagerpetidae