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The cat (''Felis catus'') is a
domestic Domestic may refer to: In the home * Anything relating to the human home or family ** A domestic animal, one that has undergone domestication ** A domestic appliance, or home appliance ** A domestic partnership ** Domestic science, sometimes c ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of the family. Cats are commonly kept as house pets but can also be
farm cat The farm cat, also known as a barn cat, is a domestic cat, usually of mixed breed, that lives primarily outdoors, in a feral or semi-feral condition on agricultural properties, usually sheltering in outbuildings. They eat assorted vermin such as ...
s or
feral cat A feral cat or a stray cat is an unowned domestic cat (''Felis catus'') that lives outdoors and avoids human contact: it does not allow itself to be handled or touched, and usually remains hidden from humans. Feral cats may breed over dozens ...
s; the feral cat ranges freely and avoids human contact. Domestic cats are valued by humans for companionship and their ability to kill
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
s. About 60
cat breeds The following list of cat breeds includes only domestic cat breeds and domestic and wild hybrids. The list includes established breeds recognized by various cat registries, new and experimental breeds, landraces being established as standardized ...
are recognized by various
cat registries A cat registry or cat breed registry, also known as a cat fancier organization, cattery federation, or cat breeders' association, is an organization that registers domestic cats (usually purebred) of many breeds, for exhibition and for breeding ...
. The cat is similar in
anatomy Anatomy () is the branch of biology concerned with the study of the structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old science, having it ...
to the other felid species: they have a strong flexible body, quick reflexes, sharp teeth, and
retractable claws A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or ta ...
adapted to killing small prey. Their night vision and sense of smell are well developed.
Cat communication Cats need to communicate with each other for bonding, and relating with each other; they need to collaborate, play, and share resources. When they communicate with people, they do so to get what they need or want, such as food, affection, or p ...
includes vocalizations like
meow A meow or miaow is a cat vocalization. ''Meows'' may have diverse tones and are sometimes chattered, murmured or whispered. Adult cats rarely meow to each other, so an adult cat meowing to human beings is probably a post-domestication exten ...
ing,
purr A purr is a tonal fluttering sound made by some species of felids and two species of genets. It varies in loudness and tone among species and in the same animal. Felids are a family of mammals that belong to the order Carnivora and are info ...
ing, trilling, hissing, growling, and grunting as well as cat-specific body language. Although the cat is a social species, they are a solitary hunter. As a
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
, they are crepuscular, i.e. most active at dawn and dusk. They can hear sounds too faint or too high in
frequency Frequency is the number of occurrences of a repeating event per unit of time. It is also occasionally referred to as ''temporal frequency'' for clarity, and is distinct from ''angular frequency''. Frequency is measured in hertz (Hz) which is eq ...
for human ears, such as those made by mice and other small mammals. They also secretes and perceive
pheromones A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
. Female domestic cats can have kittens from spring to late autumn, with litter sizes often ranging from two to five kittens. Domestic cats are bred and shown at events as registered pedigreed cats, a hobby known as
cat fancy The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members of ...
.
Population control Population control is the practice of artificially maintaining the size of any population. It simply refers to the act of limiting the size of an animal population so that it remains manageable, as opposed to the act of protecting a species from ...
of cats may be affected by
spaying Neutering, from the Latin ''neuter'' ('of neither sex'), is the removal of an animal's reproductive organ, either all of it or a considerably large part. The male-specific term is castration, while spaying is usually reserved for female animals. C ...
and neutering, but their proliferation and the abandonment of pets has resulted in large numbers of feral cats worldwide, contributing to the extinction of entire bird, mammal, and reptile species. It was long thought that cat domestication began in ancient Egypt, where cats were venerated from around 3100 BC, but recent advances in archaeology and genetics have shown that their domestication occurred in
Western Asia Western Asia, West Asia, or Southwest Asia, is the westernmost subregion of the larger geographical region of Asia, as defined by some academics, UN bodies and other institutions. It is almost entirely a part of the Middle East, and includes Ana ...
around 7500 BC. there were an estimated 220 million owned and 480 million stray cats in the world. the domestic cat was the second most popular pet in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territori ...
, with 95.6 million cats owned and around 42 million households owning at least one cat. In the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the European mainland, continental mainland. It comprises England, Scotlan ...
, 26% of adults have a cat, with an estimated population of 10.9 million pet cats


Etymology and naming

The origin of the English word ''cat'',
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlers in the mid-5th c ...
, is thought to be the
Late Latin Late Latin ( la, Latinitas serior) is the scholarly name for the form of Literary Latin of late antiquity.Roberts (1996), p. 537. English dictionary definitions of Late Latin date this period from the , and continuing into the 7th century in t ...
word , which was first used at the beginning of the 6th century. It was suggested that is derived from an
Egyptian Egyptian describes something of, from, or related to Egypt. Egyptian or Egyptians may refer to: Nations and ethnic groups * Egyptians, a national group in North Africa ** Egyptian culture, a complex and stable culture with thousands of years of ...
precursor of
Coptic Coptic may refer to: Afro-Asia * Copts, an ethnoreligious group mainly in the area of modern Egypt but also in Sudan and Libya * Coptic language, a Northern Afro-Asiatic language spoken in Egypt until at least the 17th century * Coptic alphabet ...
, "tomcat", or its feminine form suffixed with . The Late Latin word may be derived from another Afro-Asiatic or Nilo-Saharan language. The
Nubian Nubian may refer to: *Something of, from, or related to Nubia, a region along the Nile river in Southern Egypt and northern Sudan. *Nubian people *Nubian languages *Anglo-Nubian goat, a breed of goat * Nubian ibex * , several ships of the Britis ...
word "wildcat" and
Nobiin Nobiin, or Mahas, is a Northern Nubian language of the Nilo-Saharan language family. "Nobiin" is the genitive form of ''Nòòbíí'' ("Nubian") and literally means "(language) of the Nubians". Another term used is ''Noban tamen'', meaning "the ...
are possible sources or cognates. The Nubian word may be a loan from
Arabic Arabic (, ' ; , ' or ) is a Semitic language spoken primarily across the Arab world.Semitic languages: an international handbook / edited by Stefan Weninger; in collaboration with Geoffrey Khan, Michael P. Streck, Janet C. E.Watson; Walter ...
~ . However, it is "equally likely that the forms might derive from an ancient Germanic word, imported into Latin and thence to Greek and to Syriac and Arabic". The word may be derived from Germanic and Northern European languages, and ultimately be borrowed from
Uralic The Uralic languages (; sometimes called Uralian languages ) form a language family of 38 languages spoken by approximately 25million people, predominantly in Northern Eurasia. The Uralic languages with the most native speakers are Hungarian (w ...
, cf.
Northern Sami Northern may refer to the following: Geography * North, a point in direction * Northern Europe, the northern part or region of Europe * Northern Highland, a region of Wisconsin, United States * Northern Province, Sri Lanka * Northern Range, a ...
, "female
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine, Beringian ermine and ermine, is a mustelid native to Eurasia and the northern portions of North America. Because of its wide circumpolar distribution, it is listed as Least Conc ...
", and Hungarian , "lady, female stoat"; from
Proto-Uralic Proto-Uralic is the unattested reconstructed language ancestral to the modern Uralic language family. The hypothetical language is believed to have been originally spoken in a small area in about 7000–2000 BCE, and expanded to give different ...
''*käďwä'', "female (of a furred animal)". The English ''
puss Puss may refer to: Arts, entertainment, and media * ''Puss'' (magazine), Swedish magazine published between 1968 and 1973 * ''Puss'' (film), a Swedish film dated 2010 * "Puss" (song), a noise rock song dated 1992 * Tom Puss, a fictional anthro ...
'', extended as ''pussy'' and ''pussycat'', is attested from the 16th century and may have been introduced from
Dutch Dutch commonly refers to: * Something of, from, or related to the Netherlands * Dutch people () * Dutch language () Dutch may also refer to: Places * Dutch, West Virginia, a community in the United States * Pennsylvania Dutch Country People E ...
or from Low German , related to
Swedish Swedish or ' may refer to: Anything from or related to Sweden, a country in Northern Europe. Or, specifically: * Swedish language, a North Germanic language spoken primarily in Sweden and Finland ** Swedish alphabet, the official alphabet used by ...
, or
Norwegian Norwegian, Norwayan, or Norsk may refer to: *Something of, from, or related to Norway, a country in northwestern Europe * Norwegians, both a nation and an ethnic group native to Norway * Demographics of Norway *The Norwegian language, including ...
, . Similar forms exist in Lithuanian and
Irish Irish may refer to: Common meanings * Someone or something of, from, or related to: ** Ireland, an island situated off the north-western coast of continental Europe ***Éire, Irish language name for the isle ** Northern Ireland, a constituent unit ...
or . The etymology of this word is unknown, but it may have arisen from a sound used to attract a cat. A male cat is called a ''tom'' or ''tomcat'' (or a ''gib'', if neutered). An unspayed female is called a ''queen'' (or a ''molly'', if spayed), especially in a cat-breeding context. A juvenile cat is referred to as a '' kitten''. In
Early Modern English Early Modern English or Early New English (sometimes abbreviated EModE, EMnE, or ENE) is the stage of the English language from the beginning of the Tudor period to the English Interregnum and Restoration, or from the transition from Middle E ...
, the word ''kitten'' was interchangeable with the now-obsolete word ''catling''. A group of cats can be referred to as a ''clowder'' or a ''glaring''.


Taxonomy

The scientific name ''Felis catus'' was proposed by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his Nobility#Ennoblement, ennoblement in 1761 as Carl von Linné#Blunt, Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalise ...
in 1758 for a domestic cat. ''Felis catus domesticus'' was proposed by
Johann Christian Polycarp Erxleben Johann Christian Polycarp Erxleben was a German naturalist from Quedlinburg. Erxleben was professor of physics and veterinary medicine at the University of Göttingen The University of Göttingen, officially the Georg August University of ...
in 1777. ''Felis daemon'' proposed by
Konstantin Alekseevich Satunin Konstantin Alekseevich Satunin (20 May 1863–10 November 1915) was a Russian zoologist who studied and described many mammals found in Russia and Central Asia. Satunin graduated from Moscow University in 1890. From 1893 he worked at a sericultur ...
in 1904 was a black cat from the
Transcaucasus The South Caucasus, also known as Transcaucasia or the Transcaucasus, is a geographical region on the border of Eastern Europe and Western Asia, straddling the southern Caucasus Mountains. The South Caucasus roughly corresponds to modern Arme ...
, later identified as a domestic cat. In 2003, the International Commission on Zoological Nomenclature ruled that the domestic cat is a distinct species, namely ''Felis catus''. In 2007, it was considered a subspecies, ''F. silvestris catus'', of the
European wildcat The European wildcat (''Felis silvestris'') is a small wildcat species native to continental Europe, Scotland, Turkey and the Caucasus. It inhabits forests from the Iberian Peninsula, Italy, Central and Eastern Europe to the Caucasus. Its fur i ...
(''F. silvestris'') following results of
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
research. In 2017, the IUCN Cat Classification Taskforce followed the recommendation of the ICZN in regarding the domestic cat as a distinct species, ''Felis catus''.


Evolution

The domestic cat is a member of the Felidae, a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
that had a
common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. All living beings are in fact descendants of a unique ancestor commonly referred to as the last universal comm ...
about 10–15 million years ago. The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Felis'' diverged from other Felidae around 6–7 million years ago. Results of
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
research confirm that the wild ''Felis'' species evolved through sympatric or
parapatric speciation In parapatric speciation, two subpopulations of a species evolve reproductive isolation from one another while continuing to exchange genes. This mode of speciation has three distinguishing characteristics: 1) mating occurs non-randomly, 2) gene ...
, whereas the domestic cat evolved through artificial selection. The domesticated cat and its closest wild ancestor are diploid and both possess 38
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
s and roughly 20,000 genes. The
leopard cat The leopard cat (''Prionailurus bengalensis'') is a small wild cat native to continental South, Southeast, and East Asia. Since 2002 it has been listed as Least Concern on the IUCN Red List as it is widely distributed although threatened by hab ...
(''Prionailurus bengalensis'') was
tamed A tame animal is an animal that is relatively tolerant of human presence. Tameness may arise naturally (as in the case, for example, of island tameness) or due to the deliberate, human-directed process of training an animal against its initially ...
independently in China around 5500 BC. This line of partially domesticated cats leaves no trace in the domestic cat populations of today.


Domestication

The earliest known indication for the
taming A tame animal is an animal that is relatively tolerant of human presence. Tameness may arise naturally (as in the case, for example, of island tameness) or due to the deliberate, human-directed process of training an animal against its initially ...
of an
African wildcat The African wildcat (''Felis lybica'') is a small wildcat species native to Africa, West and Central Asia up to Rajasthan in India and Xinjiang in China. It has been listed as Least Concern on the IUCN Red List in 2022. In Cyprus, an African wil ...
(''F. lybica'') was excavated close by a human
Neolithic The Neolithic period, or New Stone Age, is an Old World archaeological period and the final division of the Stone Age. It saw the Neolithic Revolution, a wide-ranging set of developments that appear to have arisen independently in several p ...
grave in
Shillourokambos Shillourokambos ( el, Σιλλουρόκαμπος) is a Pre-Pottery Neolithic B (PPNB) site near Parekklisia, 6 km east of Limassol in southern Cyprus. It is located on a low plateau. Excavations began in 1992. The settlement has four phases ...
, southern
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is ge ...
, dating to about 7500–7200 BC. Since there is no evidence of native mammalian
fauna Fauna is all of the animal life present in a particular region or time. The corresponding term for plants is ''flora'', and for fungi, it is ''funga''. Flora, fauna, funga and other forms of life are collectively referred to as ''Biota (ecology ...
on Cyprus, the inhabitants of this Neolithic village most likely brought the cat and other wild mammals to the island from the
Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabia (including the Arabian Peninsula and Bahrain), Asia Minor (Asian part of Turkey except Hatay Province), East Thrace (Europ ...
ern mainland. Scientists therefore assume that African wildcats were attracted to early human settlements in the
Fertile Crescent The Fertile Crescent ( ar, الهلال الخصيب) is a crescent-shaped region in the Middle East, spanning modern-day Iraq, Syria, Lebanon, Israel, Palestine and Jordan, together with the northern region of Kuwait, southeastern region of ...
by rodents, in particular the house mouse (''Mus musculus''), and were tamed by Neolithic farmers. This mutual relationship between early farmers and tamed cats lasted thousands of years. As
agricultural practices Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people ...
spread, so did tame and domesticated cats. Wildcats of Egypt contributed to the maternal gene pool of the domestic cat at a later time. The earliest known evidence for the occurrence of the domestic cat in
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders ...
dates to around 1200 BC. Greek,
Phoenicia Phoenicia () was an ancient thalassocratic civilization originating in the Levant region of the eastern Mediterranean, primarily located in modern Lebanon. The territory of the Phoenician city-states extended and shrank throughout their histor ...
n,
Carthaginia Carthage was the capital city of Ancient Carthage, on the eastern side of the Lake of Tunis in what is now Tunisia. Carthage was one of the most important trading hubs of the Ancient Mediterranean and one of the most affluent cities of the cl ...
n and
Etrusca The Etruscan civilization () was developed by a people of Etruria in ancient Italy with a common language and culture who formed a federation of city-states. After conquering adjacent lands, its territory covered, at its greatest extent, roughl ...
n traders introduced domestic cats to southern Europe. During the
Roman Empire The Roman Empire ( la, Imperium Romanum ; grc-gre, Βασιλεία τῶν Ῥωμαίων, Basileía tôn Rhōmaíōn) was the post- Republican period of ancient Rome. As a polity, it included large territorial holdings around the Mediter ...
they were introduced to Corsica and
Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...
before the beginning of the 1st millennium. By the 5th century BC, they were familiar animals around settlements in Magna Graecia and Etruria. By the end of the Western Roman Empire in the 5th century, the Egyptian domestic cat lineage had arrived in a
Baltic Sea The Baltic Sea is an arm of the Atlantic Ocean that is enclosed by Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Poland, Russia, Sweden and the North and Central European Plain. The sea stretches from 53°N to 66°N latitude and ...
port in northern
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
. During domestication, cats have undergone only minor changes in anatomy and behavior, and they are still capable of surviving in the wild. Several natural behaviors and characteristics of wildcats may have
pre-adapted Exaptation and the related term Co-option (biology), co-option describe a shift in the function of a trait during evolution. For example, a trait can evolve because it served one particular function, but subsequently it may come to serve another. E ...
them for domestication as pets. These traits include their small size, social nature, obvious body language, love of play and relatively high intelligence. Captive ''
Leopardus ''Leopardus'' is a genus comprising eight species of small cats native to the Americas. This genus is considered the oldest branch of a genetic lineage of small cats in the Americas whose common ancestor crossed the Bering land bridge from Asia ...
'' cats may also display affectionate behavior toward humans but were not domesticated. House cats often mate with feral cats. Hybridisation between domestic and other Felinae species is also possible, producing hybrids such as the Kellas cat in
Scotland Scotland (, ) is a Countries of the United Kingdom, country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a Anglo-Scottish border, border with England to the southeast ...
. Development of
cat breeds The following list of cat breeds includes only domestic cat breeds and domestic and wild hybrids. The list includes established breeds recognized by various cat registries, new and experimental breeds, landraces being established as standardized ...
started in the mid 19th century. An analysis of the domestic cat
genome In the fields of molecular biology and genetics, a genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding g ...
revealed that the ancestral wildcat genome was significantly altered in the process of domestication, as specific
mutation In biology, a mutation is an alteration in the nucleic acid sequence of the genome of an organism, virus, or extrachromosomal DNA. Viral genomes contain either DNA or RNA. Mutations result from errors during DNA replication, DNA or viral repl ...
s were selected to develop cat breeds. Most breeds are founded on random-bred domestic cats. Genetic diversity of these breeds varies between regions, and is lowest in purebred populations, which show more than 20 deleterious
genetic disorder A genetic disorder is a health problem caused by one or more abnormalities in the genome. It can be caused by a mutation in a single gene (monogenic) or multiple genes (polygenic) or by a chromosomal abnormality. Although polygenic disorders ...
s.


Characteristics


Size

The domestic cat has a smaller
skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
and shorter bones than the
European wildcat The European wildcat (''Felis silvestris'') is a small wildcat species native to continental Europe, Scotland, Turkey and the Caucasus. It inhabits forests from the Iberian Peninsula, Italy, Central and Eastern Europe to the Caucasus. Its fur i ...
. It averages about in head-to-body length and in height, with about long tails. Males are larger than females. Adult domestic cats typically weigh between .


Skeleton

Cats have seven cervical vertebrae (as do most
mammals Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur o ...
); 13
thoracic vertebrae In vertebrates, thoracic vertebrae compose the middle segment of the vertebral column, between the cervical vertebrae and the lumbar vertebrae. In humans, there are twelve thoracic vertebrae and they are intermediate in size between the cervical ...
(humans have 12); seven
lumbar vertebrae The lumbar vertebrae are, in human anatomy, the five vertebrae between the rib cage and the pelvis. They are the largest segments of the vertebral column and are characterized by the absence of the foramen transversarium within the transverse p ...
(humans have five); three sacral vertebrae (as do most mammals, but humans have five); and a variable number of caudal vertebrae in the tail (humans have only three to five vestigial caudal vertebrae, fused into an internal
coccyx The coccyx ( : coccyges or coccyxes), commonly referred to as the tailbone, is the final segment of the vertebral column in all apes, and analogous structures in certain other mammals such as horses. In tailless primates (e.g. humans and othe ...
). The extra lumbar and thoracic vertebrae account for the cat's spinal mobility and flexibility. Attached to the spine are 13 ribs, the shoulder, and the pelvis. Unlike human arms, cat forelimbs are attached to the shoulder by free-floating
clavicle The clavicle, or collarbone, is a slender, S-shaped long bone approximately 6 inches (15 cm) long that serves as a strut between the shoulder blade and the sternum (breastbone). There are two clavicles, one on the left and one on the r ...
bones which allow them to pass their body through any space into which they can fit their head.


Skull

The cat skull is unusual among mammals in having very large
eye socket In anatomy, the orbit is the cavity or socket of the skull in which the eye and its appendages are situated. "Orbit" can refer to the bony socket, or it can also be used to imply the contents. In the adult human, the volume of the orbit is , o ...
s and a powerful specialized jaw. Within the jaw, cats have teeth adapted for killing prey and tearing meat. When it overpowers its prey, a cat delivers a lethal neck bite with its two long
canine teeth In mammalian oral anatomy, the canine teeth, also called cuspids, dog teeth, or (in the context of the upper jaw) fangs, eye teeth, vampire teeth, or vampire fangs, are the relatively long, pointed teeth. They can appear more flattened howeve ...
, inserting them between two of the prey's
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...
e and severing its
spinal cord The spinal cord is a long, thin, tubular structure made up of nervous tissue, which extends from the medulla oblongata in the brainstem to the lumbar region of the vertebral column (backbone). The backbone encloses the central canal of the sp ...
, causing irreversible paralysis and death. Compared to other felines, domestic cats have narrowly spaced canine teeth relative to the size of their jaw, which is an adaptation to their preferred prey of small rodents, which have small vertebrae. The premolar and first molar together compose the
carnassial Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified f ...
pair on each side of the mouth, which efficiently shears meat into small pieces, like a pair of scissors. These are vital in feeding, since cats' small
molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...
cannot chew food effectively, and cats are largely incapable of mastication. Although cats tend to have better teeth than most humans, with decay generally less likely because of a thicker protective layer of enamel, a less damaging saliva, less retention of food particles between teeth, and a diet mostly devoid of sugar, they are nonetheless subject to occasional tooth loss and infection.


Claws

Cats have protractible and retractable claws. In their normal, relaxed position, the claws are sheathed with the skin and fur around the paw's toe pads. This keeps the claws sharp by preventing wear from contact with the ground and allows for the silent stalking of prey. The claws on the forefeet are typically sharper than those on the hindfeet. Cats can voluntarily extend their claws on one or more paws. They may extend their claws in hunting or self-defense, climbing,
kneading In cooking (and more specifically baking), kneading is a process in the making of bread or dough, used to mix the ingredients and add strength to the final product. Its importance lies in the mixing of flour with water; when these two ingred ...
, or for extra traction on soft surfaces. Cats shed the outside layer of their claw sheaths when scratching rough surfaces. Most cats have five claws on their front paws and four on their rear paws. The
dewclaw A dewclaw is a digit – vestigial in some animals – on the foot of many mammals, birds, and reptiles (including some extinct orders, like certain theropods). It commonly grows higher on the leg than the rest of the foot, such that in digit ...
is
proximal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
to the other claws. More proximally is a protrusion which appears to be a sixth "finger". This special feature of the front paws on the inside of the wrists has no function in normal walking but is thought to be an antiskidding device used while jumping. Some cat breeds are prone to having extra digits ("
polydactyly Polydactyly or polydactylism (), also known as hyperdactyly, is an anomaly in humans and animals resulting in supernumerary fingers and/or toes. Polydactyly is the opposite of oligodactyly (fewer fingers or toes). Signs and symptoms In human ...
"). Polydactylous cats occur along North America's northeast coast and in Great Britain.


Ambulation

The cat is digitigrade. It walks on the toes, with the bones of the feet making up the lower part of the visible leg. Unlike most mammals, it uses a "pacing" gait and moves both legs on one side of the body before the legs on the other side. It registers directly by placing each hind paw close to the track of the corresponding fore paw, minimizing noise and visible tracks. This also provides sure footing for hind paws when navigating rough terrain. As it speeds up from walking to trotting, its gait changes to a "diagonal" gait: The diagonally opposite hind and fore legs move simultaneously.


Balance

Most breeds of cat are notably fond of sitting in high places, or ''perching''. A higher place may serve as a concealed site from which to hunt; domestic cats strike prey by pouncing from a perch such as a tree branch. Another possible explanation is that height gives the cat a better observation point, allowing it to survey its territory. A cat falling from heights of up to can right itself and land on its paws. During a fall from a high place, a cat reflexively twists its body and rights itself to land on its feet using its acute sense of balance and flexibility. This reflex is known as the cat righting reflex. A cat always rights itself in the same way during a fall, if it has enough time to do so, which is the case in falls of or more. How cats are able to right themselves when falling has been investigated as the "
falling cat problem The falling cat problem is a problem that consists of explaining the underlying physics behind the observation of the cat righting reflex. Although amusing and trivial to pose, the solution of the problem is not as straightforward as its stat ...
".


Coats

The cat family (Felidae) can pass down many colors and patterns to their offsprings. The domestic cat genes ''MC1R'' and ''ASIP'' allow for the variety of color in coats. The feline ASIP gene consists of three coding exons. Three novel microsatellite markers linked to ASIP were isolated from a domestic cat BAC clone containing this gene and were used to perform
linkage analysis Genetic linkage is the tendency of DNA sequences that are close together on a chromosome to be inherited together during the meiosis phase of sexual reproduction. Two genetic markers that are physically near to each other are unlikely to be separ ...
in a pedigree of 89 domestic cats that segregated for
melanism The term melanism refers to black pigment and is derived from the gr, μελανός. Melanism is the increased development of the dark-colored pigment melanin in the skin or hair. Pseudomelanism, also called abundism, is another variant of ...
.


Senses


Vision

Cats have excellent night vision and can see at only one-sixth the light level required for human vision. This is partly the result of cat eyes having a ''
tapetum lucidum The ''tapetum lucidum'' ( ; ; ) is a layer of tissue in the eye of many vertebrates and some other animals. Lying immediately behind the retina, it is a retroreflector. It reflects visible light back through the retina, increasing the light a ...
'', which reflects any light that passes through the
retina The retina (from la, rete "net") is the innermost, light-sensitive layer of tissue of the eye of most vertebrates and some molluscs. The optics of the eye create a focused two-dimensional image of the visual world on the retina, which then ...
back into the eye, thereby increasing the eye's sensitivity to dim light. Large pupils are an adaptation to dim light. The domestic cat has slit pupils, which allow it to focus bright light without
chromatic aberration In optics, chromatic aberration (CA), also called chromatic distortion and spherochromatism, is a failure of a lens to focus all colors to the same point. It is caused by dispersion: the refractive index of the lens elements varies with the w ...
. At low light, a cat's pupils expand to cover most of the exposed surface of its eyes. The domestic cat has rather poor color vision and only two types of cone cells, optimized for sensitivity to blue and yellowish green; its ability to distinguish between red and green is limited. A response to middle wavelengths from a system other than the rod cells might be due to a third type of cone. This appears to be an adaptation to low light levels rather than representing true
trichromatic Trichromacy or trichromatism is the possessing of three independent channels for conveying color information, derived from the three different types of cone cells in the eye. Organisms with trichromacy are called trichromats. The normal expl ...
vision. Cats also have a nictitating membrane, allowing them to blink without hindering their vision.


Hearing

The domestic cat's hearing is most acute in the range of 500 Hz to 32 kHz. It can detect an extremely broad range of frequencies ranging from 55 Hz to 79 kHz. It can hear a range of 10.5 octaves, while humans and dogs can hear ranges of about 9 octaves. Its hearing sensitivity is enhanced by its large movable outer ears, the
pinnae The auricle or auricula is the visible part of the ear that is outside the head. It is also called the pinna (Latin for "wing" or " fin", plural pinnae), a term that is used more in zoology. Structure The diagram shows the shape and location ...
, which amplify sounds and help detect the location of a noise. It can detect
ultrasound Ultrasound is sound waves with frequencies higher than the upper audible limit of human hearing. Ultrasound is not different from "normal" (audible) sound in its physical properties, except that humans cannot hear it. This limit varies ...
, which enables it to detect ultrasonic calls made by
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are n ...
prey. Recent research has shown that cats have socio-spatial cognitive abilities to create mental maps of owners' locations based on hearing owners' voices.


Smell

Cats have an acute sense of smell, due in part to their well-developed
olfactory bulb The olfactory bulb (Latin: ''bulbus olfactorius'') is a neural structure of the vertebrate forebrain involved in olfaction, the sense of smell. It sends olfactory information to be further processed in the amygdala, the orbitofrontal cortex ( ...
and a large surface of
olfactory mucosa The olfactory mucosa is located in the upper region of the nasal cavity and is made up of the olfactory epithelium and the underlying lamina propria, connective tissue containing fibroblasts, blood vessels, Bowman's glands and bundles of fine ax ...
, about in area, which is about twice that of humans. Cats and many other animals have a
Jacobson's organ The vomeronasal organ (VNO), or Jacobson's organ, is the paired auxiliary olfactory (smell) sense organ located in the soft tissue of the nasal septum, in the nasal cavity just above the roof of the mouth (the hard palate) in various tetrapods. T ...
in their mouths that is used in the behavioral process of flehmening. It allows them to sense certain aromas in a way that humans cannot. Cats are sensitive to
pheromones A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
such as 3-mercapto-3-methylbutan-1-ol, which they use to communicate through
urine spraying Urination, also known as micturition, is the release of urine from the urinary bladder through the urethra to the outside of the body. It is the urinary system's form of excretion. It is also known medically as micturition, voiding, uresis, ...
and marking with
scent glands Scent gland are exocrine glands found in most mammals. They produce semi-viscous secretions which contain pheromones and other semiochemical compounds. These odor-messengers indicate information such as status, territorial marking, mood, and sexu ...
. Many cats also respond strongly to plants that contain
nepetalactone Nepetalactone is a name for multiple iridoid analog stereoisomers. Nepetalactones are produced by ''Nepeta cataria'' ( catnip) and many other plants belonging to the genus '' Nepeta'', in which they protect these plants from herbivorous insects b ...
, especially
catnip ''Nepeta cataria'', commonly known as catnip, catswort, catwort, and catmint, is a species of the genus '' Nepeta'' in the family Lamiaceae, native to southern and eastern Europe, the Middle East, Central Asia, and parts of China. It is widely ...
, as they can detect that substance at less than one part per billion. About 70–80% of cats are affected by nepetalactone. This response is also produced by other plants, such as silver vine (''
Actinidia polygama ''Actinidia polygama'' (also known as , silver vine, , and cat powder) is a species of kiwifruit in the Actinidiaceae family. It grows in the mountainous areas of Korea, Japan and China at elevations between . Silver vine can reach up to high ...
'') and the herb valerian; it may be caused by the smell of these plants mimicking a pheromone and stimulating cats' social or sexual behaviors.


Taste

Cats have relatively few
taste bud Taste buds contain the taste receptor cells, which are also known as gustatory cells. The taste receptors are located around the small structures known as papillae found on the upper surface of the tongue, soft palate, upper esophagus, the c ...
s compared to humans (470 or so versus more than 9,000 on the human tongue). Domestic and wild cats share a taste receptor gene mutation that keeps their sweet taste buds from binding to sugary molecules, leaving them with no ability to taste sweetness. Their taste buds instead respond to acids,
amino acid Amino acids are organic compounds that contain both amino and carboxylic acid functional groups. Although hundreds of amino acids exist in nature, by far the most important are the alpha-amino acids, which comprise proteins. Only 22 alpha a ...
s like protein, and bitter tastes. Cats also have a distinct temperature preference for their food, preferring food with a temperature around which is similar to that of a fresh kill and routinely rejecting food presented cold or refrigerated (which would signal to the cat that the "prey" item is long dead and therefore possibly toxic or decomposing).


Whiskers

To aid with navigation and sensation, cats have dozens of movable whiskers (vibrissae) over their body, especially their faces. These provide information on the width of gaps and on the location of objects in the dark, both by touching objects directly and by sensing air currents; they also trigger protective
blink reflex The corneal reflex, also known as the blink reflex or eyelid reflex, is an involuntary blinking of the eyelids elicited by stimulation of the cornea (such as by touching or by a foreign body), though it could result from any peripheral stimulus. S ...
es to protect the eyes from damage.


Behavior

Outdoor cats are active both day and night, although they tend to be slightly more active at night. Domestic cats spend the majority of their time in the vicinity of their homes but can range many hundreds of meters from this central point. They establish
territories A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
that vary considerably in size, in one study ranging from . The timing of cats' activity is quite flexible and varied, which means house cats may be more active in the morning and evening, as a response to greater human activity at these times. Cats conserve energy by sleeping more than most animals, especially as they grow older. The daily duration of sleep varies, usually between 12 and 16 hours, with 13 and 14 being the average. Some cats can sleep as much as 20 hours. The term "cat nap" for a short rest refers to the cat's tendency to fall asleep (lightly) for a brief period. While asleep, cats experience short periods of
rapid eye movement sleep Rapid eye movement sleep (REM sleep or REMS) is a unique phase of sleep in mammals and birds, characterized by random rapid movement of the eyes, accompanied by low muscle tone throughout the body, and the propensity of the sleeper to dream ...
often accompanied by muscle twitches, which suggests they are dreaming.


Sociability

The social behavior of the domestic cat ranges from widely dispersed individuals to
feral cat colonies A feral cat or a stray cat is an unowned domestic cat (''Felis catus'') that lives outdoors and avoids human contact: it does not allow itself to be handled or touched, and usually remains hidden from humans. Feral cats may breed over dozens ...
that gather around a food source, based on groups of co-operating females. Within such groups, one cat is usually dominant over the others. Each cat in a colony holds a distinct territory, with sexually active males having the largest territories, which are about 10 times larger than those of female cats and may overlap with several females' territories. These territories are marked by
urine spraying Urination, also known as micturition, is the release of urine from the urinary bladder through the urethra to the outside of the body. It is the urinary system's form of excretion. It is also known medically as micturition, voiding, uresis, ...
, by rubbing objects at head height with secretions from facial glands, and by defecation. Between these territories are neutral areas where cats watch and greet one another without territorial conflicts. Outside these neutral areas, territory holders usually chase away stranger cats, at first by staring, hissing, and growling and, if that does not work, by short but noisy and violent attacks. Despite this colonial organization, cats do not have a social survival strategy or a
pack mentality Herd behavior is the behavior of individuals in a group acting collectively without centralized direction. Herd behavior occurs in animals in herds, packs, bird flocks, fish schools and so on, as well as in humans. Voting, demonstrations, riot ...
, and always hunt alone. Life in proximity to humans and other domestic animals has led to a symbiotic social adaptation in cats, and cats may express great affection toward humans or other animals. Ethologically, a cat's human keeper functions as if a mother surrogate. Adult cats live their lives in a kind of extended kittenhood, a form of behavioral
neoteny Neoteny (), also called juvenilization,Montagu, A. (1989). Growing Young. Bergin & Garvey: CT. is the delaying or slowing of the physiological, or somatic, development of an organism, typically an animal. Neoteny is found in modern humans compa ...
. Their high-pitched sounds may mimic the cries of a hungry human infant, making them particularly difficult for humans to ignore. Some pet cats are poorly socialized. In particular, older cats show aggressiveness toward newly arrived kittens, which include biting and scratching; this type of behavior is known as feline asocial aggression.
Redirected aggression Displaced aggression, also referred to as redirected aggression, occurs when an animal or human is fearful or agitated by external stimulus, external stimuli, a provocation, or perception, but is unable or unwilling to direct their aggression t ...
is a common form of aggression which can occur in multiple cat households. In redirected aggression there is usually something that agitates the cat: this could be a sight, sound, or another source of stimuli which causes a heightened level of anxiety or arousal. If the cat cannot attack the stimuli, it may direct anger elsewhere by attacking or directing aggression to the nearest cat, dog, human or other being. Domestic cats' scent rubbing behavior toward humans or other cats is thought to be a feline means for social bonding.


Communication

Domestic cats use many vocalizations for communication, including
purr A purr is a tonal fluttering sound made by some species of felids and two species of genets. It varies in loudness and tone among species and in the same animal. Felids are a family of mammals that belong to the order Carnivora and are info ...
ing, trilling, hissing, growling/snarling, grunting, and several different forms of meowing. Their
body language Body language is a type of communication in which physical behaviors, as opposed to words, are used to express or convey information. Such behavior includes facial expressions, body posture, gestures, eye movement, touch and the use of space. Th ...
, including position of ears and tail, relaxation of the whole body, and kneading of the paws, are all indicators of mood. The tail and ears are particularly important social signal mechanisms in cats. A raised tail indicates a friendly greeting, and flattened ears indicates hostility. Tail-raising also indicates the cat's position in the group's
social hierarchy Social stratification refers to a society's categorization of its people into groups based on socioeconomic factors like wealth, income, race, education, ethnicity, gender, occupation, social status, or derived power (social and political). As ...
, with dominant individuals raising their tails less often than subordinate ones. Feral cats are generally silent. Nose-to-nose touching is also a common greeting and may be followed by social grooming, which is solicited by one of the cats raising and tilting its head.
Purring A purr is a tonal fluttering sound made by some species of felids and two species of genets. It varies in loudness and tone among species and in the same animal. Felids are a family of mammals that belong to the order Carnivora and are info ...
may have developed as an evolutionary advantage as a signaling mechanism of reassurance between mother cats and
nursing Nursing is a profession within the health care sector focused on the care of individuals, families, and communities so they may attain, maintain, or recover optimal health and quality of life. Nurses may be differentiated from other health ...
kittens, who are thought to use it as a care-soliciting signal. Post-nursing cats also often purr as a sign of contentment: when being petted, becoming relaxed, or eating. Even though purring as popularly interpreted as indicative of pleasure, it has been recorded in a wide variety of circumstances, most of which involve physical contact between the cat and another, presumably trusted individual. Some cats have been observed to purr continuously when chronically ill or in apparent pain. The exact mechanism by which cats purr has long been elusive, but it has been proposed that purring is generated via a series of sudden build-ups and releases of pressure as the glottis is opened and closed, which causes the
vocal folds In humans, vocal cords, also known as vocal folds or voice reeds, are folds of throat tissues that are key in creating sounds through vocalization. The size of vocal cords affects the pitch of voice. Open when breathing and vibrating for speec ...
to separate forcefully. The laryngeal muscles in control of the glottis are thought to be driven by a
neural oscillator In biology, the nervous system is the highly complex part of an animal that coordinates its actions and sensory information by transmitting signals to and from different parts of its body. The nervous system detects environmental changes th ...
which generates a cycle of contraction and release every 30-40 milliseconds (giving a frequency of 33 to 25 Hz).


Grooming

Cats are known for spending considerable amounts of time licking their coats to keep them clean. The cat's tongue has backward-facing spines about 500 μm long, which are called papillae. These contain
keratin Keratin () is one of a family of structural fibrous proteins also known as ''scleroproteins''. Alpha-keratin (α-keratin) is a type of keratin found in vertebrates. It is the key structural material making up scales, hair, nails, feathers, ho ...
which makes them rigid so the papillae act like a hairbrush. Some cats, particularly longhaired cats, occasionally regurgitate
hairball A hairball is a small collection of hair or fur formed in the stomach of animals, and uncommonly in humans, that is occasionally vomited up when it becomes too big. Hairballs are primarily a tight elongated cylinder of packed fur, but may inclu ...
s of fur that have collected in their stomachs from grooming. These clumps of fur are usually sausage-shaped and about long. Hairballs can be prevented with remedies that ease elimination of the hair through the gut, as well as regular grooming of the coat with a comb or stiff brush.


Fighting

Among domestic cats, males are more likely to fight than females. Among feral cats, the most common reason for cat fighting is competition between two males to mate with a female. In such cases, most fights are won by the heavier male. Another common reason for fighting in domestic cats is the difficulty of establishing territories within a small home. Female cats also fight over territory or to defend their kittens. Neutering will decrease or eliminate this behavior in many cases, suggesting that the behavior is linked to
sex hormone Sex hormones, also known as sex steroids, gonadocorticoids and gonadal steroids, are steroid hormones that interact with vertebrate steroid hormone receptors. The sex hormones include the androgens, estrogens, and progestogens. Their effect ...
s. When cats become aggressive, they try to make themselves appear larger and more threatening by raising their fur, arching their backs, turning sideways and hissing or spitting. Often, the ears are pointed down and back to avoid damage to the inner ear and potentially listen for any changes behind them while focused forward. Cats may also vocalize loudly and bare their teeth in an effort to further intimidate their opponents. Fights usually consist of grappling and delivering powerful slaps to the face and body with the forepaws as well as bites. Cats also throw themselves to the ground in a defensive posture to rake their opponent's belly with their powerful hind legs. Serious damage is rare, as the fights are usually short in duration, with the loser running away with little more than a few scratches to the face and ears. Fights for mating rights are typically more severe and injuries may include deep puncture wounds and lacerations. Normally, serious injuries from fighting are limited to infections of scratches and bites, though these can occasionally kill cats if untreated. In addition, bites are probably the main route of transmission of
feline immunodeficiency virus Feline immunodeficiency virus (FIV) is a ''Lentivirus'' that affects cats worldwide, with 2.5% to 4.4% of felines being infected. FIV was first isolated in 1986, by Niels C Pedersen and Janet K. Yamamoto at the UC Davis School of Veterinary M ...
. Sexually active males are usually involved in many fights during their lives, and often have decidedly battered faces with obvious scars and cuts to their ears and nose. Cats are willing to threaten animals larger than them to defend their territory, such as dogs and
foxes Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...
.


Hunting and feeding

The shape and structure of cats' cheeks is insufficient to allow them to take in liquids using suction. Therefore, when drinking they lap with the tongue to draw liquid upward into their mouths. Lapping at a rate of four times a second, the cat touches the smooth tip of its tongue to the surface of the water, and quickly retracts it like a corkscrew, drawing water upward. Feral cats and free-fed house cats consume several small meals in a day. The frequency and size of meals varies between individuals. They select food based on its temperature, smell and texture; they dislike chilled foods and respond most strongly to moist foods rich in amino acids, which are similar to meat. Cats reject novel flavors (a response termed
neophobia Neophobia is the fear of anything new, especially a persistent and abnormal fear. In its milder form, it can manifest as the unwillingness to try new things or break from routine. In the context of children the term is generally used to indicate a ...
) and learn quickly to avoid foods that have tasted unpleasant in the past. It is also a common misconception that cats like milk/cream, as they tend to avoid sweet food and milk. Most adult cats are
lactose intolerant Lactose intolerance is a common condition caused by a decreased ability to digest lactose, a sugar found in dairy products. Those affected vary in the amount of lactose they can tolerate before symptoms develop. Symptoms may include abdominal ...
; the sugar in milk is not easily digested and may cause soft stools or
diarrhea Diarrhea, also spelled diarrhoea, is the condition of having at least three loose, liquid, or watery bowel movements each day. It often lasts for a few days and can result in dehydration due to fluid loss. Signs of dehydration often begin w ...
. Some also develop odd eating habits and like to eat or chew on things like wool, plastic, cables, paper, string, aluminum foil, or even coal. This condition, pica, can threaten their health, depending on the amount and toxicity of the items eaten. Cats hunt small prey, primarily birds and rodents, and are often used as a form of pest control. Cats use two hunting strategies, either stalking prey actively, or waiting in ambush until an animal comes close enough to be captured. The strategy used depends on the prey species in the area, with cats waiting in ambush outside burrows, but tending to actively stalk birds. Domestic cats are a major predator of wildlife in the United States, killing an estimated 1.3 to 4.0 billion birds and 6.3 to 22.3 billion mammals annually. Certain species appear more susceptible than others; in one English village, for example, 30% of house sparrow mortality was linked to the domestic cat. In the recovery of ringed robins (''
Erithacus rubecula The European robin (''Erithacus rubecula''), known simply as the robin or robin redbreast in Great Britain & Ireland, is a small insectivorous passerine bird that belongs to the chat subfamily of the Old World flycatcher family. About in len ...
'') and dunnocks (''
Prunella modularis The dunnock (''Prunella modularis'') is a small passerine, or perching bird, found throughout temperate Europe and into Asian Russia. Dunnocks have also been successfully introduced into New Zealand. It is by far the most widespread member of th ...
'') in Britain, 31% of deaths were a result of cat predation. In parts of North America, the presence of larger carnivores such as coyotes which prey on cats and other small predators reduces the effect of predation by cats and other small predators such as opossums and
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight of ...
s on bird numbers and variety. Perhaps the best-known element of cats' hunting behavior, which is commonly misunderstood and often appalls cat owners because it looks like torture, is that cats often appear to "play" with prey by releasing and recapturing it. This
cat and mouse Cat and mouse, often expressed as cat-and-mouse game, is an English-language idiom that means "a contrived action involving constant pursuit, near captures, and repeated escapes." The "cat" is unable to secure a definitive victory over the "mouse ...
behavior is due to an instinctive imperative to ensure that the prey is weak enough to be killed without endangering the cat. Another poorly understood element of cat hunting behavior is the presentation of prey to human guardians. One explanation is that cats adopt humans into their social group and share excess kill with others in the group according to the
dominance hierarchy In biology, a dominance hierarchy (formerly and colloquially called a pecking order) is a type of social hierarchy that arises when members of animal social groups interact, creating a ranking system. A dominant higher-ranking individual is so ...
, in which humans are reacted to as if they are at or near the top. Another explanation is that they attempt to teach their guardians to hunt or to help their human as if feeding "an elderly cat, or an inept kitten". This hypothesis is inconsistent with the fact that male cats also bring home prey, despite males having negligible involvement in raising kittens.


Play

Domestic cats, especially young kittens, are known for their love of play. This behavior mimics hunting and is important in helping kittens learn to stalk, capture, and kill prey. Cats also engage in play fighting, with each other and with humans. This behavior may be a way for cats to practice the skills needed for real combat, and might also reduce any fear they associate with launching attacks on other animals. Cats also tend to play with toys more when they are hungry. Owing to the close similarity between play and hunting, cats prefer to play with objects that resemble prey, such as small furry toys that move rapidly, but rapidly lose interest. They become habituated to a toy they have played with before. String is often used as a toy, but if it is eaten, it can become caught at the base of the cat's tongue and then move into the intestines, a medical emergency which can cause serious illness, even death. Owing to the risks posed by cats eating string, it is sometimes replaced with a
laser pointer A laser pointer or laser pen is a small handheld device with a power source (usually a battery) and a laser diode emitting a very narrow coherent low-powered laser beam of visible light, intended to be used to highlight something of interest by ...
's dot, which cats may chase.


Reproduction

Female cats, called queens, are
polyestrous The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous ...
with several
estrus The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestrous ...
cycles during a year, lasting usually 21 days. They are usually ready to mate between early February and August. Several males, called tomcats, are attracted to a female in heat. They fight over her, and the victor wins the right to mate. At first, the female rejects the male, but eventually, the female allows the male to mate. The female utters a loud yowl as the male pulls out of her because a male cat's
penis A penis (plural ''penises'' or ''penes'' () is the primary sexual organ that male animals use to inseminate females (or hermaphrodites) during copulation. Such organs occur in many animals, both vertebrate and invertebrate, but males d ...
has a band of about 120–150 backward-pointing
penile spines Many mammalian species have developed keratinized penile spines along the glans and/or shaft, which may be involved in sexual selection. These spines have been described as being simple, single-pointed structures (macaques) or complex with two ...
, which are about long; upon withdrawal of the penis, the spines may provide the female with increased sexual stimulation, which acts to induce ovulation. After mating, the female cleans her
vulva The vulva (plural: vulvas or vulvae; derived from Latin for wrapper or covering) consists of the external female sex organs. The vulva includes the mons pubis (or mons veneris), labia majora, labia minora, clitoris, vestibular bulbs, vulv ...
thoroughly. If a male attempts to mate with her at this point, the female attacks him. After about 20 to 30 minutes, once the female is finished grooming, the cycle will repeat. Because ovulation is not always triggered by a single mating, females may not be impregnated by the first male with which they mate. Furthermore, cats are superfecund; that is, a female may mate with more than one male when she is in heat, with the result that different kittens in a litter may have different fathers. The
morula A morula (Latin, ''morus'': mulberry) is an early-stage embryo consisting of a solid ball of cells called blastomeres, contained in mammals, and other animals within the zona pellucida shell. The blastomeres are the daughter cells of the zygot ...
forms 124 hours after conception. At 148 hours, early blastocysts form. At 10–12 days, implantation occurs. The gestation of queens lasts between 64 and 67 days, with an average of 65 days. Data on the reproductive capacity of more than 2,300 free-ranging queens were collected during a study between May 1998 and October 2000. They had one to six kittens per
litter Litter consists of waste products that have been discarded incorrectly, without consent, at an unsuitable location. Litter can also be used as a verb; to litter means to drop and leave objects, often man-made, such as aluminum cans, paper cups ...
, with an average of three kittens. They produced a mean of 1.4 litters per year, but a maximum of three litters in a year. Of 169 kittens, 127 died before they were six months old due to a trauma caused in most cases by dog attacks and road accidents. The first litter is usually smaller than subsequent litters. Kittens are weaned between six and seven weeks of age. Queens normally reach sexual maturity at 5–10 months, and males at 5–7 months. This varies depending on breed. Kittens reach
puberty Puberty is the process of physical changes through which a child's body matures into an adult body capable of sexual reproduction. It is initiated by hormonal signals from the brain to the gonads: the ovaries in a girl, the testes in a bo ...
at the age of 9–10 months. Cats are ready to go to new homes at about 12 weeks of age, when they are ready to leave their mother. They can be surgically sterilized (spayed or
castrated Castration is any action, surgical, chemical, or otherwise, by which an individual loses use of the testicles: the male gonad. Surgical castration is bilateral orchiectomy (excision of both testicles), while chemical castration uses pharmac ...
) as early as seven weeks to limit unwanted reproduction. This surgery also prevents undesirable sex-related behavior, such as aggression, territory marking (spraying urine) in males and yowling (calling) in females. Traditionally, this surgery was performed at around six to nine months of age, but it is increasingly being performed before
puberty Puberty is the process of physical changes through which a child's body matures into an adult body capable of sexual reproduction. It is initiated by hormonal signals from the brain to the gonads: the ovaries in a girl, the testes in a bo ...
, at about three to six months. In the United States, about 80% of household cats are neutered.


Lifespan and health

The average lifespan of pet cats has risen in recent decades. In the early 1980s, it was about seven years, rising to 9.4 years in 1995 and about 15 years in 2021. Some cats have been reported as surviving into their 30s,Example: with the oldest known cat, Creme Puff, dying at a verified age of 38. Neutering increases life expectancy: one study found castrated male cats live twice as long as intact males, while spayed female cats live 62% longer than intact females. Having a cat
neutered Neutering, from the Latin ''neuter'' ('of neither sex'), is the removal of an animal's reproductive organ, either all of it or a considerably large part. The male-specific term is castration, while spaying is usually reserved for female animals. C ...
confers health benefits, because castrated males cannot develop
testicular cancer Testicular cancer is cancer that develops in the testicles, a part of the male reproductive system. Symptoms may include a lump in the testicle, or swelling or pain in the scrotum. Treatment may result in infertility. Risk factors include an ...
, spayed females cannot develop
uterine The uterus (from Latin ''uterus'', plural ''uteri'') or womb () is the organ in the reproductive system of most female mammals, including humans that accommodates the embryonic and fetal development of one or more embryos until birth. The ...
or
ovarian cancer Ovarian cancer is a cancerous tumor of an ovary. It may originate from the ovary itself or more commonly from communicating nearby structures such as fallopian tubes or the inner lining of the abdomen. The ovary is made up of three different c ...
, and both have a reduced risk of
mammary cancer A mammary tumor is a neoplasm originating in the mammary gland. It is a common finding in older female dogs and cats that are not spayed, but they are found in other animals as well. The mammary glands in dogs and cats are associated with their ...
.


Disease

About 250 heritable
genetic disorder A genetic disorder is a health problem caused by one or more abnormalities in the genome. It can be caused by a mutation in a single gene (monogenic) or multiple genes (polygenic) or by a chromosomal abnormality. Although polygenic disorders ...
s have been identified in cats, many similar to human
inborn errors of metabolism Inborn errors of metabolism form a large class of genetic diseases involving congenital disorders of enzyme activities. The majority are due to defects of single genes that code for enzymes that facilitate conversion of various substances ( substra ...
. The high level of similarity among the
metabolism Metabolism (, from el, μεταβολή ''metabolē'', "change") is the set of life-sustaining chemical reactions in organisms. The three main functions of metabolism are: the conversion of the energy in food to energy available to run c ...
of mammals allows many of these feline diseases to be diagnosed using
genetic tests Genetic testing, also known as DNA testing, is used to identify changes in DNA sequence or chromosome structure. Genetic testing can also include measuring the results of genetic changes, such as RNA analysis as an output of gene expression, o ...
that were originally developed for use in humans, as well as the use of cats as
animal model An animal model (short for animal disease model) is a living, non-human, often genetic-engineered animal used during the research and investigation of human disease, for the purpose of better understanding the disease process without the risk of ha ...
s in the study of the human diseases. Diseases affecting domestic cats include acute infections, parasitic infestations, injuries, and chronic diseases such as kidney disease,
thyroid disease Thyroid disease is a medical condition that affects the function of the thyroid gland. The thyroid gland is located at the front of the neck and produces thyroid hormones that travel through the blood to help regulate many other organs, meaning ...
, and arthritis.
Vaccinations Vaccination is the administration of a vaccine to help the immune system develop immunity from a disease. Vaccines contain a microorganism or virus in a weakened, live or killed state, or proteins or toxins from the organism. In stimulating ...
are available for many infectious diseases, as are treatments to eliminate parasites such as worms, ticks, and fleas.


Ecology


Habitats

The domestic cat is a
cosmopolitan species In biogeography, cosmopolitan distribution is the term for the range of a taxon that extends across all or most of the world in appropriate habitats. Such a taxon, usually a species, is said to exhibit cosmopolitanism or cosmopolitism. The extr ...
and occurs across much of the world. It is adaptable and now present on all continents except
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
, and on 118 of the 131 main groups of islands, even on the isolated Kerguelen Islands. Due to its ability to thrive in almost any terrestrial habitat, it is among the world's most invasive species. It lives on small islands with no human inhabitants. Feral cats can live in forests, grasslands, tundra, coastal areas, agricultural land, scrublands, urban areas, and wetlands. The unwantedness that leads to the domestic cat being treated as an invasive species is twofold. On one hand, as it is little altered from the wildcat, it can readily interbreed with the wildcat. This hybridization poses a danger to the genetic distinctiveness of some wildcat populations, particularly in
Scotland Scotland (, ) is a Countries of the United Kingdom, country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a Anglo-Scottish border, border with England to the southeast ...
and
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the ...
, possibly also the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
, and where protected natural areas are close to human-dominated landscapes, such as Kruger National Park in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...
. On the other hand, and perhaps more obviously, its introduction to places where no native felines are present contributes to the decline of native species.


Ferality

Feral cats are domestic cats that were born in or have reverted to a wild state. They are unfamiliar with and wary of humans and roam freely in urban and rural areas. The numbers of feral cats is not known, but estimates of the United States feral population range from 25 to 60 million. Feral cats may live alone, but most are found in large
colonies In modern parlance, a colony is a territory subject to a form of foreign rule. Though dominated by the foreign colonizers, colonies remain separate from the administration of the original country of the colonizers, the '' metropolitan state'' ...
, which occupy a specific territory and are usually associated with a source of food. Famous feral cat colonies are found in Rome around the
Colosseum The Colosseum ( ; it, Colosseo ) is an oval amphitheatre in the centre of the city of Rome, Italy, just east of the Roman Forum. It is the largest ancient amphitheatre ever built, and is still the largest standing amphitheatre in the world t ...
and Forum Romanum, with cats at some of these sites being fed and given medical attention by volunteers. Public attitudes toward feral cats vary widely, from seeing them as free-ranging pets to regarding them as vermin. Some feral cats can be successfully socialized and 're-tamed' for adoption; young cats, especially kittens and cats that have had prior experience and contact with humans are the most receptive to these efforts.


Impact on wildlife

On islands, birds can contribute as much as 60% of a cat's diet. In nearly all cases, the cat cannot be identified as the sole cause for reducing the numbers of island birds, and in some instances, eradication of cats has caused a "
mesopredator release The mesopredator release hypothesis is an ecological theory used to describe the interrelated population dynamics between apex predators and mesopredators within an ecosystem, such that a collapsing population of the former results in dramatically ...
" effect; where the suppression of top carnivores creates an abundance of smaller predators that cause a severe decline in their shared prey. Domestic cats are a contributing factor to the decline of many species, a factor that has ultimately led, in some cases, to extinction. The South Island piopio,
Chatham rail The Chatham rail (''Cabalus modestus'') is an extinct flightless species of bird in the family Rallidae. It was endemic to Chatham, Mangere and Pitt Islands, in the Chatham archipelago of New Zealand. The Chatham rail and the Dieffenbach's ...
, and the
New Zealand merganser The New Zealand merganser (''Mergus australis''), also known as Auckland merganser or Auckland Islands merganser, was a typical merganser which is now extinct. Description This duck was similar in size to the red-breasted merganser (''Mergus s ...
are a few from a long list, with the most extreme case being the flightless
Lyall's wren Lyall's wren or the Stephens Island wren (''Traversia lyalli'') is a small, extinct, flightless passerine belonging to the family Acanthisittidae, the New Zealand wrens. It was once found throughout New Zealand, but when it came to the attention ...
, which was driven to extinction only a few years after its discovery. One feral cat in New Zealand killed 102
New Zealand lesser short-tailed bat The New Zealand lesser short-tailed bat (''Mystacina tuberculata'') is a small-sized omnivorous mammal endemic to the islands of New Zealand. It is one of two extant and three overall terrestrial mammal species unique to New Zealand. The NZ lesse ...
s in seven days. In the US, feral and free-ranging domestic cats kill an estimated 6.3 – 22.3 billion mammals annually. In Australia, the impact of cats on mammal populations is even greater than the impact of habitat loss. More than one million reptiles are killed by feral cats each day, representing 258 species. Cats have contributed to the extinction of the Navassa curly-tailed lizard and '' Chioninia coctei.''


Interaction with humans

Cats are common
pet A pet, or companion animal, is an animal kept primarily for a person's company or entertainment rather than as a working animal, livestock, or a laboratory animal. Popular pets are often considered to have attractive appearances, intelligence ...
s throughout the world, and their worldwide population as of 2007 exceeded 500 million. Cats have been used for millennia to control rodents, notably around grain stores and aboard ships, and both uses extend to the present day. As well as being kept as pets, cats are also used in the international fur trade and leather industries for making coats, hats, blankets, and stuffed toys; and shoes, gloves, and musical instruments respectively (about 24 cats are needed to make a cat-fur coat). This use has been outlawed in the United States since 2000 and in the European Union (as well as the United Kingdom) since 2007. Cat pelts have been used for superstitious purposes as part of the practice of
witchcraft Witchcraft traditionally means the use of magic or supernatural powers to harm others. A practitioner is a witch. In medieval and early modern Europe, where the term originated, accused witches were usually women who were believed to have ...
, and are still made into blankets in Switzerland as
traditional medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
thought to cure rheumatism. A few attempts to build a cat census have been made over the years, both through associations or national and international organizations (such as that of the
Canadian Federation of Humane Societies Humane Canada is the largest national animal welfare organization in Canada, representing humane societies, SPCAs, and animal rescue organizations. Humane Canada aims to speak with a strong national voice, promoting the welfare and humane treat ...
) and over the Internet, but such a task does not seem simple to achieve. General estimates for the global population of domestic cats range widely from anywhere between 200 million to 600 million.
Walter Chandoha Walter George Chandoha (November 30, 1920 – January 11, 2019) was a prolific photographer, known especially for his photographs of animals and particularly of cats. Additional subjects for which he is known are fruits, vegetables, flowers, an ...
made his career photographing cats after his 1949 images of ''Loco'', an especially charming stray taken in, were published around the world. He is reported to have photographed 90,000 cats during his career and maintained an archive of 225,000 images that he drew from for publications during his lifetime.


Shows

A
cat show A cat show is a judged event where the owners of cats compete to win titles in various cat registering organizations by entering their cats to be judged after a breed standard. Both pedigreed and companion (or moggy) cats are admissible, althoug ...
is a judged event in which the owners of cats compete to win titles in various cat-registering organizations by entering their cats to be judged after a breed standard. It is often required that a cat must be healthy and vaccinated in order to participate in a cat show. Both
pedigreed Purebreds are "cultivated varieties" of an animal species achieved through the process of selective breeding. When the lineage of a purebred animal is recorded, that animal is said to be " pedigreed". Purebreds breed true-to-type which means the ...
and non-
purebred Purebreds are " cultivated varieties" of an animal species achieved through the process of selective breeding. When the lineage of a purebred animal is recorded, that animal is said to be "pedigreed". Purebreds breed true-to-type which means the ...
companion ("moggy") cats are admissible, although the rules differ depending on the organization. Competing cats are compared to the applicable breed standard, and assessed for temperament.


Infection

Cats can be infected or infested with
virus A virus is a submicroscopic infectious agent that replicates only inside the living cells of an organism. Viruses infect all life forms, from animals and plants to microorganisms, including bacteria and archaea. Since Dmitri Ivanovsk ...
es,
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of prokaryotic microorganisms. Typically a few micrometr ...
,
fungus A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from t ...
,
protozoans Protozoa (singular: protozoan or protozoon; alternative plural: protozoans) are a group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic tissues and debris. Histo ...
,
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
s or worms that can transmit diseases to humans. In some cases, the cat exhibits no symptoms of the disease. The same disease can then become evident in a human. The likelihood that a person will become diseased depends on the age and
immune In biology, immunity is the capability of multicellular organisms to resist harmful microorganisms. Immunity involves both specific and nonspecific components. The nonspecific components act as barriers or eliminators of a wide range of pathogens ...
status of the person. Humans who have cats living in their home or in close association are more likely to become infected. Others might also acquire infections from cat feces and
parasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has ...
s exiting the cat's body. Some of the infections of most concern include salmonella, cat-scratch disease and
toxoplasmosis Toxoplasmosis is a parasitic disease caused by '' Toxoplasma gondii'', an apicomplexan. Infections with toxoplasmosis are associated with a variety of neuropsychiatric and behavioral conditions. Occasionally, people may have a few weeks or mont ...
.


History and mythology

In ancient Egypt, cats were worshipped, and the goddess
Bastet Bastet or Bast ( egy, bꜣstjt, cop, Ⲟⲩⲃⲁⲥⲧⲉ, Oubaste , Phoenician: 𐤀𐤁𐤎𐤕, romanized: ’bst, or 𐤁𐤎𐤕, romanized: bst) was a goddess of ancient Egyptian religion, worshipped as early as the Second Dynasty (2 ...
often depicted in cat form, sometimes taking on the war-like aspect of a lioness. The Greek historian
Herodotus Herodotus ( ; grc, , }; BC) was an ancient Greek historian and geographer A geographer is a physical scientist, social scientist or humanist whose area of study is geography, the study of Earth's natural environment and human society ...
reported that killing a cat was forbidden, and when a household cat died, the entire family mourned and shaved their eyebrows. Families took their dead cats to the sacred city of
Bubastis Bubastis ( Bohairic Coptic: ''Poubasti''; Greek: ''Boubastis'' or ''Boubastos''), also known in Arabic as Tell-Basta or in Egyptian as Per-Bast, was an ancient Egyptian city. Bubastis is often identified with the biblical ''Pi-Beseth'' ( h ...
, where they were embalmed and buried in sacred repositories. Herodotus expressed astonishment at the domestic cats in Egypt, because he had only ever seen wildcats. Ancient Greeks and Romans kept
weasel Weasels are mammals of the genus ''Mustela'' of the family Mustelidae. The genus ''Mustela'' includes the least weasels, polecats, stoats, ferrets and European mink. Members of this genus are small, active predators, with long and slender b ...
s as pets, which were seen as the ideal rodent-killers. The earliest unmistakable evidence of the Greeks having domestic cats comes from two coins from Magna Graecia dating to the mid-fifth century BC showing Iokastos and Phalanthos, the legendary founders of
Rhegion Reggio di Calabria ( scn, label= Southern Calabrian, Riggiu; el, label=Calabrian Greek, Ρήγι, Rìji), usually referred to as Reggio Calabria, or simply Reggio by its inhabitants, is the largest city in Calabria. It has an estimated popula ...
and Taras respectively, playing with their pet cats. The usual ancient Greek word for 'cat' was , meaning 'thing with the waving tail'. Cats are rarely mentioned in
ancient Greek literature Ancient Greek literature is literature written in the Ancient Greek language from the earliest texts until the time of the Byzantine Empire. The earliest surviving works of ancient Greek literature, dating back to the early Archaic period, are ...
.
Aristotle Aristotle (; grc-gre, Ἀριστοτέλης ''Aristotélēs'', ; 384–322 BC) was a Greek philosopher and polymath during the Classical period in Ancient Greece. Taught by Plato, he was the founder of the Peripatetic school of ph ...
remarked in his ''
History of Animals ''History of Animals'' ( grc-gre, Τῶν περὶ τὰ ζῷα ἱστοριῶν, ''Ton peri ta zoia historion'', "Inquiries on Animals"; la, Historia Animalium, "History of Animals") is one of the major texts on biology by the ancient Gr ...
'' that "female cats are naturally
lecherous Lascivious behavior is sexual behavior or conduct that is considered crude and offensive, or contrary to local moral or other standards of appropriate behavior. In this sense "lascivious" is similar in meaning to "lewd", "indecent", "lecherous", ...
." The Greeks later syncretized their own goddess
Artemis In ancient Greek mythology and religion, Artemis (; grc-gre, Ἄρτεμις) is the goddess of the hunt, the wilderness, wild animals, nature, vegetation, childbirth, care of children, and chastity. She was heavily identified wit ...
with the Egyptian goddess Bastet, adopting Bastet's associations with cats and ascribing them to Artemis. In
Ovid Pūblius Ovidius Nāsō (; 20 March 43 BC – 17/18 AD), known in English as Ovid ( ), was a Roman poet who lived during the reign of Augustus. He was a contemporary of the older Virgil and Horace, with whom he is often ranked as one of the th ...
's ''
Metamorphoses The ''Metamorphoses'' ( la, Metamorphōsēs, from grc, μεταμορφώσεις: "Transformations") is a Latin narrative poem from 8 CE by the Roman poet Ovid. It is considered his ''magnum opus''. The poem chronicles the history of the ...
'', when the deities flee to Egypt and take animal forms, the goddess Diana turns into a cat. Cats eventually displaced
weasel Weasels are mammals of the genus ''Mustela'' of the family Mustelidae. The genus ''Mustela'' includes the least weasels, polecats, stoats, ferrets and European mink. Members of this genus are small, active predators, with long and slender b ...
s as the pest control of choice because they were more pleasant to have around the house and were more enthusiastic hunters of mice. During the
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the late 5th to the late 15th centuries, similar to the post-classical period of global history. It began with the fall of the Western Roman Empire ...
, many of Artemis's associations with cats were grafted onto the
Virgin Mary Mary; arc, ܡܪܝܡ, translit=Mariam; ar, مريم, translit=Maryam; grc, Μαρία, translit=María; la, Maria; cop, Ⲙⲁⲣⲓⲁ, translit=Maria was a first-century Jewish woman of Nazareth, the wife of Joseph and the mother of ...
. Cats are often shown in icons of Annunciation and of the
Holy Family The Holy Family consists of the Child Jesus, the Virgin Mary and Saint Joseph. The subject became popular in art from the 1490s on, but veneration of the Holy Family was formally begun in the 17th century by Saint François de Laval, the fir ...
and, according to
Italian folklore Folklore of Italy refers to the folklore and urban legends of Italy. Within the Italian territory, various peoples have followed one another over time, each of which has left its mark on current culture. Some tales also come from Christianizat ...
, on the same night that Mary gave birth to
Jesus Jesus, likely from he, יֵשׁוּעַ, translit=Yēšūaʿ, label= Hebrew/ Aramaic ( AD 30 or 33), also referred to as Jesus Christ or Jesus of Nazareth (among other names and titles), was a first-century Jewish preacher and religiou ...
, a cat in
Bethlehem Bethlehem (; ar, بيت لحم ; he, בֵּית לֶחֶם '' '') is a city in the central West Bank, Palestine, about south of Jerusalem. Its population is approximately 25,000,Amara, 1999p. 18.Brynen, 2000p. 202. and it is the capital o ...
gave birth to a kitten. Domestic cats were spread throughout much of the rest of the world during the
Age of Discovery The Age of Discovery (or the Age of Exploration), also known as the early modern period, was a period largely overlapping with the Age of Sail, approximately from the 15th century to the 17th century in European history, during which seafarin ...
, as ships' cats were carried on
sailing ship A sailing ship is a sea-going vessel that uses sails mounted on masts to harness the power of wind and propel the vessel. There is a variety of sail plans that propel sailing ships, employing square-rigged or fore-and-aft sails. Some ships ...
s to control shipboard rodents and as good-luck charms. Several ancient religions believed cats are exalted souls, companions or guides for humans, that are all-knowing but mute so they cannot influence decisions made by humans. In Japan, the cat is a symbol of good fortune. In Norse mythology,
Freyja In Norse paganism, Freyja (Old Norse "(the) Lady") is a goddess associated with love, beauty, fertility, sex, war, gold, and seiðr (magic for seeing and influencing the future). Freyja is the owner of the necklace Brísingamen, rides a chario ...
, the goddess of love, beauty, and fertility, is depicted as riding a chariot drawn by cats. In Jewish legend, the first cat was living in the house of the first man Adam as a pet that got rid of mice. The cat was once partnering with the first dog before the latter broke an oath they had made which resulted in enmity between the descendants of these two animals. It is also written that neither cats nor
foxes Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...
are represented in the water, while every other animal has an incarnation species in the water. Although no species are sacred in Islam, cats are revered by Muslims. Some Western writers have stated
Muhammad Muhammad ( ar, مُحَمَّد;  570 – 8 June 632 CE) was an Arab religious, social, and political leader and the founder of Islam. According to Islamic doctrine, he was a prophet divinely inspired to preach and confirm the mo ...
had a favorite cat, ''
Muezza The domestic cat is a revered animal in Islam. Admired for their cleanliness, cats are considered "the quintessential pet" by Muslims. Origins of reverence Cats have been venerated in the Near East since antiquity. Islam also has that trad ...
''. He is reported to have loved cats so much, "he would do without his cloak rather than disturb one that was sleeping on it". The story has no origin in early Muslim writers, and seems to confuse a story of a later Sufi saint,
Ahmed ar-Rifa'i Ahmad ( ar, أحمد, ʾAḥmad) is an Arabic male given name common in most parts of the Muslim world. Other spellings of the name include Ahmed and Ahmet. Etymology The word derives from the root (ḥ-m-d), from the Arabic (), from the ve ...
, centuries after Muhammad. One of the companions of Muhammad was known as
Abu Hurayrah Abu Hurayra ( ar, أبو هريرة, translit=Abū Hurayra; –681) was one of the companions of Islamic prophet Muhammad and, according to Sunni Islam, the most prolific narrator of hadith. He was known by the ''kunyah'' Abu Hurayrah "Fathe ...
("father of the kitten"), in reference to his documented affection to cats.


Superstitions and rituals

Many cultures have negative superstitions about cats. An example would be the belief that encountering a black cat ("crossing one's path") leads to bad luck, or that cats are witches'
familiars In European folklore of the medieval and early modern periods, familiars (sometimes referred to as familiar spirits) were believed to be supernatural entities that would assist witches and cunning folk in their practice of magic. According to r ...
used to augment a witch's powers and skills. The killing of cats in Medieval
Ypres Ypres ( , ; nl, Ieper ; vls, Yper; german: Ypern ) is a Belgian city and municipality in the province of West Flanders. Though the Dutch name is the official one, the city's French name is most commonly used in English. The municipality c ...
,
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...
, is commemorated in the innocuous present-day
Kattenstoet The ''Kattenstoet'' (lit. "Festival of the Cats") is a parade in Ypres, Belgium, devoted to the cat. It has been held regularly on the second Sunday of May since 1955. Most recently, the 45th edition took place on 13 May 2018, with the next sch ...
(cat parade). In mid-16th century France, cats would be burnt alive as a form of entertainment. According to Norman Davies, the assembled people "shrieked with laughter as the animals, howling with pain, were singed, roasted, and finally
carbonized Carbonized were a Sweden, Swedish avant-garde metal band formed 1988 in Saltsjöbaden. The band was formed by Lars Rosenberg in 1988, with Dismember (band), Dismember vocalist Matti Kärki. Joined by drummer Piotr Wawrzeniuk, the trio was compl ...
".
James Frazer Sir James George Frazer (; 1 January 1854 – 7 May 1941) was a Scottish social anthropologist and folklorist influential in the early stages of the modern studies of mythology and comparative religion. Personal life He was born on 1 Janua ...
wrote that "It was the custom to burn a basket, barrel, or sack full of live cats, which was hung from a tall mast in the midst of the bonfire; sometimes a
fox Foxes are small to medium-sized, omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull, upright, triangular ears, a pointed, slightly upturned snout, and a long bushy tail (or ''brush''). Twelve sp ...
was burned. The people collected the embers and ashes of the fire and took them home, believing that they brought good luck. The French kings often witnessed these spectacles and even lit the bonfire with their own hands. In 1648
Louis XIV , house = Bourbon , father = Louis XIII , mother = Anne of Austria , birth_date = , birth_place = Château de Saint-Germain-en-Laye, Saint-Germain-en-Laye, France , death_date = , death_place = Palace of Ver ...
, crowned with a wreath of roses and carrying a bunch of roses in his hand, kindled the fire, danced at it and partook of the banquet afterwards in the town hall. But this was the last occasion when a monarch presided at the midsummer bonfire in Paris. At
Metz Metz ( , , lat, Divodurum Mediomatricorum, then ) is a city in northeast France located at the confluence of the Moselle and the Seille rivers. Metz is the prefecture of the Moselle department and the seat of the parliament of the Grand ...
midsummer fires were lighted with great pomp on the esplanade, and a dozen cats, enclosed in wicker cages, were burned alive in them, to the amusement of the people. Similarly, at Gap, in the department of the
Hautes-Alpes Hautes-Alpes (; oc, Auts Aups; en, Upper Alps) is a department in the Provence-Alpes-Côte d'Azur region of Southeastern France. It is located in the heart of the French Alps, after which it is named. Hautes-Alpes had a population of 141,220 ...
, cats used to be roasted over the midsummer bonfire." According to a myth in many cultures, cats have multiple lives. In many countries, they are believed to have nine lives, but in Italy, Germany, Greece, Brazil and some Spanish-speaking regions, they are said to have seven lives, while in Arabic traditions, the number of lives is six. The myth is attributed to the natural suppleness and swiftness cats exhibit to escape life-threatening situations. Also lending credence to this myth is the fact that falling cats often land on their feet, using an instinctive
righting reflex The righting reflex, also known as the labyrinthine righting reflex, is a reflex that corrects the orientation of the body when it is taken out of its normal upright position. It is initiated by the vestibular system, which detects that the body is ...
to twist their bodies around. Nonetheless, cats can still be injured or killed by a high fall. (Press release.)


Gallery

File:Persian Baby Cat.jpg, alt=Baby Persian Cat, A
Persian Persian may refer to: * People and things from Iran, historically called ''Persia'' in the English language ** Persians, the majority ethnic group in Iran, not to be conflated with the Iranic peoples ** Persian language, an Iranian language of the ...
kitten.


See also

*
Aging in cats Aging in cats is the process by which cats change over the course of their natural lifespans. As cats age, they undergo changes in health and behavior. Dental disease and loss of olfaction are common as cats age, affecting eating habits. Arthriti ...
*
Ailurophobia Ailurophobia (''pronunciation:'' aɪˌlʊər əˈfoʊ bi ə) is the persistent and excessive fear or hatred of cats. Like other specific phobias, the exact cause of ailurophobia is unknown, and potential treatment generally involves therapy.Milos ...
* Animal testing on cats *
Animal track __notoc__ An animal track is an imprint left behind in soil, snow, or mud, or on some other ground surface, by an animal walking across it. Animal tracks are used by hunters in tracking their prey and by naturalists to identify animals living ...
* Cancer in cats *
Cat bite Cat bites are bites inflicted upon humans, other cats, and other animals by the domestic cat (). Data from the United States show that cat bites represent between 5–15% of all animal bites inflicted to humans, but it has been argued that this fig ...
*
Cat café A cat café is a theme café whose attraction is cats who can be watched and played with. Patrons pay a cover fee, generally hourly, and thus cat cafés can be seen as a form of supervised indoor pet rental. "Cat café" has been officially reco ...
*
Cat collar A cat collar is a piece of material put around the neck of a cat. Cat collars are used for identification, fashion, protection (as from fleas), restraint, or to warn off prey, and may be worn by cats that are indoor-only as well as cats with outdoo ...
* Cat lady *
Cat lover culture Cat culture describes the culture that surrounds cat lovers. Cat fancy is a hobby involving the appreciation, promotion, or breeding of cats. For some, cats can become an obsession. Some refer to themselves as "cat people". Description One as ...
*
Cat meat The cat (''Felis catus'') is a domestic species of small carnivorous mammal. It is the only domesticated species in the family Felidae and is commonly referred to as the domestic cat or house cat to distinguish it from the wild members o ...
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Cats and the Internet Images and videos of domestic cats make up some of the most viewed content on the World Wide Web, particularly image macros in the form of lolcats. ThoughtCatalog has described cats as the "unofficial mascot of the Internet". The subject has ...
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Cats in Australia Cats are an invasive species in Australia. Because they are not native to Australia and were only introduced by colonisers as pets in the early 1800s, native Australian animals did not co-evolve with them. , some 3.8 million domestic cats and u ...
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Cats in New Zealand Cats are a popular pet in New Zealand. Cat ownership is occasionally raised as a controversial conservation issue due to the predation of endangered species, such as birds and lizards, by feral cats. Domesticated cats The domestic cat (''Fe ...
* Cats in the United States * Dog–cat relationship *
Dried cat In some European cultures it was customary to place the dried or desiccated body of a cat inside the walls of a newly built home to ward off evil spirits or as a good luck charm. It was believed that the cats had a sixth sense and that putting a ...
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List of cat breeds The following list of cat breeds includes only domestic cat breeds and domestic and wild hybrids. The list includes established breeds recognized by various cat registries, new and experimental breeds, landraces being established as standardized b ...
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List of cat documentaries List of cat documentaries, television series and cartoons includes ''serious'' documentaries, television series and cartoons, in alphabetical order, related to cats . Documentaries # ''Cats: Caressing the Tiger'' (1991), National Geographic # ''C ...
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List of individual cats This is a list of famous cats which achieved some degree of popularity either in their own right or by association with someone famous. Before the modern era * Nedjem or Nojem ( Egyptian: ''nḏm'' "Sweet One" or "Sweetie"), 15th century BC. The ...
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List of fictional cats and felines This is a list of fictional cats and felines and is a subsidiary to the list of fictional animals. It includes a limited selection of notable felines from various works, organized by medium. More complete lists are accessible by clicking on the "Ma ...
* Perlorian *
Pet door A pet door or pet flap (also referred to in more specific terms, such as cat flap, cat door, kitty door, dog flap, dog door, or doggy/doggie door) is a small opening to allow pets to enter and exit a building on their own without needing a human t ...
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Pet first aid Pet first aid refers to emergency treatment administered to an injured or sick domestic animal before professional medical care is available. Much of the first aid First aid is the first and immediate assistance given to any person with eith ...
* Popular cat names


References


External links

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High-Resolution Images of the Cat Brain

Biodiversity Heritage Library bibliography
for ''Felis catus''
Catpert. The Cat Expert
nbsp;– Cat articles * View th
cat genome
in
Ensembl Ensembl genome database project is a scientific project at the European Bioinformatics Institute, which provides a centralized resource for geneticists, molecular biologists and other researchers studying the genomes of our own species and other v ...
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Scientific American ''Scientific American'', informally abbreviated ''SciAm'' or sometimes ''SA'', is an American popular science magazine. Many famous scientists, including Albert Einstein and Nikola Tesla, have contributed articles to it. In print since 1845, it ...
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The Origin of the Cat
, 20 August 1881, pp. 120 {{Authority control Mammals described in 1758 Animal models Articles containing video clips Felis Taxa named by Carl Linnaeus Cosmopolitan mammals