Desmatosuchine
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Desmatosuchinae is a major
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
of
aetosaur Aetosaurs () are heavily armored reptiles belonging to the extinct order Aetosauria (; from Greek, (aetos, "eagle") and (, "lizard")). They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs ...
s within the
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
Desmatosuchia Aetosaurs () are heavily armored reptiles belonging to the extinct order (biology), order Aetosauria (; from Ancient Greek, Greek, (aetos, "eagle") and (, "lizard")). They were medium- to large-sized Omnivore, omnivorous or Herbivore, herbivoro ...
. It is a
stem-based taxon Phylogenetic nomenclature is a method of nomenclature for taxa in biology that uses phylogenetic definitions for taxon names as explained below. This contrasts with the traditional approach, in which taxon names are defined by a '' type'', which ...
defined as all aetosaurs more closely related to ''
Desmatosuchus ''Desmatosuchus'' (, from Greek δεσμός ''desmos'' 'link' + σοῦχος ''soûkhos'' 'crocodile') is an extinct genus of archosaur belonging to the Order Aetosauria. It lived during the Late Triassic. Description ''Desmatosuchus'' was a ...
'' than to ''
Stagonolepis ''Stagonolepis'' is an extinct genus of stagonolepidid aetosaur known from the Late Triassic (Carnian stage) Hassberge Formation of Germany, the Drawno Beds of Poland, and the Lossiemouth Sandstone of Scotland. Supposed fossils from North and ...
,'' ''
Aetosaurus ''Aetosaurus'' is an extinct genus of pseudosuchian reptile belonging to the order Aetosauria. It is generally considered to be the most primitive aetosaur. Three species are currently recognized: ''A. ferratus'', the type species from Germany an ...
'', or ''
Paratypothorax ''Paratypothorax'' is an extinct genus of aetosaur, known from a single species, ''Paratypothorax andressorum''. It was a broadly distributed member of the group found in Germany, North America, and possibly parts of Gondwana. The best specimen ...
''. The clade Desmatosuchinae has often been restricted to a few closely-related aetosaurs with spiny armor, such as ''Desmatosuchus'', ''
Longosuchus ''Longosuchus'' (meaning "Long's crocodile") is an extinct genus of aetosaur from the Late Triassic of North America and Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of Nort ...
'', and ''
Lucasuchus ''Lucasuchus'' is an extinct genus of aetosaur. Fossils have been found from the Bull Canyon Formation of the Dockum Group outcropping in the Revuelto Creek locality in Quay County, New Mexico. All specimens date back to the Norian stage of the ...
''. It was later expanded to include a number of ''
Stagonolepis ''Stagonolepis'' is an extinct genus of stagonolepidid aetosaur known from the Late Triassic (Carnian stage) Hassberge Formation of Germany, the Drawno Beds of Poland, and the Lossiemouth Sandstone of Scotland. Supposed fossils from North and ...
''-like aetosaurs with less specialized armor. Under this more expansive usage, the strongly-supported clade encompassing "traditional" desmatosuchines (''sensu stricto'') was given a new name, Desmatosuchini.
Synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have ...
that diagnose this clade can be found in the
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amp ...
s. These include tongue-and-groove articulations for lateral plates present in dorsal presacral paramedian plates and large spikes on the lateral cervical, dorsal, and caudal plates.


Timeline of generic descriptions

ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:1890 till:2035 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:50 start:1890 ScaleMinor = unit:year increment:10 start:1890 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:1900s value:rgb(0.94,0.25,0.24) id:2000s value:rgb(0.2,0.7,0.79) id:2000syears value:rgb(0.52,0.81,0.91) id:1900syears value:rgb(0.95,0.56,0.45) id:1700s value:rgb(0.5,0.78,0.31) id:1700syears value:rgb(0.63,0.78,0.65) id:latecretaceous value:rgb(0.74,0.82,0.37) id:1800syears value:rgb(0.95,0.98,0.11) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:1800s value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:NAM2 bar:NAM3 bar:NAM4 bar:NAM5 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: 1890 till: 1900 color:1800syears text:
90s The 90s ran from 90 AD to 99 AD. Significant people * Titus Flavius Domitianus, Roman Emperor (AD 81– 96) * Nerva, Roman Emperor (AD 96 AD 96 ( XCVI) was a leap year starting on Friday (link will display the full calendar) of the Jul ...
from: 1900 till: 1910 color:1900syears text: 00s from: 1910 till: 1920 color:1900syears text:
10s The 10s decade ran from January 1, AD 10, to December 31, AD 19. In Europe, the decade saw the end of the Early Imperial campaigns in Germania when Roman forces led by Germanicus defeated Germanic tribes in the Battle of Idistaviso in 16 AD. I ...
from: 1920 till: 1930 color:1900syears text: 20s from: 1930 till: 1940 color:1900syears text:
30s The prokaryotic small ribosomal subunit, or 30 S subunit, is the smaller subunit of the 70S ribosome found in prokaryotes. It is a complex of the 16S ribosomal RNA (rRNA) and 19 proteins. This complex is implicated in the binding of transfer ...
from: 1940 till: 1950 color:1900syears text:
40s The eukaryotic small ribosomal subunit (40S) is the smaller subunit of the eukaryotic 80S ribosomes, with the other major component being the large ribosomal subunit (60S). The "40S" and "60S" names originate from the convention that ribosomal pa ...
from: 1950 till: 1960 color:1900syears text:
50s The 50s decade ran from January 1, 50, to December 31, 59. It was the sixth decade in the Anno Domini/Common Era, if the nine-year period from 1 AD to 9 AD is considered as a "decade". Significant people * Claudius, Roman Emperor (AD 41â ...
from: 1960 till: 1970 color:1900syears text: 60s from: 1970 till: 1980 color:1900syears text:
70s Ribosomes ( ) are macromolecular machines, found within all cells, that perform biological protein synthesis (mRNA translation). Ribosomes link amino acids together in the order specified by the codons of messenger RNA (mRNA) molecules to for ...
from: 1980 till: 1990 color:1900syears text:
80s Ribosomes are a large and complex molecular machine that catalyzes the synthesis of proteins, referred to as translation. The ribosome selects aminoacylated transfer RNAs (tRNAs) based on the sequence of a protein-encoding messenger RNA (mRNA) ...
from: 1990 till: 2000 color:1900syears text:
90s The 90s ran from 90 AD to 99 AD. Significant people * Titus Flavius Domitianus, Roman Emperor (AD 81– 96) * Nerva, Roman Emperor (AD 96 AD 96 ( XCVI) was a leap year starting on Friday (link will display the full calendar) of the Jul ...
from: 2000 till: 2010 color:2000syears text: 00s from: 2010 till: 2020 color:2000syears text:
10s The 10s decade ran from January 1, AD 10, to December 31, AD 19. In Europe, the decade saw the end of the Early Imperial campaigns in Germania when Roman forces led by Germanicus defeated Germanic tribes in the Battle of Idistaviso in 16 AD. I ...
from: 2020 till: 2030 color:2000syears text: 20s from: 2030 till: 2035 color:2000syears text:
30s The prokaryotic small ribosomal subunit, or 30 S subunit, is the smaller subunit of the 70S ribosome found in prokaryotes. It is a complex of the 16S ribosomal RNA (rRNA) and 19 proteins. This complex is implicated in the binding of transfer ...
bar:eratop from: 1890 till: 1900 color:1800s text:
19th 19 (nineteen) is the natural number following 18 and preceding 20. It is a prime number. Mathematics 19 is the eighth prime number, and forms a sexy prime with 13, a twin prime with 17, and a cousin prime with 23. It is the third full re ...
from: 1900 till: 2000 color:1900s text: 20th from: 2000 till: 2035 color:2000s text: 21st PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:1900s bar:NAM1 at:1990 mark:(line,black) text: Longosuchus meadei color:1900s bar:NAM3 at:1995 mark:(line,black) text: Lucasuchus hunti color:1900s bar:NAM5 at:2008 mark:(line,black) text: Sierritasuchus macalpini color:1800s bar:NAM1 at:1892 mark:(line,black) text: Desmatosuchus haplocerus color:1900s bar:NAM2 at:1921 mark:(line,black) text:
Desmatosuchus ''Desmatosuchus'' (, from Greek δεσμός ''desmos'' 'link' + σοῦχος ''soûkhos'' 'crocodile') is an extinct genus of archosaur belonging to the Order Aetosauria. It lived during the Late Triassic. Description ''Desmatosuchus'' was a ...
& D. spurensis color:1900s bar:NAM4 at:2005 mark:(line,black) text: Desmatosuchus smalli PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: 1890 till: 1900 color:1800syears text:
90s The 90s ran from 90 AD to 99 AD. Significant people * Titus Flavius Domitianus, Roman Emperor (AD 81– 96) * Nerva, Roman Emperor (AD 96 AD 96 ( XCVI) was a leap year starting on Friday (link will display the full calendar) of the Jul ...
from: 1900 till: 1910 color:1900syears text: 00s from: 1910 till: 1920 color:1900syears text:
10s The 10s decade ran from January 1, AD 10, to December 31, AD 19. In Europe, the decade saw the end of the Early Imperial campaigns in Germania when Roman forces led by Germanicus defeated Germanic tribes in the Battle of Idistaviso in 16 AD. I ...
from: 1920 till: 1930 color:1900syears text: 20s from: 1930 till: 1940 color:1900syears text:
30s The prokaryotic small ribosomal subunit, or 30 S subunit, is the smaller subunit of the 70S ribosome found in prokaryotes. It is a complex of the 16S ribosomal RNA (rRNA) and 19 proteins. This complex is implicated in the binding of transfer ...
from: 1940 till: 1950 color:1900syears text:
40s The eukaryotic small ribosomal subunit (40S) is the smaller subunit of the eukaryotic 80S ribosomes, with the other major component being the large ribosomal subunit (60S). The "40S" and "60S" names originate from the convention that ribosomal pa ...
from: 1950 till: 1960 color:1900syears text:
50s The 50s decade ran from January 1, 50, to December 31, 59. It was the sixth decade in the Anno Domini/Common Era, if the nine-year period from 1 AD to 9 AD is considered as a "decade". Significant people * Claudius, Roman Emperor (AD 41â ...
from: 1960 till: 1970 color:1900syears text: 60s from: 1970 till: 1980 color:1900syears text:
70s Ribosomes ( ) are macromolecular machines, found within all cells, that perform biological protein synthesis (mRNA translation). Ribosomes link amino acids together in the order specified by the codons of messenger RNA (mRNA) molecules to for ...
from: 1980 till: 1990 color:1900syears text:
80s Ribosomes are a large and complex molecular machine that catalyzes the synthesis of proteins, referred to as translation. The ribosome selects aminoacylated transfer RNAs (tRNAs) based on the sequence of a protein-encoding messenger RNA (mRNA) ...
from: 1990 till: 2000 color:1900syears text:
90s The 90s ran from 90 AD to 99 AD. Significant people * Titus Flavius Domitianus, Roman Emperor (AD 81– 96) * Nerva, Roman Emperor (AD 96 AD 96 ( XCVI) was a leap year starting on Friday (link will display the full calendar) of the Jul ...
from: 2000 till: 2010 color:2000syears text: 00s from: 2010 till: 2020 color:2000syears text:
10s The 10s decade ran from January 1, AD 10, to December 31, AD 19. In Europe, the decade saw the end of the Early Imperial campaigns in Germania when Roman forces led by Germanicus defeated Germanic tribes in the Battle of Idistaviso in 16 AD. I ...
from: 2020 till: 2030 color:2000syears text: 20s from: 2030 till: 2035 color:2000syears text:
30s The prokaryotic small ribosomal subunit, or 30 S subunit, is the smaller subunit of the 70S ribosome found in prokaryotes. It is a complex of the 16S ribosomal RNA (rRNA) and 19 proteins. This complex is implicated in the binding of transfer ...
bar:era from: 1890 till: 1900 color:1800s text:
19th 19 (nineteen) is the natural number following 18 and preceding 20. It is a prime number. Mathematics 19 is the eighth prime number, and forms a sexy prime with 13, a twin prime with 17, and a cousin prime with 23. It is the third full re ...
from: 1900 till: 2000 color:1900s text: 20th from: 2000 till: 2035 color:2000s text: 21st


References

{{Taxonbar, from=Q3494921 Aetosaurs Late Triassic first appearances