Diguetia
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''Diguetia'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of coneweb spiders that was first described by
Eugène Simon Eugène Louis Simon (; 30 April 1848 – 17 November 1924) was a French naturalist who worked particularly on insects and spiders, but also on birds and plants. He is by far the most prolific spider taxonomist in history, describing over 4, ...
in 1895. Members of this genus are six-eyed spiders that are either white or patterned. They are common in the southwestern United States and Mexico, and one
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
is found in Argentina. In the United States, species have been found in Arizona, California, Nevada, New Mexico, and Texas. These spiders build a tubular retreat at the tip of their tent-like webs. Once an insect is caught in the web, the spider bites it and injects
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
to stop its prey from moving, later wrapping it in silk. Both males and females use
stridulation Stridulation is the act of producing sound by rubbing together certain body parts. This behavior is mostly associated with insects, but other animals are known to do this as well, such as a number of species of fish, snakes and spiders. The mech ...
while mating, with females also stridulating when harassed. Two species of jumping spiders feed on its eggs. There are eleven ''Diguetia'' species.


Identification

The species in the genus are haplogynes that have flat oval
carapace A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...
s and six eyes which are arranged into three groups of two, known as dyads. The
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the torso. ...
is either white or has patterns, with some species having bands on their legs. ''Diguetia'' species have three
tarsal claws This glossary describes the terms used in formal descriptions of spiders; where applicable these terms are used in describing other arachnids. Links within the glossary are shown . Terms A Abdomen or opisthosoma: One of the two main body parts ...
. Although all spiders may have such features, only spiders of this genus have a combination of those features.


Habitat and diet

The genus is common in the southwestern United States and Mexico, with the exception of one Argentinian species. In the United States, species have been found in Arizona, California, Nevada, New Mexico, and Texas. The most common species in the U.S. are '' D. canities'' and the smaller '' D. albolineata''. These spiders build an unusual tent-like web, often between
cacti A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Greek ...
, that somewhat resembles the webs of
Linyphiidae Linyphiidae, spiders commonly known as sheet weavers (from the shape of their webs), or money spiders (in the United Kingdom, Ireland, Australia, New Zealand, and in Portugal, from the superstition that if such a spider is seen running on you, ...
. They build a tubular retreat at the tip, camouflaged with leaf litter or similar substances, where thin egg sacs are positioned. The retreat is built in the center out of dead plant material, parts of insects, and insect exoskeleton remains. A horizontal piece of web stretches from the entrance and the spider uses it as platform to move from the bottom. Surrounding the web and retreat are threads which are connected to vegetation. Its web is able to survive strong winds. Once an insect is caught in the web, the spider bites it and injects
venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
to stop its prey from moving. The spider later wraps silk around its prey to hold it still or to bring the insect's wings together by its body. The feeding of the genus is believed to be close to that of earlier discovered spiders of webs that are suspended in the air due to them evolving from only using venom to capture prey, to using both venom and web. This form of capture allows the spider to bite the insect several times.


Reproduction

Not much is known about its reproduction cycle, but it is known that both males and females use
stridulation Stridulation is the act of producing sound by rubbing together certain body parts. This behavior is mostly associated with insects, but other animals are known to do this as well, such as a number of species of fish, snakes and spiders. The mech ...
while mating; if a female is harassed, it stridulates for a short period, making a loud enough noise for humans to hear. The females become mature during the beginning of summer and they produce their first egg sacs in August, continuing to produce them through September. The egg sacs are shaped like a discus and are stacked together. As many as 10 egg sacs are produced per female, each egg sac holding 150 to 250 eggs, for a total of over a thousand eggs. Less than 1% of the eggs will survive to maturity. The female will at first protect her eggs from
predators Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
, such as from the jumping spiders '' Metaphidippus manni'' and '' Habronattus tranquillus''. However, when she dies, the jumping spiders will invade and lay their eggs in the retreat. The eggs of the jumping spiders hatch first, and both their offspring and the adults will feed on the ''Diguetia'' eggs and juveniles. Larva of the clerid beetle '' Phyllobaenus discoideus'' is also a potential predator of ''Diguetia'' eggs.


Species

it contains eleven species: *''
Diguetia albolineata ''Diguetia albolineata'' is a species of desertshrub spider in the family Diguetidae Coneweb spiders (Diguetidae) are six-eyed haplogyne spiders that live in tangled space webs, fashioning a cone-like central retreat where they hide and lay ...
'' (
O. Pickard-Cambridge Octavius Pickard-Cambridge FRS (3 November 1828 – 9 March 1917) was an English clergyman and zoologist. He was a keen arachnologist who described and named more than 900 species of spider. Life and work Pickard-Cambridge was born in Blox ...
, 1895)
– Southern California to Texas, Mexico *'' Diguetia andersoni''
Gertsch Willis John Gertsch (October 4, 1906 – December 12, 1998) was an American arachnologist. He described over 1,000 species of spiders, scorpions, and other arachnids, including the Brown recluse spider and the Tooth cave spider. Gertsch was born ...
, 1958
Mojave Desert The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily in ...
*'' Diguetia canities'' (McCook, 1890) ( type) – Southwestern United States, Mexico **'' Diguetia c. dialectica''
Chamberlin The Chamberlin is an electro-mechanical keyboard instrument that was a precursor to the Mellotron. It was developed and patented by the American inventor Harry Chamberlin from 1949 to 1956, when the first model was introduced. There are severa ...
, 1924
– Mexico **'' Diguetia c. mulaiki'' Gertsch, 1958 – USA *'' Diguetia catamarquensis'' ( Mello-Leitão, 1941) – Argentina *'' Diguetia imperiosa'' Gertsch & Mulaik, 1940 – Arizona to southern Texas, Mexico *'' Diguetia mojavea'' Gertsch, 1958 – Southern California and Nevada *'' Diguetia propinqua'' (O. Pickard-Cambridge, 1896) – Mexico *''
Diguetia signata ''Diguetia signata'' is a species of desertshrub spider in the family Diguetidae Coneweb spiders (Diguetidae) are six-eyed haplogyne spiders that live in tangled space webs, fashioning a cone-like central retreat where they hide and lay eggs. ...
'' Gertsch, 1958 – Southern California to New Mexico, Mexico *''
Diguetia stridulans ''Diguetia'' is a genus of coneweb spiders that was first described by Eugène Simon in 1895. Members of this genus are six-eyed spiders that are either white or patterned. They are common in the southwestern United States and Mexico, and one s ...
'' Chamberlin, 1924 – Mexico


References


External links


Diguetia
at
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* {{Taxonbar, from=Q2091737 Araneomorphae genera Diguetidae Taxa named by Eugène Simon