Dromatheriidae is an

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

prozostrodontia Prozostrodontia is a clade of cynodonts including mammaliaforms and their closest relatives such as Tritheledontidae and Tritylodontidae. It was erected as a node-based taxon by Liu and Olsen (2010) and defined as the least inclusive clade cont ...

cynodont The cynodonts () (clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variety ...

s, closely related to

mammals Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...

. Members of the family are known from the

Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...

(

Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...

Rhaetian The Rhaetian is the latest age of the Triassic Period (in geochronology) or the uppermost stage of the Triassic System (in chronostratigraphy). It was preceded by the Norian and succeeded by the Hettangian (the lowermost stage or earliest age ...

) of

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

and

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

. Apart from a few jaw fragments, dromatheriids are mainly known from their sectorial (flesh-slicing) postcanine teeth. The teeth were fairly typical among early prozostrodontians, as they were labiolingually compressed (flattened sideways), with a single

root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the sur ...

and

crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...

hosting a longitudinal row of sharp cusps. Dromatheriids in particular have a very narrow and symmetrical crown (when seen from above) without a prominent cingulum (a ridge or array of cuspules adjacent to the main cusps). Dromatheriid teeth on average have four main cusps, though some have as few as two ('' Dromatherium'') or three (''

Tricuspes ''Tricuspes'' is an extinct genus of cynodonts that lived in what would be Europe during the Triassic from 203.6 to 199.6 mya, existing for approximately . Three species are known: ''Tricuspes tubingensis'' (Huene, 1933), ''Tricuspes sigogneaua ...

''), or as many as six ('' Inditherium'', '' Pseudotriconodon''). Although the teeth have a single root, a vertical furrow on each side of the root appears to be a trait incipient towards the two fully divided roots of

mammaliaforms Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recen ...

. Making note of this condition, some authors have suggested that dromatheriids are a

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

group ancestral to mammaliaforms. Other studies instead consider the closest relatives of dromatheriids to be the " therioherpetids" '' Therioherpeton'' and '' Meurthodon'', which may even be placed within the family. However, the broader cusps of ''Therioherpeton'' and the divided root of ''Meurthodon'' dissuade their position within Dromatheriidae.

References

Prehistoric prozostrodonts Prehistoric therapsid families {{Paleo-therapsid-stub