Dlusskyidris
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''Dlusskyidris'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus of ant in the Formicidae subfamily Sphecomyrminae, and is one of the five genera placed in the tribe Sphecomyrmini. The genus contains a single described species, ''Dlusskyidris zherichini'', and is known from three Late Cretaceous fossils which have been found in
northern Russia Russian North (russian: Русский Север) is an ethnocultural region situated in the northwestern part of Russia. It spans the regions of Arkhangelsk Oblast, the Republic of Karelia, Komi Republic, Vologda Oblast and Nenets Autonomo ...
.


History and classification

The three males of ''Dlusskyidris'' have been identified from adults that are preserved as inclusions in Taymyr amber. The ambers of the peninsula occur in the upper levels of the
Kheta Formation Kheta may refer to: * Kheta (river), a river in Russia * , a settlement in Taymyrsky Dolgano-Nenetsky District, Russia * Kheta of Mewar, 14th-century Indian ruler * Kheta Ram Kheta Ram aka Khetha Ram / KhetaRam (born 20 September 1986) is an I ...
, which is exposed in a number of locations in the Taimyr region. Age estimates of the Kheta Formation are between the
Coniacian The Coniacian is an age or stage in the geologic timescale. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series and spans the time between 89.8 ± 1 Ma and 86.3 ± 0.7 Ma (million years ago). The Coniacian is preceded by t ...
and Santonian, and the ambers are found consistently in the uppermost units, giving a Santonian age range for the inclusions. The ''Dlusskyidris'' adults were collected from the Yantardakh locality, which is approximately upstream from the confluence of the
Maimecha River The Maymecha, also known as Medvezhye (russian: Маймеча, Медвежье), is a river in Taymyrsky Dolgano-Nenetsky District of Krasnoyarsk Krai in Russia, right tributary of the Kheta ( Khatanga basin). The length of the river is . The ar ...
with the Kheta River, on the bank of the Maimecha. Based on the flora and fauna of the Ledyanaya and Mutino formations which surround the Kheta Formation, the paleoforest likely has a humid and warm temperate climate with the trees growing along river banks. While the resin producing trees have not been identified, the resins were likely dropped into the river systems and buried quickly in deltaic sediments. The three males were first studied by Russian paleoentomologist
Gennady Dlussky Gennady ( rus, Геннадий, p=ɡʲɪˈnadʲɪj), also spelled Gennadi or Gennadiy, is a Russian male name. They are derived from the Greek given name Gennadius. People *Gennady Gladkov, Soviet and Russian composer *Gennady Golovkin, Kazakh bo ...
, who published the genus and species description in a 1975 paper. In that paper Dlussky named the genus "''Palaeomyrmex''" and the species "''Palaeomyrmex zherichini''". However the genus name ''Palaeomyrmex'' had already been used over a hundred years earlier by Oswald Heer for '' Palaeomyrmex prodromus''. Due to the name being
preoccupied The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linn ...
, the species was moved to the new genus name ''Dlusskyidris'' in 1994 by
Barry Bolton Barry Bolton is an English Myrmecology, myrmecologist, an expert on the classification, systematics, and taxonomy of ants, who long worked at the Natural History Museum, London. He is known especially for monographs on African and Asian ants, and ...
. Russian paleoentomologist K. S. Perfilieva re-examined the holotype and paratype fossils 2011. Perfilieva noted that the visibility of the wings and the refractive index of the amber had been altered due to years of storage in castor oil. The oil had seeped into the amber, filling in air bubbles and making the wings nearly invisible under light microscope examination.


Description

The ''Dlusskyidris'' males are distinguished from other Sphecomyrminae males based on several features. There are spurs present on the tibias of the hind and mid legs. The tergite of the end abdominal segment is triangular in shape, as is the subgenital plate. There are well developed cerci on the abdomen, the
paramere Parameres ('side parts') are part of the external reproductive organs of male insects and the term was first used by Karl Wilhelm Verhoeff, Verhoeff in 1893 for the lateral genital lobes in Coleoptera. The primary phallic lobes which appear in th ...
s are narrow with slight curving in the middle and pointed tips. In Dlussky's original description, he considered the wing venation primitive and similar to that of Armaniidae genera with complete cells 1r+2r, 3r and 2rm. Perfilieva noted however that two different versions of the venation were presented by Dlussky.


References

{{Taxonbar, from=Q18102432 Sphecomyrminae Cretaceous insects Fossil ant genera Fossil taxa described in 1975 Fossil taxa described in 1994 Cretaceous insects of Asia Taimyr amber