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Dinosauromorpha is a
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
of
avemetatarsalian Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Dinos ...
archosaur Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian d ...
s (reptiles closer to birds than to crocodilians) that includes the Dinosauria (dinosaurs) and some of their close relatives. It was originally defined to include dinosauriforms and lagerpetids, with later formulations specifically excluding pterosaurs from the group.
Bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweigh ...
s are the only dinosauromorphs which survive to the present day.


Classification

The name "Dinosauromorpha" was briefly coined by Michael J. Benton in 1985. It was considered an alternative name for the group "Ornithosuchia", which was named by Jacques Gauthier to correspond to archosaurs closer to dinosaurs than to crocodilians. Although "Ornithosuchia" was later recognized as a
misnomer A misnomer is a name that is incorrectly or unsuitably applied. Misnomers often arise because something was named long before its correct nature was known, or because an earlier form of something has been replaced by a later form to which the name ...
(since
ornithosuchids Ornithosuchidae is an extinct family of pseudosuchian archosaurs (distant relatives of modern crocodilians) from the Triassic period. Ornithosuchids were quadrupedal and facultatively bipedal (e.g. like chimpanzees), meaning that they had the a ...
are now considered closer to crocodilians than to dinosaurs), it was still a more popular term than Dinosauromorpha in the 1980s. The group encompassed by Gauthier's "Ornithosuchia" and Benton's "Dinosauromorpha" is now given the name
Avemetatarsalia Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. ...
. In 1991,
Paul Sereno Paul Callistus Sereno (born October 11, 1957) is a professor of paleontology at the University of Chicago and a National Geographic "explorer-in-residence" who has discovered several new dinosaur species on several continents, including at sites ...
redefined Dinosauromorpha as a node-based
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
, defined by a
last common ancestor In biology and genetic genealogy, the most recent common ancestor (MRCA), also known as the last common ancestor (LCA) or concestor, of a set of organisms is the most recent individual from which all the organisms of the set are descended. The ...
and its descendants. In his definition, Dinosauromorpha included the
last common ancestor In biology and genetic genealogy, the most recent common ancestor (MRCA), also known as the last common ancestor (LCA) or concestor, of a set of organisms is the most recent individual from which all the organisms of the set are descended. The ...
of ''
Lagerpeton ''Lagerpeton'' is a genus of lagerpetid avemetatarsalian, comprising a single species, ''L. chanarensis''. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, ''Lagerpeton'' anatomy is somewhat incompletely kn ...
'' (a lagerpetid), ''
Marasuchus ''Marasuchus'' (meaning "Mara crocodile") is a genus of basal dinosauriform archosaur which is possibly synonymous with '' Lagosuchus''. Both genera lived during the Late Triassic in what is now La Rioja Province, Argentina. ''Marasuchus'' conta ...
'' (a possible
junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...
of ''
Lagosuchus ''Lagosuchus'' is an extinct genus of avemetatarsalian archosaur from the Late Triassic of Argentina. The type species of ''Lagosuchus'', ''Lagosuchus talampayensis,'' is based on a small partial skeleton recovered from the early Carnian-age Cha ...
),'' ''
Pseudolagosuchus ''Lewisuchus'' is a genus of archosaur that lived during the Late Triassic (early Carnian). As a silesaurid dinosauriform, it was a member of the group of reptiles most commonly considered to be the closest relatives of dinosaurs (possibly tr ...
'' (now considered a synonym of the
silesaurid Silesauridae is an extinct family of Triassic dinosauriforms. It is most commonly considered to be a clade of non-dinosaur dinosauriforms, and the sister group of dinosaurs. Some studies have instead suggested that most or all silesaurids compri ...
'' Lewisuchus''), Dinosauria (including Aves), and all its descendants. This definition was intended to correspond to a clade including lagerpetids and crownward bird-line archosaurs, but not pterosaurs or other archosaurs. In 2011, Dinosauromorpha was redefined by
Sterling Nesbitt Sterling Nesbitt (born March 25, 1982, in Mesa, Arizona) is an American paleontologist best known for his work on the origin and early evolutionary patterns of archosaurs. He is currently an associate professor at Virginia Tech in the Department of ...
to be a branch-based clade, defined by including reptiles closer to one group than to another. Under this definition, Dinosauromorpha included all reptiles closer to dinosaurs (represented by ''
Passer domesticus The house sparrow (''Passer domesticus'') is a bird of the sparrow family Passeridae, found in most parts of the world. It is a small bird that has a typical length of and a mass of . Females and young birds are coloured pale brown and grey, an ...
'', the house sparrow), rather than pterosaurs (represented by ''
Pterodactylus ''Pterodactylus'' (from Greek () meaning 'winged finger') is an extinct genus of pterosaurs. It is thought to contain only a single species, ''Pterodactylus antiquus'', which was the first pterosaur to be named and identified as a flying rept ...
''), ornithosuchids (represented by ''
Ornithosuchus ''Ornithosuchus'' (meaning "bird crocodile") is an extinct genus of pseudosuchians from the Late Triassic (Carnian) Lossiemouth Sandstone of Scotland. It was originally thought to be the ancestor to the carnosaurian dinosaurs (such as ''Allosau ...
''), or other
pseudosuchians Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Prior to ...
(represented by ''
Crocodylus niloticus ''Crocodylus'' is a genus of true crocodiles in the family Crocodylidae. Taxonomy The generic name, ''Crocodylus'', was proposed by Josephus Nicolaus Laurenti in 1768. ''Crocodylus'' contains 13–14 extant (living) species and 5 extinct spe ...
'', the Nile crocodile). Nesbitt's study supported the hypothesis that Pterosauromorpha (pterosaurs and their potential relatives) was the sister group of Dinosauromorpha. Pterosauromorphs and dinosauromorphs together formed the group
Ornithodira Avemetatarsalia (meaning "bird metatarsals") is a clade of diapsid reptiles containing all archosaurs more closely related to birds than to crocodilians. The two most successful groups of avemetatarsalians were the dinosaurs and pterosaurs. Dinos ...
, which encompasses almost all avemetatarsalians. Dinosauriformes was coined in 1992 by F.E. Novas, who used it to encompass dinosaurs, ''Lagosuchus'', "''Pseudolagosuchus''" (=''Lewisuchus''), and the herrerasaurids, which he did not consider to be "eudinosaurs" (true dinosaurs like ornithischians and
saurischia Saurischia ( , meaning "reptile-hipped" from the Greek ' () meaning 'lizard' and ' () meaning 'hip joint') is one of the two basic divisions of dinosaurs (the other being Ornithischia), classified by their hip structure. Saurischia and Ornithis ...
ns). Contrary to Novas, most paleontologists since 1992 have considered herrerasaurids to be true dinosaurs, though many other dinosaur-like reptiles still fall within his definition of Dinosauriformes. Novas (1992) defined Dinosauriformes as a node-based clade containing the most recent common ancestor of ''Lagosuchus'' and Dinosauria, and all its descendants. Nesbitt (2011) provided a roughly equivalent definition, using ''Marasuchus'' and ''Passer domesticus'' (the house sparrow, a representative of dinosaurs). In his analysis, Dinosauriformes included dinosaurs, silesaurids, and ''Marasuchus'', but not lagerpetids, which were considered to be an earlier-branching family of dinosauromorphs.


Phylogeny

A phylogenetic analysis by Andrea Cau in 2018 resolved two different topologies for dinosaur origins, depending on whether it was run using parsimony or
bayesian inference Bayesian inference is a method of statistical inference in which Bayes' theorem is used to update the probability for a hypothesis as more evidence or information becomes available. Bayesian inference is an important technique in statistics, a ...
. Cau coined the term Dracohors for the clade uniting all taxa closer to the theropod '' Megalosaurus bucklandi'' than the basal form '' Marasuchus lilloensis''. The name is derived from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
words for "dragon" and "cohort", ''dracos'' and ''cohors''. Under parsimony results, Dracohors included only Silesauridae and Dinosauria, the latter including the groups
Herrerasauria Saurischia ( , meaning "reptile-hipped" from the Greek ' () meaning 'lizard' and ' () meaning 'hip joint') is one of the two basic divisions of dinosaurs (the other being Ornithischia), classified by their hip structure. Saurischia and Ornithis ...
,
Sauropodomorpha Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lon ...
,
Theropoda Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ca ...
and Ornithischia, along with the basal form ''
Eodromaeus ''Eodromaeus'' (meaning "dawn runner") is an extinct genus of probable basal theropod dinosaurs from the Late Triassic of Argentina. Like many other of the earliest-known dinosaurs, it hails from the Carnian-age (~230 Ma) Ischigualasto Form ...
''. However, under bayesian results, Herrerasauria placed outside Dinosauria within Dracohors, and Dinosauriformes, Dinosauromorpha, and Pan-Aves were synonyms, with ''Marasuchus'' in a clade with lagerpetids. ''Pisanosaurus'' was resolved within Silesauridae. Cau identified the synapomorphies of Dracohors as:
The anterior tympanic recess, the axial epipophyses, the centrodiapophyseal laminae in the presacral vertebrae, the relative size enlargement of the postacetabular process of ilium, the elongation of the pubis, the proximal sulcus and the reduction of the ligament tuber in the femoral head, and the further reduction in length of the fourth metatarsal and toe compared to the third.
Following the discovery and description of more cranial and postcranial material of the genera '' Kongonaphon'', ''
Ixalerpeton ''Ixalerpeton'' (meaning "leaping reptile") is a genus of Lagerpetidae, lagerpetid Avemetatarsalia, avemetatarsalian containing one species, ''I. polesinensis''. It lived in the Late Triassic of Brazil alongside the sauropodomorpha, sauropodomorp ...
'' and ''
Lagerpeton ''Lagerpeton'' is a genus of lagerpetid avemetatarsalian, comprising a single species, ''L. chanarensis''. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, ''Lagerpeton'' anatomy is somewhat incompletely kn ...
'', it was found that lagerpetids shared many features with the basal taxa of Pterosauria. Features of the
maxillary bone The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The t ...
, teeth,
braincase In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skul ...
and forelimb meant that the
2020 2020 was heavily defined by the COVID-19 pandemic, which led to global Social impact of the COVID-19 pandemic, social and Economic impact of the COVID-19 pandemic, economic disruption, mass cancellations and postponements of events, COVID- ...
phylogenetic analysis of Ezcurra and colleagues placed Lagerpetidae next to pterosaurs within Pterosauromorpha, removing the family from Dinosauromorpha. The contents of Dinosauromorpha was thus restricted to only Silesauridae, Dinosauria, and individual genera like ''
Lagosuchus ''Lagosuchus'' is an extinct genus of avemetatarsalian archosaur from the Late Triassic of Argentina. The type species of ''Lagosuchus'', ''Lagosuchus talampayensis,'' is based on a small partial skeleton recovered from the early Carnian-age Cha ...
''. Simultaneously, Rodrigo Müller and Maurício Garcia published novel results that reduced the family Silesauridae to a grade of basal dinosaurs in Ornithischia. ''
Pisanosaurus ''Pisanosaurus'' () is an extinct genus of early dinosauriform, likely an ornithischian or silesaurid, from the Late Triassic of Argentina. It was a small, lightly built, ground-dwelling herbivore, that could grow up to an estimated long. Onl ...
'', considered by various authors to be either a silesaurid or basal ornithischian, was found to be intermediate between the grade of silesaurids and true ornithischians, explaining its peculiar combination of silesaurid and ornithischian features that has resulted in its phylogenetic inconsistency. '' Lewisuchus'', a carnivorous form, was found to be the most primitive form of ornithischian, which was almost universally considered to be an only-herbivorous clade before. Dinosauromorpha was reduced to only including Lagerpetidae and ''Lagosuchus'' as a result of the reclassification of silesaurids. Below are the results of: Cau (2018, parsimony results): Cau (2018, bayesian results): Ezcurra ''et al.'' (2020): Müller and Garcia (2020): A variety of individual species and taxa have at times been found to place within Dinosauromorpha and its subgroups, but outside Dinosauria. The taxon ''Marasuchus'' has been consistently recovered as a dinosauromorph between lagerpetids and silesaurids, but may also be a junior synonym of the coexisting form ''
Lagosuchus ''Lagosuchus'' is an extinct genus of avemetatarsalian archosaur from the Late Triassic of Argentina. The type species of ''Lagosuchus'', ''Lagosuchus talampayensis,'' is based on a small partial skeleton recovered from the early Carnian-age Cha ...
'', another dinosauromorph. ''Pisanosaurus'', traditionally considered an ornithischian, was recovered in an unpublished analysis as a dinosauriform outside other clades, but has since been recovered only as a member of Silesauridae or Ornithischia. ''
Saltopus ''Saltopus'' ("hopping foot") is a genus of very small bipedal dinosauriform containing the single species ''Saltopus elginensis'' from the late Triassic period of Scotland. It is one of the most famous Elgin Reptiles. Description ''Saltopus ...
'', an enigmatic taxon from the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
of
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to the ...
, has been placed closer to dinosaurs than ''Marasuchus'', in a polytomy with Silesauridae and Dinosauria, as a sister taxon to ''Marasuchus'', or within Dinosauria as a basal saurischian. The British taxon ''
Agnosphitys ''Agnosphitys'' (; "unknown begetter"; sometimes mistakenly called ''Agnostiphys'' or ''Agnosphytis'') is a genus of dinosauriform that lived during the Late Triassic. It contains only one species, the type species ''A. cromhallensis''. Its remai ...
'' was originally described as a dinosauriform closer to Dinosauria than ''
Herrerasaurus ''Herrerasaurus'' is a genus of saurischian dinosaur from the Late Triassic period. This genus was one of the earliest dinosaurs from the fossil record. Its name means "Herrera's lizard", after the rancher who discovered the first specimen in ...
'',Nicholas C. Fraser, Kevin Padian, Gordon M. Walkden and A. L. M. Davis, 2002. Basal dinosauriform remains from Britain and the diagnosis of the Dinosauria. Palaeontology. 45(1), 79-95. but has also been classified as a dinosauriform more derived than silesaurids but basal to
Herrerasauridae Herrerasauridae is a family of carnivorous dinosaurs, possibly basal to either theropods or even all of saurischians, or even their own branching from dracohors, separate from dinosauria altogether. They are among the oldest known dinosaurs, ...
and Dinosauria, a silesaurid, or a basal saurischian. The genus ''
Nyasasaurus ''Nyasasaurus'' (meaning " Lake Nyasa lizard") is an extinct genus of avemetatarsalian archosaur from the putatively Middle Triassic Manda Formation of Tanzania that may be the earliest known dinosaur. The type species ''Nyasasaurus parringtoni' ...
'' from the early Late Triassic of
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
is known from multiple incomplete specimens, making it difficult to classify. It has been found as the direct sister taxon of Dinosauria, the basalmost ornithischian, a basal theropod, or a deeply-nested sauropodomorph.


Origins

Dinosauromorphs appeared putatively around 242 to 244 million years ago by the
Anisian In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from million years ago until million years ago. The Anisian Age succeeds the Olenekian Age (part of the Lower Triassic ...
stage of the
Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between Ma and ...
, splitting from other ornithodires.
Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a un ...
footprints reported in October 2010 from the Świętokrzyskie (Holy Cross) Mountains of Poland may belong to a dinosauromorph. If so, the origin of dinosauromorphs would be pushed back into the Early
Olenekian In the geologic timescale, the Olenekian is an age in the Early Triassic epoch; in chronostratigraphy, it is a stage in the Lower Triassic series. It spans the time between Ma and Ma (million years ago). The Olenekian is sometimes divided i ...
, around 249 Ma. The oldest Polish footprints are from a small quadrupedal animal named ''
Prorotodactylus ''Prorotodactylus'' is a dinosauromorph or pterosauromorph ichnogenus known from fossilized footprints found in Poland and France. The prints may have been made by a dinosauromorph that was a precursor to the dinosaurs, possibly closely related ...
'', but footprints belonging to the
ichnogenus An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxa'' comes from the Greek ίχνος, ''ichnos'' meaning ''track'' and ταξις, ''taxis'' meaning ...
'' Sphingopus'' that have been found from Early Anisian strata show that moderately large bipedal dinosauromorphs had appeared by 246 Ma. The tracks show that the dinosaur lineage appeared soon after the Permian-Triassic extinction event. Their age suggests that the rise of dinosaurs was slow and drawn out across much of the Triassic. The oldest known dinosauromorph is ''
Asilisaurus ''Asilisaurus'' ( ); from Swahili, ''asili'' ("ancestor" or "foundation"), and Greek, (, "lizard") is an extinct genus of silesaurid archosaur. The type species is ''Asilisaurus kongwe.'' ''Asilisaurus'' fossils were uncovered in the Manda Bed ...
'', a
silesaurid Silesauridae is an extinct family of Triassic dinosauriforms. It is most commonly considered to be a clade of non-dinosaur dinosauriforms, and the sister group of dinosaurs. Some studies have instead suggested that most or all silesaurids compri ...
which may have lived as early as the
Anisian In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from million years ago until million years ago. The Anisian Age succeeds the Olenekian Age (part of the Lower Triassic ...
age of the middle
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period ...
period, about 245 million years ago, although it is possible that ''Nyasasaurus'' is a dinosaur of the same age, pushing the origins of the groups back further. Putative basal dinosauromorphs include ''
Saltopus ''Saltopus'' ("hopping foot") is a genus of very small bipedal dinosauriform containing the single species ''Saltopus elginensis'' from the late Triassic period of Scotland. It is one of the most famous Elgin Reptiles. Description ''Saltopus ...
'', ''
Marasuchus ''Marasuchus'' (meaning "Mara crocodile") is a genus of basal dinosauriform archosaur which is possibly synonymous with '' Lagosuchus''. Both genera lived during the Late Triassic in what is now La Rioja Province, Argentina. ''Marasuchus'' conta ...
'', the perhaps identical ''
Lagosuchus ''Lagosuchus'' is an extinct genus of avemetatarsalian archosaur from the Late Triassic of Argentina. The type species of ''Lagosuchus'', ''Lagosuchus talampayensis,'' is based on a small partial skeleton recovered from the early Carnian-age Cha ...
'', the lagerpetid ''
Lagerpeton ''Lagerpeton'' is a genus of lagerpetid avemetatarsalian, comprising a single species, ''L. chanarensis''. First described from the Chañares Formation of Argentina by A. S. Romer in 1971, ''Lagerpeton'' anatomy is somewhat incompletely kn ...
'' from the
Ladinian The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic). ...
of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
and ''
Dromomeron ''Dromomeron'' (meaning "running femur") is a genus of lagerpetid avemetatarsalian which lived around 220 to 211.9 ± 0.7 million years ago. The genus contains species known from Late Triassic-age rocks of the Southwestern United States and nort ...
'' from the
Norian The Norian is a division of the Triassic Period. It has the rank of an age (geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic defi ...
of
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
, and
Texas Texas (, ; Spanish language, Spanish: ''Texas'', ''Tejas'') is a state in the South Central United States, South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2 ...
(all in the United States), '' Ixalerpeton polesinensis'' and an unnamed form from the
Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...
(
Santa Maria Formation The Santa Maria Formation is a sedimentary rock formation found in Rio Grande do Sul, Brazil. It is primarily Carnian in age (Late Triassic), and is notable for its fossils of cynodonts, "rauisuchian" pseudosuchians, and early dinosaurs and oth ...
) of
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, and the
silesaurid Silesauridae is an extinct family of Triassic dinosauriforms. It is most commonly considered to be a clade of non-dinosaur dinosauriforms, and the sister group of dinosaurs. Some studies have instead suggested that most or all silesaurids compri ...
s, which include '' Silesaurus'' from the
Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followed by t ...
of
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
, '' Eucoelophysis'' from the Carnian-Norian of New Mexico, '' Lewisuchus'' and the perhaps identical ''
Pseudolagosuchus ''Lewisuchus'' is a genus of archosaur that lived during the Late Triassic (early Carnian). As a silesaurid dinosauriform, it was a member of the group of reptiles most commonly considered to be the closest relatives of dinosaurs (possibly tr ...
'' from the Ladinian of
Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...
, '' Sacisaurus'' from the Norian of
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
, '' Technosaurus'' from the Carnian of Texas, ''
Asilisaurus ''Asilisaurus'' ( ); from Swahili, ''asili'' ("ancestor" or "foundation"), and Greek, (, "lizard") is an extinct genus of silesaurid archosaur. The type species is ''Asilisaurus kongwe.'' ''Asilisaurus'' fossils were uncovered in the Manda Bed ...
'' from the
Anisian In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from million years ago until million years ago. The Anisian Age succeeds the Olenekian Age (part of the Lower Triassic ...
of
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
, and ''
Diodorus Diodorus Siculus, or Diodorus of Sicily ( grc-gre, Διόδωρος ;  1st century BC), was an ancient Greek historian. He is known for writing the monumental universal history ''Bibliotheca historica'', in forty books, fifteen of which su ...
'' from the Carnian(?) to Norian of
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to ...
.


References


External links

* * * {{Portal bar, Dinosaurs, Reptiles, Animals, Paleontology, Biology Anisian first appearances Extant Middle Triassic first appearances Taxa named by Michael Benton