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Deer or true deer are
hoof The hoof (plural: hooves) is the tip of a toe of an ungulate mammal, which is covered and strengthened with a thick and horny keratin covering. Artiodactyls are even-toed ungulates, species whose feet have an even number of digits, yet the rumin ...
ed
ruminant Ruminants (suborder Ruminantia) are ungulate, hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by Enteric fermentation, fermenting it in a specialized stomach prior to digestion, principally ...
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or ...
s forming the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Cervidae. The two main groups of deer are the
Cervinae The Cervinae or the Old World deer, are a subfamily of deer. Alternatively, they are known as the plesiometacarpal deer, due to their ankle structure being different from the telemetacarpal deer of the Capreolinae. Classification and species The ...
, including the
muntjac Muntjacs ( ), also known as the barking deer or rib-faced deer, (URL is Google Books) are small deer of the genus ''Muntiacus'' native to South Asia and Southeast Asia. Muntjacs are thought to have begun appearing 15–35 million years ago, ...
, the
elk The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The common ...
(wapiti), the
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
, and the
fallow deer ''Dama'' is a genus of deer in the subfamily Cervinae, commonly referred to as fallow deer. Name The name fallow is derived from the deer's pale brown colour. The Latin word ''dāma'' or ''damma'', used for roe deer, gazelles, and antelopes, ...
; and the
Capreolinae The Capreolinae, Odocoileinae, or the New World deer are a subfamily of deer. Alternatively, they are known as the telemetacarpal deer, due to their bone structure being different from the plesiometacarpal deer subfamily Cervinae. The telemetac ...
, including the
reindeer Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subspe ...
(caribou),
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...
, the
roe deer The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
, and the
moose The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
. Male deer of all species (except the
water deer The water deer (''Hydropotes inermis'') is a small deer superficially more similar to a musk deer than a true deer. Native to China and Korea, there are two subspecies: the Chinese water deer (''Hydropotes inermis inermis'') and the Korean water ...
), as well as female reindeer, grow and shed new
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on male ...
s each year. In this they differ from permanently
horn Horn most often refers to: *Horn (acoustic), a conical or bell shaped aperture used to guide sound ** Horn (instrument), collective name for tube-shaped wind musical instruments *Horn (anatomy), a pointed, bony projection on the head of various ...
ed
antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mammals ...
, which are part of a different family (
Bovidae The Bovidae comprise the biological family of cloven-hoofed, ruminant mammals that includes cattle, bison, buffalo, antelopes, and caprines. A member of this family is called a bovid. With 143 extant species and 300 known extinct species, ...
) within the same order of
even-toed ungulate The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
s (Artiodactyla). The
musk deer Musk deer can refer to any one, or all seven, of the species that make up ''Moschus'', the only extant genus of the family Moschidae. Despite being commonly called deer, they are not true deer belonging to the family Cervidae, but rather their fa ...
(
Moschidae Moschidae is a family of pecoran even-toed ungulates, containing the musk deer (''Moschus'') and its extinct relatives. They are characterized by long 'saber teeth' instead of horns, antlers or ossicones, modest size (''Moschus'' only reaches ; ...
) of Asia and
chevrotain Chevrotains, or mouse-deer, are small even-toed ungulates that make up the family Tragulidae, the only extant members of the infraorder Tragulina. The 10  extant species are placed in three genera, but several species also are known only ...
s (
Tragulidae The asterisk ( ), from Late Latin , from Ancient Greek , ''asteriskos'', "little star", is a typographical symbol. It is so called because it resembles a conventional image of a heraldic star. Computer scientists and mathematicians often voc ...
) of tropical African and Asian forests are separate families that are also in the ruminant clade
Ruminantia Ruminants ( suborder Ruminantia) are hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The ...
; they are not especially closely related to Cervidae. Deer appear in art from
Paleolithic The Paleolithic or Palaeolithic (), also called the Old Stone Age (from Greek: παλαιός ''palaios'', "old" and λίθος ''lithos'', "stone"), is a period in human prehistory that is distinguished by the original development of stone too ...
cave painting In archaeology, Cave paintings are a type of parietal art (which category also includes petroglyphs, or engravings), found on the wall or ceilings of caves. The term usually implies prehistoric origin, and the oldest known are more than 40,000 ye ...
s onwards, and they have played a role in mythology, religion, and literature throughout history, as well as in
heraldry Heraldry is a discipline relating to the design, display and study of armorial bearings (known as armory), as well as related disciplines, such as vexillology, together with the study of ceremony, rank and pedigree. Armory, the best-known branch ...
, such as red deer that appear in the
coat of arms of Åland The coat of arms of Åland features a golden red deer on a blue field. This is traditionally surmounted by a comital coronet of the elder Swedish style. History First known symbol of the Åland islands is a seal from 1326 depicting Saint Olaf ...
. Their economic importance includes the use of their meat as
venison Venison originally meant the meat of a game animal but now refers primarily to the meat of antlered ungulates such as elk or deer (or antelope in South Africa). Venison can be used to refer to any part of the animal, so long as it is edible, in ...
, their skins as soft, strong buckskin, and their antlers as handles for knives. Deer
hunting Hunting is the human activity, human practice of seeking, pursuing, capturing, or killing wildlife or feral animals. The most common reasons for humans to hunt are to harvest food (i.e. meat) and useful animal products (fur/hide (skin), hide, ...
has been a popular activity since the Middle Ages and remains a resource for many families today.


Etymology and terminology

The word ''deer'' was originally broad in meaning, becoming more specific with time.
Old English Old English (, ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the early Middle Ages. It was brought to Great Britain by Anglo-Saxon settlement of Britain, Anglo ...
''dēor'' and
Middle English Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old English p ...
''der'' meant a wild animal of any kind. Cognates of Old English ''dēor'' in other dead
Germanic languages The Germanic languages are a branch of the Indo-European language family spoken natively by a population of about 515 million people mainly in Europe, North America, Oceania and Southern Africa. The most widely spoken Germanic language, Engli ...
have the general sense of ''animal'', such as
Old High German Old High German (OHG; german: Althochdeutsch (Ahd.)) is the earliest stage of the German language, conventionally covering the period from around 750 to 1050. There is no standardised or supra-regional form of German at this period, and Old High ...
''tior'',
Old Norse Old Norse, Old Nordic, or Old Scandinavian, is a stage of development of North Germanic languages, North Germanic dialects before their final divergence into separate Nordic languages. Old Norse was spoken by inhabitants of Scandinavia and t ...
''djur'' or ''dȳr'',
Gothic Gothic or Gothics may refer to: People and languages *Goths or Gothic people, the ethnonym of a group of East Germanic tribes **Gothic language, an extinct East Germanic language spoken by the Goths **Crimean Gothic, the Gothic language spoken b ...
''dius'',
Old Saxon Old Saxon, also known as Old Low German, was a Germanic language and the earliest recorded form of Low German (spoken nowadays in Northern Germany, the northeastern Netherlands, southern Denmark, the Americas and parts of Eastern Europe). It i ...
''dier'', and
Old Frisian Old Frisian was a West Germanic language spoken between the 8th and 16th centuries along the North Sea coast, roughly between the mouths of the Rhine and Weser rivers. The Frisian settlers on the coast of South Jutland (today's Northern Friesl ...
''diar''. This general sense gave way to the modern English sense by the end of the Middle English period, around 1500. All modern Germanic languages save English and Scots retain the more general sense: for example, German ''Tier'' and Norwegian ''dyr'' mean ''animal''. For many types of deer in modern English usage, the male is a ''buck'' and the female a ''doe'', but the terms vary with dialect, and according to the size of the species. The male
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
is a ''stag'', while for other large species the male is a ''bull'', the female a ''cow'', as in cattle. In older usage, the male of any species is a ''
hart Hart often refers to: * Hart (deer) Hart may also refer to: Organizations * Hart Racing Engines, a former Formula One engine manufacturer * Hart Skis, US ski manufacturer * Hart Stores, a Canadian chain of department stores * Hart's Reptile W ...
'', especially if over five years old, and the female is a ''hind'', especially if three or more years old. The young of small species is a ''fawn'' and of large species a ''
calf Calf most often refers to: * Calf (animal), the young of domestic cattle. * Calf (leg), in humans (and other primates), the back portion of the lower leg Calf or calves may also refer to: Biology and animal byproducts * Veal, meat from calves * ...
''; a very small young may be a ''kid''. A castrated male is a ''havier''. A group of any species is a ''herd''. The
adjective In linguistics, an adjective (list of glossing abbreviations, abbreviated ) is a word that generally grammatical modifier, modifies a noun or noun phrase or describes its referent. Its semantic role is to change information given by the noun. Tra ...
of relation is ''
cervine Deer or true deer are hoofed ruminant mammals forming the family (biology), family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, inclu ...
''; like the family name ''Cervidae'', this is from la, cervus, meaning ''stag'' or ''deer''.


Distribution

Deer live in a variety of
biome A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
s, ranging from
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
to the
tropical rainforest Tropical rainforests are rainforests that occur in areas of tropical rainforest climate in which there is no dry season – all months have an average precipitation of at least 60 mm – and may also be referred to as ''lowland equatori ...
. While often associated with forests, many deer are
ecotone An ecotone is a transition area between two biological communities, where two communities meet and integrate. It may be narrow or wide, and it may be local (the zone between a field and forest) or regional (the transition between forest and gras ...
species that live in transitional areas between forests and thickets (for cover) and prairie and savanna (open space). The majority of large deer species inhabit temperate mixed deciduous forest, mountain mixed coniferous forest, tropical seasonal/dry forest, and savanna habitats around the world. Clearing open areas within forests to some extent may actually benefit deer populations by exposing the
understory In forestry and ecology, understory (American English), or understorey (Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the forest canopy without penetrating it to any great extent, but abov ...
and allowing the types of grasses, weeds, and herbs to grow that deer like to eat. Access to adjacent croplands may also benefit deer. Adequate forest or brush cover must still be provided for populations to grow and thrive. Deer are widely distributed, with indigenous representatives in all continents except
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
and
Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...
, though
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
has only one native deer, the
Barbary stag The Barbary stag (''Cervus elaphus barbarus''), also known as the Atlas deer or African elk, is a subspecies of the red deer that is native to North Africa. It is the only deer known to be native to Africa, aside from '' Megaceroides algericus'', ...
, a subspecies of
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or hart, and a female is called a hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Iran, and parts of wes ...
that is confined to the
Atlas Mountains The Atlas Mountains are a mountain range in the Maghreb in North Africa. It separates the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range. It stretches around through Moroc ...
in the northwest of the continent. Another extinct species of deer, ''
Megaceroides algericus ''Megaceroides algericus'' is an extinct species of deer known from the Late Pleistocene to the Holocene of North Africa. It is one of only two species of deer known to have been native to the African continent, alongside the Barbary stag, a subs ...
,'' was present in
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
until 6000 years ago.
Fallow deer ''Dama'' is a genus of deer in the subfamily Cervinae, commonly referred to as fallow deer. Name The name fallow is derived from the deer's pale brown colour. The Latin word ''dāma'' or ''damma'', used for roe deer, gazelles, and antelopes, ...
have been introduced to
South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...
. Small species of
brocket deer Brockets or brocket deer are the species of deer in the genus ''Mazama''. They are medium to small in size, and are found in the Yucatán Peninsula, Central and South America, and the island of Trinidad. Most species are primarily found in forest ...
and
pudú The pudus (Mapudungun ''püdü'' or ''püdu'', es, pudú, ) are two species of South American deer from the genus ''Pudu'', and are the world's smallest deer. The chevrotains (mouse-deer; Tragulidae) are smaller, but they are not true deer. The ...
s of
Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known as ...
and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
, and
muntjac Muntjacs ( ), also known as the barking deer or rib-faced deer, (URL is Google Books) are small deer of the genus ''Muntiacus'' native to South Asia and Southeast Asia. Muntjacs are thought to have begun appearing 15–35 million years ago, ...
s of
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an area ...
generally occupy dense forests and are less often seen in open spaces, with the possible exception of the
Indian muntjac The Indian muntjac or the common muntjac (''Muntiacus muntjak''), also called the southern red muntjac and barking deer, is a deer species native to South and Southeast Asia. It is listed as Least Concern on the IUCN Red List. In popular local l ...
. There are also several species of deer that are highly specialized and live almost exclusively in mountains, grasslands, swamps, and "wet" savannas, or riparian corridors surrounded by
desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
s. Some deer have a circumpolar distribution in both
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
and
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
. Examples include the
caribou Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subspe ...
that live in Arctic
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
and
taiga Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruce ...
(boreal forests) and
moose The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
that inhabit taiga and adjacent areas. Huemul deer (
taruca The taruca (''Hippocamelus antisensis''), also known as the Peruvian guemal, north Andean deer, north Andean huemul, northern huemul or northern guemal, is a mid sized deer species that inhabits the high regions of the Andes mountains in South Am ...
and
Chilean huemul The south Andean deer (''Hippocamelus bisulcus''), also known as the southern guemal, south Andean huemul, southern huemul, or Chilean ''huemul'' or '' güemul'' ( , ), is an endangered species of deer native to the mountains of Argentina and Chi ...
) of
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southe ...
's
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
fill the ecological niches of the
ibex An ibex (plural ibex, ibexes or ibices) is any of several species of wild goat (genus ''Capra''), distinguished by the male's large recurved horns, which are transversely ridged in front. Ibex are found in Eurasia, North Africa and East Africa. ...
and
wild goat The wild goat (''Capra aegagrus'') is a wild goat species, inhabiting forests, shrublands and rocky areas ranging from Turkey and the Caucasus in the west to Turkmenistan, Afghanistan and Pakistan in the east. It has been listed as near threatene ...
, with the fawns behaving more like
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
kids. The highest concentration of large deer species in temperate North America lies in the Canadian Rocky Mountain and
Columbia Mountain Columbia may refer to: * Columbia (personification), the historical female national personification of the United States, and a poetic name for America Places North America Natural features * Columbia Plateau, a geologic and geographic region in ...
regions between Alberta and British Columbia where all five North American deer species (
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...
,
mule deer The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer. Unlike the related whit ...
,
caribou Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subspe ...
,
elk The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The common ...
, and
moose The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
) can be found. This region has several clusters of national parks including
Mount Revelstoke National Park Mount Revelstoke National Park is a national park located adjacent to the city of Revelstoke, British Columbia, Canada. The park is relatively small for a national park, covering . It is located in the Selkirk Mountains and was founded in 1914. ...
,
Glacier National Park (Canada) Glacier National Park is part of a system of 43 parks and park reserves across Canada, and one of seven national parks in British Columbia. Established in 1886, the park encompasses , and includes a portion of the Selkirk Mountains which are pa ...
,
Yoho National Park Yoho National Park ( ) is a National Parks of Canada, national park of Canada. It is located within the Canadian Rockies, Rocky Mountains along the western slope of the Continental Divide of the Americas in southeastern British Columbia, bordered ...
, and
Kootenay National Park Kootenay National Park is a national park of Canada located in southeastern British Columbia. The park consists of of the Canadian Rockies, including parts of the Kootenay and Park mountain ranges, the Kootenay River and the entirety of the Ve ...
on the British Columbia side, and
Banff National Park Banff National Park is Canada's oldest National Parks of Canada, national park, established in 1885 as Rocky Mountains Park. Located in Alberta's Rockies, Alberta's Rocky Mountains, west of Calgary, Banff encompasses of mountainous terrain, wi ...
,
Jasper National Park Jasper National Park is a national park in Alberta, Canada. It is the largest national park within Alberta's Rocky Mountains spanning . It was established as a national park in 1930 and declared a UNESCO World Heritage Site in 1984. Its locatio ...
, and
Glacier National Park (U.S.) Glacier National Park is an American national park located in northwestern Montana, on the Canada–United States border, adjacent to the Canadian provinces of Alberta and British Columbia. The park encompasses more than and includes parts of t ...
on the Alberta and Montana sides. Mountain slope habitats vary from moist coniferous/mixed forested habitats to dry subalpine/pine forests with alpine meadows higher up. The foothills and river valleys between the mountain ranges provide a mosaic of cropland and deciduous parklands. The rare woodland caribou have the most restricted range living at higher altitudes in the subalpine meadows and
alpine tundra Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated alpine climate, harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alp ...
areas of some of the mountain ranges. Elk and mule deer both migrate between the alpine meadows and lower coniferous forests and tend to be most common in this region. Elk also inhabit river valley bottomlands, which they share with White-tailed deer. The White-tailed deer have recently expanded their range within the foothills and river valley bottoms of the Canadian Rockies owing to conversion of land to cropland and the clearing of coniferous forests allowing more deciduous vegetation to grow up the mountain slopes. They also live in the aspen parklands north of Calgary and Edmonton, where they share habitat with the moose. The adjacent
Great Plains The Great Plains (french: Grandes Plaines), sometimes simply "the Plains", is a broad expanse of flatland in North America. It is located west of the Mississippi River and east of the Rocky Mountains, much of it covered in prairie, steppe, an ...
grassland habitats are left to herds of elk,
American bison The American bison (''Bison bison'') is a species of bison native to North America. Sometimes colloquially referred to as American buffalo or simply buffalo (a different clade of bovine), it is one of two extant species of bison, alongside the ...
, and
pronghorn The pronghorn (, ) (''Antilocapra americana'') is a species of artiodactyl (even-toed, hoofed) mammal indigenous to interior western and central North America. Though not an antelope, it is known colloquially in North America as the American a ...
. The
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
n Continent (including the Indian Subcontinent) boasts the most species of deer in the world, with most species being found in Asia. Europe, in comparison, has lower diversity in plant and animal species. Many national parks and protected reserves in Europe have populations of red deer,
roe deer The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
, and fallow deer. These species have long been associated with the continent of Europe, but also inhabit
Asia Minor Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
, the
Caucasus Mountains The Caucasus Mountains, : pronounced * hy, Կովկասյան լեռներ, : pronounced * az, Qafqaz dağları, pronounced * rus, Кавка́зские го́ры, Kavkázskiye góry, kɐfˈkasːkʲɪje ˈɡorɨ * tr, Kafkas Dağla ...
, and Northwestern
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
. "European" fallow deer historically lived over much of Europe during the Ice Ages, but afterwards became restricted primarily to the Anatolian Peninsula, in present-day Turkey. Present-day fallow deer populations in Europe are a result of historic man-made introductions of this species, first to the Mediterranean regions of Europe, then eventually to the rest of Europe. They were initially park animals that later escaped and reestablished themselves in the wild. Historically, Europe's deer species shared their deciduous forest habitat with other herbivores, such as the extinct
tarpan The term tarpan (''Equus ferus ferus'') refers to free-ranging horses of the Russian steppe from the 18th to the 20th century. It is generally unknown whether those horses represented genuine wild horses, feral domestic horses or hybrids. The las ...
(forest horse), extinct
aurochs The aurochs (''Bos primigenius'') ( or ) is an extinct cattle species, considered to be the wild ancestor of modern domestic cattle. With a shoulder height of up to in bulls and in cows, it was one of the largest herbivores in the Holocen ...
(forest ox), and the endangered
wisent The European bison (''Bison bonasus'') or the European wood bison, also known as the wisent ( or ), the zubr (), or sometimes colloquially as the European buffalo, is a European species of bison. It is one of two extant species of bison, along ...
(European bison). Good places to see deer in Europe include the
Scottish Highlands The Highlands ( sco, the Hielands; gd, a’ Ghàidhealtachd , 'the place of the Gaels') is a historical region of Scotland. Culturally, the Highlands and the Lowlands diverged from the Late Middle Ages into the modern period, when Lowland Sco ...
, the
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
n
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
, the
wetlands A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...
between
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
,
Hungary Hungary ( hu, Magyarország ) is a landlocked country in Central Europe. Spanning of the Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia a ...
, and the
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
, and some National Parks, including
Doñana National Park Doñana National Park or Parque Nacional y Natural de Doñana is a natural reserve in Andalucía, southern Spain, in the provinces of Huelva (most of its territory), Cádiz and Seville. It covers , of which are a protected area. The park is an ...
in
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
, the
Veluwe The Veluwe () is a forest-rich ridge of hills (1100 km2) in the province of Gelderland in the Netherlands. The Veluwe features many different landscapes, including woodland, heath, some small lakes and Europe's largest sand drifts. The Ve ...
in the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
, the
Ardennes The Ardennes (french: Ardenne ; nl, Ardennen ; german: Ardennen; wa, Årdene ; lb, Ardennen ), also known as the Ardennes Forest or Forest of Ardennes, is a region of extensive forests, rough terrain, rolling hills and ridges primarily in Be ...
in
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...
, and
Białowieża National Park '' , iucn_category = II , photo = Bialowieza National Park in Poland0029.JPG , photo_caption = Fallen tree in the Białowieża Forest Park logo with European bison , map = Poland , relief = 1 , map_caption = Location in Poland , location = ...
in
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...
.
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
,
Eastern Europe Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russ ...
, and the
Caucasus Mountains The Caucasus Mountains, : pronounced * hy, Կովկասյան լեռներ, : pronounced * az, Qafqaz dağları, pronounced * rus, Кавка́зские го́ры, Kavkázskiye góry, kɐfˈkasːkʲɪje ˈɡorɨ * tr, Kafkas Dağla ...
have forest areas that are not only home to sizable deer populations but also other animals that were once abundant such as the wisent,
Eurasian lynx The Eurasian lynx (''Lynx lynx'') is a medium-sized wild cat widely distributed from Northern, Central and Eastern Europe to Central Asia and Siberia, the Tibetan Plateau and the Himalayas. It inhabits temperate and boreal forests up to an eleva ...
,
Iberian lynx The Iberian lynx (''Lynx pardinus'') is a wild cat species endemic to the Iberian Peninsula in southwestern Europe. It is listed as Endangered on the IUCN Red List. In the 20th century, the Iberian lynx population had declined because of overhun ...
,
wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
, and
brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is kno ...
s. The highest concentration of large deer species in temperate Asia occurs in the mixed deciduous forests, mountain coniferous forests, and taiga bordering North Korea, Manchuria (Northeastern China), and the Ussuri Region (Russia). These are among some of the richest deciduous and coniferous forests in the world where one can find
Siberian roe deer The Siberian roe deer, eastern roe deer, or Asian roe (''Capreolus pygargus''), is a species of roe deer found in northeastern Asia. In addition to Siberia and Mongolia, it is found in Kazakhstan, the Tian Shan Mountains of Kyrgyzstan, eastern Ti ...
,
sika deer The sika deer (''Cervus nippon''), also known as the Northern spotted deer or the Japanese deer, is a species of deer native to much of East Asia and introduced to other parts of the world. Previously found from northern Vietnam in the south to ...
, elk, and moose. Asian caribou occupy the northern fringes of this region along the Sino-Russian border. Deer such as the sika deer,
Thorold's deer Thorold's deer (''Cervus albirostris'')Pitraa, Fickela, Meijaard, Groves (2004). ''Evolution and phylogeny of old world deer.'' Molecular Phylogenetics and Evolution 33: 880–895. is a threatened species of deer found in grassland, shrubland, an ...
,
Central Asian red deer The Central Asian red deer (''Cervus hanglu''), also known as the Tarim red deer is a deer species native to Central Asia, where it used to be widely distributed, but is scattered today with small population units in several countries. It has bee ...
, and elk have historically been farmed for their antlers by
Han Chinese The Han Chinese () or Han people (), are an East Asian ethnic group native to China. They constitute the world's largest ethnic group, making up about 18% of the global population and consisting of various subgroups speaking distinctive va ...
,
Turkic peoples The Turkic peoples are a collection of diverse ethnic groups of West, Central, East, and North Asia as well as parts of Europe, who speak Turkic languages.. "Turkic peoples, any of various peoples whose members speak languages belonging t ...
,
Tungusic peoples Tungusic peoples are an ethno-linguistic group formed by the speakers of Tungusic languages (or Manchu–Tungus languages). They are native to Siberia and Northeast Asia. The Tungusic phylum is divided into two main branches, northern (Evenic or ...
,
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
ns, and
Koreans Koreans ( South Korean: , , North Korean: , ; see names of Korea) are an East Asian ethnic group native to the Korean Peninsula. Koreans mainly live in the two Korean nation states: North Korea and South Korea (collectively and simply refe ...
. Like the
Sami people Acronyms * SAMI, ''Synchronized Accessible Media Interchange'', a closed-captioning format developed by Microsoft * Saudi Arabian Military Industries, a government-owned defence company * South African Malaria Initiative, a virtual expertise net ...
of Finland and Scandinavia, the Tungusic peoples, Mongolians, and Turkic peoples of Southern Siberia, Northern Mongolia, and the Ussuri Region have also taken to raising semi-domesticated herds of Asian caribou. The highest concentration of large deer species in the tropics occurs in Southern Asia in India's Indo-Gangetic Plain Region and
Nepal Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne, सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is mai ...
's Terai Region. These fertile plains consist of tropical seasonal moist deciduous, dry deciduous forests, and both dry and wet savannas that are home to
chital The chital or cheetal (''Axis axis''; ), also known as the spotted deer, chital deer, and axis deer, is a deer species native to the Indian subcontinent. It was first described and given a binomial name by German naturalist Johann Christian Po ...
,
hog deer ''Axis'' is a genus of deer occurring in South and Southeast Asia. As presently defined by most authorities, four species are placed in the genus. Three of the four species are called hog deer. The genus name is a word mentioned in Pliny the Eld ...
,
barasingha The barasingha (''Rucervus duvaucelii''), also known as the swamp deer, is a deer species distributed in the Indian subcontinent. Populations in northern and central India are fragmented, and two isolated populations occur in southwestern Nepal. ...
, Indian sambar, and
Indian muntjac The Indian muntjac or the common muntjac (''Muntiacus muntjak''), also called the southern red muntjac and barking deer, is a deer species native to South and Southeast Asia. It is listed as Least Concern on the IUCN Red List. In popular local l ...
. Grazing species such as the endangered barasingha and very common chital are gregarious and live in large herds. Indian sambar can be gregarious but are usually solitary or live in smaller herds. Hog deer are solitary and have lower densities than Indian muntjac. Deer can be seen in several national parks in India, Nepal, and Sri Lanka of which
Kanha National Park Kanha Tiger Reserve, also known as Kanha–Kisli National Park, is one of the tiger reserves of India and the largest national park of the state of Madhya Pradesh. The present-day Kanha area is divided into two protected areas, Hallon and Banja ...
,
Dudhwa National Park The Dudhwa National Park is a national park in the Terai belt of marshy grasslands in northern Uttar Pradesh, India. It stretches over an area of , with a buffer zone of . It is part of the Dudhwa Tiger Reserve in the Kheri and Lakhimpur distric ...
, and
Chitwan National Park , iucn_category = II , location = Central Terai of Nepal , established = 1973 , nearest_city = Bharatpur , map = Nepal Bagmati Province#Nepal#India#South Asia , relief = 1 , label = Chitwan National Park , label_position = top , coordina ...
are most famous. Sri Lanka's
Wilpattu National Park Wilpattu National Park (''Willu-pattu'', "Land of Lakes") is a national park in Sri Lanka. The unique feature of this park is the existence of "Willus" (natural lakes) – natural, sand-rimmed water basins or depressions that fill with rainwater. ...
and
Yala National Park Yala (යාල) National Park is the most visited and second largest national park in Sri Lanka, bordering the Indian Ocean. The park consists of five blocks, two of which are now open to the public, and also adjoining parks. The blocks have ind ...
have large herds of Indian sambar and chital. The Indian sambar are more gregarious in Sri Lanka than other parts of their range and tend to form larger herds than elsewhere. The Chao Praya River Valley of Thailand was once primarily tropical seasonal moist deciduous forest and wet savanna that hosted populations of hog deer, the now-extinct
Schomburgk's deer The Schomburgk's deer (''Rucervus schomburgki'') is an extinct species of deer once endemic to central Thailand. It was described by Edward Blyth in 1863 and named after Sir Robert H. Schomburgk, who was the British consul in Bangkok from 1857 t ...
,
Eld's deer Eld's deer (''Rucervus eldii'' or ''Panolia eldii''Pitraa, Fickela, Meijaard, Groves (2004). ''Evolution and phylogeny of old world deer.'' Molecular Phylogenetics and Evolution 33: 880–895.), also known as the thamin or brow-antlered deer, is ...
, Indian sambar, and Indian muntjac. Both the hog deer and Eld's deer are rare, whereas Indian sambar and Indian muntjac thrive in protected national parks, such as
Khao Yai Khao Yai National Park ( th, อุทยานแห่งชาติเขาใหญ่, , ) is a national park in Thailand. Established in 1962 as Thailand's first national park, it is the third largest national park in Thailand. Descr ...
. Many of these South Asian and Southeast Asian deer species also share their habitat with other
herbivores A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
, such as
Asian elephant The Asian elephant (''Elephas maximus''), also known as the Asiatic elephant, is the only living species of the genus ''Elephas'' and is distributed throughout the Indian subcontinent and Southeast Asia, from India in the west, Nepal in the no ...
s, the various Asian rhinoceros species, various antelope species (such as
nilgai The nilgai (''Boselaphus tragocamelus'') (, literally meaning "blue cow") is the largest Asian antelope and is ubiquitous across the northern Indian subcontinent. It is the sole member of the genus ''Boselaphus'' and was described by Peter Sim ...
,
four-horned antelope The four-horned antelope (''Tetracerus quadricornis''), or ''chousingha'', is a small antelope found in India and Nepal. Its four horns distinguish it from most other bovids, which have two horns (with a few exceptions, such as the Jacob sheep). ...
,
blackbuck The blackbuck (''Antilope cervicapra''), also known as the Indian antelope, is an antelope native to India and Nepal. It inhabits grassy plains and lightly forested areas with perennial water sources. It stands up to high at the shoulder. Mal ...
, and
Indian gazelle The chinkara (''Gazella bennettii''), also known as the Indian gazelle, is a gazelle species native to Iran, Afghanistan, Pakistan and India. Taxonomy The following six subspecies are considered valid: * Deccan chinkara (''G. b. bennettii'') ...
in India), and wild oxen (such as
wild Asian water buffalo The wild water buffalo (''Bubalus arnee''), also called Asian buffalo, Asiatic buffalo and wild buffalo, is a large bovine native to the Indian subcontinent and Southeast Asia. It has been listed as ''Endangered'' in the IUCN Red List since 198 ...
,
gaur The gaur (''Bos gaurus''; ), also known as the Indian bison, is a bovine native to South Asia and Southeast Asia, and has been listed as Vulnerable on the IUCN Red List since 1986. The global population was estimated at a maximum of 21,000 m ...
,
banteng The banteng (''Bos javanicus''; ), also known as tembadau, is a species of cattle found in Southeast Asia. The head-and-body length is between . Wild banteng are typically larger and heavier than their domesticated counterparts, but are otherw ...
, and
kouprey The kouprey (''Bos sauveli''), also known as forest ox is a forest-dwelling, wild bovine species native to Southeast Asia. A young male was sent to the Paris Zoological Park in 1937 and was described by the French zoologist Achille Urbain who d ...
). One way that different herbivores can survive together in a given area is for each species to have different food preferences, although there may be some overlap. As a result of
acclimatisation society Acclimatisation societies were voluntary associations in the 19th and 20th centuries that encouraged the introduced species, introduction of non-native species in various places around the world, in the hope that they would acclimatization, acclimat ...
releases in the 19th century, Australia has six
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived there ...
of deer that have established sustainable wild populations. They are fallow deer, red deer, sambar, hog deer, rusa, and chital. Red deer were introduced into New Zealand in 1851 from English and Scottish stock. Many have been domesticated in
deer farm A deer farm (technically a ranch) is fenced piece of a land suitable for grazing that is populated with deer species, such as elk, moose, reindeer, or especially white-tailed deer, raised as livestock. New Zealand is the largest supplier of farm- ...
s since the late 1960s and are common farm animals there now. Seven other species of deer were introduced into New Zealand but none are as widespread as red deer.


Description

Deer constitute the second most diverse family of artiodactyla after bovids. Though of a similar build, deer are strongly distinguished from
antelope The term antelope is used to refer to many species of even-toed ruminant that are indigenous to various regions in Africa and Eurasia. Antelope comprise a wastebasket taxon defined as any of numerous Old World grazing and browsing hoofed mammals ...
s by their
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on male ...
s, which are temporary and regularly regrown unlike the permanent
horn Horn most often refers to: *Horn (acoustic), a conical or bell shaped aperture used to guide sound ** Horn (instrument), collective name for tube-shaped wind musical instruments *Horn (anatomy), a pointed, bony projection on the head of various ...
s of bovids. Characteristics typical of deer include long, powerful legs, a diminutive tail and long ears. Deer exhibit a broad variation in physical proportions. The
largest Large means of great size. Large may also refer to: Mathematics * Arbitrarily large, a phrase in mathematics * Large cardinal, a property of certain transfinite numbers * Large category, a category with a proper class of objects and morphisms (or ...
extant deer is the
moose The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
, which is nearly tall and weighs up to . The elk stands at the shoulder and weighs . The northern pudu is the smallest deer in the world; it reaches merely at the shoulder and weighs . The southern pudu is only slightly taller and heavier.
Sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
is quite pronounced – in most species males tend to be larger than females, and, except for the reindeer, only males possess antlers. Coat colour generally varies between red and brown, though it can be as dark as chocolate brown in the tufted deer or have a grayish tinge as in elk. Different species of brocket deer vary from gray to reddish brown in coat colour. Several species such as the chital, the fallow deer and the sika deer feature white spots on a brown coat. Coat of reindeer shows notable geographical variation. Deer undergo two
moult In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...
s in a year; for instance, in red deer the red, thin-haired summer coat is gradually replaced by the dense, greyish brown winter coat in autumn, which in turn gives way to the summer coat in the following spring. Moulting is affected by the
photoperiod Photoperiodism is the physiological reaction of organisms to the length of night or a dark period. It occurs in plants and animals. Plant photoperiodism can also be defined as the developmental responses of plants to the relative lengths of light a ...
. Deer are also excellent jumpers and swimmers. Deer are
ruminant Ruminants (suborder Ruminantia) are ungulate, hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by Enteric fermentation, fermenting it in a specialized stomach prior to digestion, principally ...
s, or cud-chewers, and have a four-chambered stomach. Some deer, such as those on the island of
Rùm Rùm (), a Scottish Gaelic name often anglicised to Rum (), is one of the Small Isles of the Inner Hebrides, in the district of Lochaber, Scotland. For much of the 20th century the name became Rhum, a spelling invented by the former owner, Sir ...
, do consume meat when it is available. Nearly all deer have a facial gland in front of each eye. The gland contains a strongly scented
pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
, used to
mark Mark may refer to: Currency * Bosnia and Herzegovina convertible mark, the currency of Bosnia and Herzegovina * East German mark, the currency of the German Democratic Republic * Estonian mark, the currency of Estonia between 1918 and 1927 * Fi ...
its home range. Bucks of a wide range of species open these glands wide when angry or excited. All deer have a
liver The liver is a major Organ (anatomy), organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the Protein biosynthesis, synthesis of proteins and biochemicals necessary for ...
without a
gallbladder In vertebrates, the gallbladder, also known as the cholecyst, is a small hollow organ where bile is stored and concentrated before it is released into the small intestine. In humans, the pear-shaped gallbladder lies beneath the liver, although ...
. Deer also have a
tapetum lucidum The ''tapetum lucidum'' ( ; ; ) is a layer of tissue in the eye of many vertebrates and some other animals. Lying immediately behind the retina, it is a retroreflector. It reflects visible light back through the retina, increasing the light a ...
, which gives them sufficiently good
night vision Night vision is the ability to see in low-light conditions, either naturally with scotopic vision or through a night-vision device. Night vision requires both sufficient spectral range and sufficient intensity range. Humans have poor night vi ...
.


Antlers

All male deer possess
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on male ...
s, with the exception of the
water deer The water deer (''Hydropotes inermis'') is a small deer superficially more similar to a musk deer than a true deer. Native to China and Korea, there are two subspecies: the Chinese water deer (''Hydropotes inermis inermis'') and the Korean water ...
, in which males have long tusk-like canines that reach below the lower jaw. Females generally lack antlers, though female reindeer bear antlers smaller and less branched than those of the males. Occasionally females in other species may develop antlers, especially in telemetacarpal deer such as European roe deer, red deer, white-tailed deer and mule deer and less often in plesiometacarpal deer. A study of antlered female white-tailed deer noted that antlers tend to be small and malformed, and are shed frequently around the time of parturition. The fallow deer and the various subspecies of the reindeer have the largest as well as the heaviest antlers, both in absolute terms as well as in proportion to body mass (an average of eight grams per kilogram of body mass); the tufted deer, on the other hand, has the smallest antlers of all deer, while the pudú has the lightest antlers with respect to body mass (0.6 g per kilogram of body mass). The structure of antlers show considerable variation; while fallow deer and elk antlers are palmate (with a broad central portion), white-tailed deer antlers include a series of tines sprouting upward from a forward-curving main beam, and those of the pudú are mere spikes. Antler development begins from the pedicel, a bony structure that appears on the top of the skull by the time the animal is a year old. The pedicel gives rise to a spiky antler the following year, that is replaced by a branched antler in the third year. This process of losing a set of antlers to develop a larger and more branched set continues for the rest of the life. The antlers emerge as soft tissues (known as
velvet antler Velvet antler is the whole cartilaginous antler in a precalcified growth stage of the Cervidae family including the species of deer such as elk, moose, and caribou. Velvet antler is covered in a hairy, velvet-like "skin" known as velvet and its t ...
s) and progressively harden into bony structures (known as hard antlers), following mineralisation and blockage of
blood vessel The blood vessels are the components of the circulatory system that transport blood throughout the human body. These vessels transport blood cells, nutrients, and oxygen to the tissues of the body. They also take waste and carbon dioxide away ...
s in the tissue, from the tip to the base. Antlers might be one of the most exaggerated male
secondary sexual characteristic Secondary sex characteristics are features that appear during puberty in humans, and at sexual maturity in other animals. These characteristics are particularly evident in the sexual dimorphism, sexually dimorphic phenotypic traits that distinguis ...
s, and are intended primarily for reproductive success through
sexual selection Sexual selection is a mode of natural selection in which members of one biological sex mate choice, choose mates of the other sex to mating, mate with (intersexual selection), and compete with members of the same sex for access to members of t ...
and for combat. The tines (forks) on the antlers create grooves that allow another male's antlers to lock into place. This allows the males to wrestle without risking injury to the face. Antlers are correlated to an individual's position in the social hierarchy and its behaviour. For instance, the heavier the antlers, the higher the individual's status in the social hierarchy, and the greater the delay in shedding the antlers; males with larger antlers tend to be more aggressive and dominant over others. Antlers can be an
honest signal Within evolutionary biology, signalling theory is a body of theoretical work examining communication between individuals, both within species and across species. The central question is when organisms with conflicting interests, such as in sex ...
of genetic quality; males with larger antlers relative to body size tend to have increased resistance to
pathogen In biology, a pathogen ( el, πάθος, "suffering", "passion" and , "producer of") in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an infectious agent, or simply a germ ...
s and higher reproductive capacity. In elk in
Yellowstone National Park Yellowstone National Park is an American national park located in the western United States, largely in the northwest corner of Wyoming and extending into Montana and Idaho. It was established by the 42nd U.S. Congress with the Yellowston ...
, antlers also provide protection against predation by
wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
. Homology of tines, that is, the branching structure of antlers among species, have been discussed before the 1900s. Recently, a new method to describe the branching structure of antlers and determining homology of tines was developed.


Teeth

Most deer bear 32 teeth; the corresponding
dental formula Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolo ...
is: . The elk and the reindeer may be exceptions, as they may retain their upper canines and thus have 34 teeth (dental formula: ). The Chinese water deer, tufted deer, and
muntjac Muntjacs ( ), also known as the barking deer or rib-faced deer, (URL is Google Books) are small deer of the genus ''Muntiacus'' native to South Asia and Southeast Asia. Muntjacs are thought to have begun appearing 15–35 million years ago, ...
have enlarged upper
canine teeth In mammalian oral anatomy, the canine teeth, also called cuspids, dog teeth, or (in the context of the upper jaw) fangs, eye teeth, vampire teeth, or vampire fangs, are the relatively long, pointed teeth. They can appear more flattened howeve ...
forming sharp tusks, while other species often lack upper canines altogether. The cheek teeth of deer have crescent ridges of enamel, which enable them to grind a wide variety of vegetation. The teeth of deer are adapted to feeding on vegetation, and like other ruminants, they lack upper
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...
s, instead having a tough pad at the front of their upper jaw.


Biology


Diet

Deer are
browsers Browse, browser or browsing may refer to: Programs *Web browser, a program used to access the World Wide Web *Code browser, a program for navigating source code *File browser or file manager, a program used to manage files and related objects *Ha ...
, and feed primarily on foliage of
grasses Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
,
sedges The Cyperaceae are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large, with some 5,500 known species described in about 90 genera, the largest being the "true sedges" genus ''Carex'' wit ...
,
forbs A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
,
shrubs A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
and
trees In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are u ...
, secondarily on
lichens A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among hypha, filaments of multiple fungus, fungi species in a mutualism (biology), mutualistic relationship.ruminant Ruminants (suborder Ruminantia) are ungulate, hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by Enteric fermentation, fermenting it in a specialized stomach prior to digestion, principally ...
standards, and high nutrition requirements. Rather than eating and digesting vast quantities of low-grade fibrous food as, for example,
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated s ...
and
cattle Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus ''Bos''. Adult females are referred to as cows and adult mal ...
do, deer select easily digestible shoots, young leaves, fresh grasses, soft twigs, fruit,
fungi A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from ...
, and
lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.calcium Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar to ...
and phosphate in order to support antler growth, and this further necessitates a nutrient-rich diet. There are some reports of deer engaging in carnivorous activity, such as eating dead
alewives The alewife (''Alosa pseudoharengus'') is an anadromous species of herring found in North America. It is one of the "typical" North American shads, attributed to the subgenus ''Pomolobus'' of the genus ''Alosa''. As an adult it is a marin ...
along lakeshores or depredating the nests of
northern bobwhite The northern bobwhite (''Colinus virginianus''), also known as the Virginia quail or (in its home range) bobwhite quail, is a ground-dwelling bird native to Canada, the United States, Mexico, and Cuba, with introduced populations elsewhere in th ...
s.


Reproduction

Nearly all cervids are so-called uniparental species: the young, known in most species as fawns, are only cared for by the mother, most often called a doe. A doe generally has one or two fawns at a time (triplets, while not unknown, are uncommon). Mating season typically begins in later August and lasts until December. Some species mate until early March. The
gestation period In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once it ...
is anywhere up to ten months for the European roe deer. Most fawns are born with their fur covered with white spots, though in many species they lose these spots by the end of their first winter. In the first twenty minutes of a fawn's life, the fawn begins to take its first steps. Its mother licks it clean until it is almost free of scent, so
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill th ...
s will not find it. Its mother leaves often to graze, and the fawn does not like to be left behind. Sometimes its mother must gently push it down with her foot.Deer – info and games
Sheppard Software.
The fawn stays hidden in the grass for one week until it is strong enough to walk with its mother. The fawn and its mother stay together for about one year. A male usually leaves and never sees his mother again, but females sometimes come back with their own fawns and form small herds.


Disease

In some areas of the UK, deer (especially
fallow deer ''Dama'' is a genus of deer in the subfamily Cervinae, commonly referred to as fallow deer. Name The name fallow is derived from the deer's pale brown colour. The Latin word ''dāma'' or ''damma'', used for roe deer, gazelles, and antelopes, ...
due to their
gregarious behaviour Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother wasp ...
) have been implicated as a possible reservoir for transmission of
bovine tuberculosis Bovines ( subfamily Bovinae) comprise a diverse group of 10 genera of medium to large-sized ungulates, including cattle, bison, African buffalo, water buffalos, and the four-horned and spiral-horned antelopes. The evolutionary relationship betw ...
, a disease which in the UK in 2005 cost £90 million in attempts to eradicate. In New Zealand, deer are thought to be important as vectors picking up ''M. bovis'' in areas where brushtail possums ''
Trichosurus vulpecula The common brushtail possum (''Trichosurus vulpecula'', from the Greek for "furry tailed" and the Latin for "little fox", previously in the genus ''Phalangista'') is a nocturnal, semiarboreal marsupial of the family Phalangeridae, native to Austr ...
'' are infected, and transferring it to previously uninfected possums when their carcasses are scavenged elsewhere. The white-tailed deer ''
Odocoileus virginianus The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...
'' has been confirmed as the sole maintenance host in the Michigan outbreak of bovine tuberculosis which remains a significant barrier to the US nationwide eradication of the disease in livestock. Moose and deer can carry
rabies Rabies is a viral disease that causes encephalitis in humans and other mammals. Early symptoms can include fever and tingling at the site of exposure. These symptoms are followed by one or more of the following symptoms: nausea, vomiting, vi ...
. Docile moose may suffer from
brain worm ''Dicrocoelium dendriticum'', the lancet liver fluke, is a parasite fluke that tends to live in cattle or other grazing mammals. History of discovery Much of what is presently known about ''Dicrocoelium dendriticum'' is the result of the wor ...
, a
helminth Parasitic worms, also known as helminths, are large macroparasites; adults can generally be seen with the naked eye. Many are intestinal worms that are soil-transmitted and infect the gastrointestinal tract. Other parasitic worms such as schi ...
which drills holes through the brain in its search for a suitable place to lay its eggs. A government biologist states that "They move around looking for the right spot and never really find it." Deer appear to be immune to this parasite; it passes through the digestive system and is excreted in the feces. The parasite is not screened by the moose intestine, and passes into the brain where damage is done that is externally apparent, both in behaviour and in gait. Deer, elk and moose in North America may suffer from
chronic wasting disease Chronic wasting disease (CWD), sometimes called zombie deer disease, is a transmissible spongiform encephalopathy (TSE) affecting deer. TSEs are a family of diseases thought to be caused by misfolded proteins called prions and include similar dis ...
, which was identified at a
Colorado Colorado (, other variants) is a state in the Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of t ...
laboratory in the 1960s and is believed to be a prion disease. Out of an abundance of caution hunters are advised to avoid contact with
specified risk material Specified risk material (SRM) is any of various tissues of ruminant animals that cannot be inspected and passed for human food because scientists have determined that BSE-causing prions concentrate there. The term was referred to in the United Ki ...
(SRM) such as the brain, spinal column or lymph nodes. Deboning the meat when butchering and sanitizing the knives and other tools used to butcher are amongst other government recommendations.


Evolution

Deer are believed to have evolved from antlerless,
tusk Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share c ...
ed ancestors that resembled modern
duiker A duiker is a small to medium-sized brown antelope native to sub-Saharan Africa, found in heavily wooded areas. The 22 extant species, including three sometimes considered to be subspecies of the other species, form the subfamily Cephalophina ...
s and diminutive deer in the early
Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...
, and gradually developed into the first antlered cervoids (the
superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...
of cervids and related extinct families) in the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
. Eventually, with the development of antlers, the tusks as well as the upper
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, whe ...
s disappeared. Thus, evolution of deer took nearly 30 million years. Biologist
Valerius Geist Valerius Geist (2 February 1938 – 6 July 2021) was a Canadian biologist and a professor emeritus in the Faculty of Environmental Design at the University of Calgary. He was a specialist on the biology, behavior, and social dynamics of North Ame ...
suggests evolution to have occurred in stages. There are not many prominent fossils to trace this evolution, but only fragments of skeletons and antlers that might be easily confused with false antlers of non-cervid species.


Eocene

The
ruminant Ruminants (suborder Ruminantia) are ungulate, hoofed herbivorous grazing or browsing mammals that are able to acquire nutrients from plant-based food by Enteric fermentation, fermenting it in a specialized stomach prior to digestion, principally ...
s, ancestors of the Cervidae, are believed to have evolved from ''
Diacodexis ''Diacodexis'' is an extinct genus of small herbivore mammals belonging to the family Dichobunidae which lived in North America, Europe and Pakistan from 55.4 mya to 46.2 mya and existing for approximately . Description ''Diacodexis'' is the ...
'', the earliest known artiodactyl (even-toed ungulate), 50–55 Mya in the Eocene. ''Diacodexis'', nearly the size of a
rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit speci ...
, featured the
talus bone The talus (; Latin for ankle or ankle bone), talus bone, astragalus (), or ankle bone is one of the group of foot bones known as the tarsus. The tarsus forms the lower part of the ankle joint. It transmits the entire weight of the body from the ...
characteristic of all modern
even-toed ungulate The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
s. This ancestor and its relatives occurred throughout North America and Eurasia, but were on the decline by at least 46 Mya. Analysis of a nearly complete skeleton of ''Diacodexis'' discovered in 1982 gave rise to speculation that this ancestor could be closer to the non-ruminants than the ruminants. '' Andromeryx'' is another prominent prehistoric ruminant, but appears to be closer to the tragulids.


Oligocene

The formation of the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
and the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
brought about significant geographic changes. This was the chief reason behind the extensive diversification of deer-like forms and the emergence of cervids from the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
to the early
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Eumeryx'' and the North American ''
Leptomeryx ''Leptomeryx'' is an extinct genus of ruminant of the family Leptomerycidae, endemic to North America during the Eocene through Oligocene 38–24.8 Mya, existing for approximately . It was a small deer-like ruminant with somewhat slender body. ...
''. The latter resembled modern-day bovids and cervids in dental morphology (for instance, it had
brachyodont The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...
molars), while the former was more advanced. Other deer-like forms included the North American ''
Blastomeryx ''Blastomeryx'' is an extinct genus of musk deer endemic to North America. It lived during the Miocene epoch 20.4—10.3 mya, existing for approximately . There may be only one species, ''Blastomeryx gemmifer''.Prothero, 2007 (p. 221-226) ''B ...
'' and the European '' Dremotherium''; these sabre-toothed animals are believed to have been the direct ancestors of all modern antlered deer, though they themselves lacked antlers. Another contemporaneous form was the four-horned
protoceratid Protoceratidae is an extinct family of herbivorous North American artiodactyls (even-toed ungulates) that lived during the Eocene through Pliocene at around 46.2—4.9 Mya, existing for about 41 million years. Classification Protoceratidae was ...
''
Protoceras ''Protoceras'' ('first horns') is an extinct genus of Artiodactyla, of the family Protoceratidae, endemic to North America. It lived from the Oligocene to the Early Miocene 33.3—16.0 Ma, existing for approximately . Morphology ''Protoceras ...
'', that was replaced by ''
Syndyoceras ''Syndyoceras'' is a small extinct genus of Artiodactyla, of the family Protoceratidae, endemic to central North America from the Miocene epoch (24.8—20.6 Ma), existing for approximately . Taxonomy ''Syndyoceras'' was named by Barbour (1 ...
'' in the Miocene; these animals were unique in having a horn on the nose. Late Eocene fossils dated approximately 35 million years ago, which were found in North America, show that ''Syndyoceras'' had bony skull outgrowths that resembled non-deciduous antlers.


Miocene

Fossil evidence suggests that the earliest members of the superfamily Cervoidea appeared in Eurasia in the Miocene. ''
Dicrocerus ''Dicrocerus elegans'' (Its name is Greek for "fork antler") is an extinct species of deer found in France, Europe (related species in Asia). ''Dicrocerus'' probably came from Asia, from the region where true deer are believed to have originated ...
'', ''
Euprox ''Euprox'' is an extinct genus of deer that lived in Eurasia during the Miocene. Taxonomy The type species ''Euprox furcatus'' was originally under the genus ''Prox'', but that name was preoccupied. Depéret assigned it to the related genus ''Di ...
'' and ''
Heteroprox ''Heteroprox'' is an extinct genus of deer from the Miocene of Europe. Description ''Heteroprox'' was a mid-sized deer that would have been similar in appearance to a muntjac Muntjacs ( ), also known as the barking deer or rib-faced deer, ( ...
'' were probably the first antlered cervids. ''Dicrocerus'' featured single-forked antlers that were shed regularly. '' Stephanocemas'' had more developed and diffuse ("crowned") antlers. ''
Procervulus ''Procervulus'' is an extinct species of deer found in Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is a ...
'' (
Palaeomerycidae The Palaeomerycidae are an extinct family of ruminants in the order Artiodactyla. Palaeomerycids lived in North America, Europe, Africa and Asia from 33 to 4.9 million years ago (from the Late Eocene to Pliocene epochs), existing for about 28 mi ...
) also possessed antlers that were not shed. Contemporary forms such as the merycodontines eventually gave rise to the modern pronghorn. The Cervinae emerged as the first group of extant cervids around 7–9 Mya, during the late Miocene in central Asia. The tribe Muntiacini made its appearance as ''
Muntiacus leilaoensis Muntjacs ( ), also known as the barking deer or rib-faced deer, (URL is Google Books) are small deer of the genus ''Muntiacus'' native to South Asia and Southeast Asia. Muntjacs are thought to have begun appearing 15–35 million years ago, ...
'' around 7–8 Mya; The early muntjacs varied in size–as small as hares or as large as fallow deer. They had tusks for fighting and antlers for defence. Capreolinae followed soon after; Alceini appeared 6.4–8.4 Mya. Around this period, the
Tethys Ocean The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...
disappeared to give way to vast stretches of grassland; these provided the deer with abundant protein-rich vegetation that led to the development of ornamental antlers and allowed populations to flourish and colonise areas. As antlers had become pronounced, the canines were either lost or became poorly represented (as in elk), probably because diet was no longer
browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
-dominated and antlers were better display organs. In muntjac and tufted deer, the antlers as well as the canines are small. The tragulids possess long canines to this day.


Pliocene

With the onset of the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58† ''
Cervocerus novorossiae ''Cervocerus'' is a deer genus that lived about 7 million years ago, at the boundary between the Miocene and Pliocene periods. ''Cervocerus novorossiae'' was found in China and the Republic of Moldova. External links ''Cervocerus'' on data.gbi ...
'', appeared around the transition from Miocene to Pliocene (4.2–6 Mya) in Eurasia; cervine fossils from early Pliocene to as late as the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
have been excavated in China and the Himalayas. While ''Cervus'' and ''Dama'' appeared nearly 3 Mya, ''Axis'' emerged during the late Pliocene–Pleistocene. The tribes Capreolini and Rangiferini appeared around 4–7 Mya. Around 5 Mya, the rangiferina ''
Bretzia ''Bretzia'' is an extinct genus of deer that was endemic to North America. Two species have been described. Taxonomy and evolution The genus ''Bretzia'' was named in 1974 by paleontologist Eric Paul Gustafson and his colleague Willis Fry. It wa ...
'' and '' Eocoileus'' were the first cervids to reach North America. This implies the Bering Strait could be crossed during the late Miocene–Pliocene; this appears highly probable as the
camelid Camelids are members of the biological family Camelidae, the only currently living family in the suborder Tylopoda. The seven extant members of this group are: dromedary camels, Bactrian camels, wild Bactrian camels, llamas, alpacas, vicuñas, ...
s migrated into Asia from North America around the same time. Deer invaded South America in the late Pliocene (2.5–3 Mya) as part of the
Great American Interchange The Great American Biotic Interchange (commonly abbreviated as GABI), also known as the Great American Interchange and the Great American Faunal Interchange, was an important late Cenozoic paleozoogeographic biotic interchange event in which lan ...
, thanks to the recently formed
Isthmus of Panama The Isthmus of Panama ( es, Istmo de Panamá), also historically known as the Isthmus of Darien (), is the narrow strip of land that lies between the Caribbean Sea and the Pacific Ocean, linking North and South America. It contains the country ...
, and emerged successful due to the small number of competing ruminants in the continent.


Pleistocene

Large deer with impressive antlers evolved during the early Pleistocene, probably as a result of abundant resources to drive evolution. The early Pleistocene cervid ''
Eucladoceros ''Eucladoceros'' (Greek for "well-branched antler") or bush-antlered deer is an extinct genus of deer whose fossils have been discovered in Europe, the Middle East and Central Asia. This genus was formally described by Hugh Falconer in 1868.Fal ...
'' was comparable in size to the modern elk. ''
Megaloceros ''Megaloceros'' (from Greek: + , literally "Great Horn"; see also Lister (1987)) is an extinct genus of deer whose members lived throughout Eurasia from the early Pleistocene to the beginning of the Holocene and were important herbivores durin ...
'' (Pliocene–Pleistocene) featured the
Irish elk The Irish elk (''Megaloceros giganteus''), also called the giant deer or Irish deer, is an extinct species of deer in the genus ''Megaloceros'' and is one of the largest deer that ever lived. Its range extended across Eurasia during the Pleisto ...
(''M. giganteus''), one of the largest known cervids. The Irish elk reached at the shoulder and had heavy antlers that spanned from tip to tip. These large animals are thought to have faced extinction due to conflict between
sexual selection Sexual selection is a mode of natural selection in which members of one biological sex mate choice, choose mates of the other sex to mating, mate with (intersexual selection), and compete with members of the same sex for access to members of t ...
for large antlers and body and
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Charle ...
for a smaller form. Meanwhile, the moose and reindeer radiated into North America from Siberia.


Taxonomy and classification

Deer constitute the
artiodactyl The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poster ...
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Cervidae. This family was first described by German zoologist
Georg August Goldfuss Georg August Goldfuss (Goldfuß, 18 April 1782 – 2 October 1848) was a German palaeontologist, zoologist and botanist. Goldfuss was born at Thurnau near Bayreuth. He was educated at Erlangen, where he graduated PhD in 1804 and became profe ...
in ''Handbuch der Zoologie'' (1820). Three
subfamilies In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end subfamily botanical names with "-oideae", and zoologi ...
were recognised: Capreolinae (first described by the English zoologist
Joshua Brookes Joshua Brookes (24 November 1761 – 10 January 1833) was a British anatomist and naturalist. Early life Brookes studied under William Hunter, William Hewson, Andrew Marshall, and John Sheldon, in London. He then attended the practice of ...
in 1828), Cervinae (described by Goldfuss) and Hydropotinae (first described by French zoologist
Édouard Louis Trouessart Édouard Louis Trouessart (25 August 1842 – 30 June 1927) was a French zoologist born in Angers. He studied military medicine in Strasbourg, but was forced to leave school due to serious health problems. In 1864 he started work as ''préparateur ...
in 1898). Other attempts at the classification of deer have been based on morphological and genetic differences. The Anglo-Irish naturalist
Victor Brooke Sir Victor Alexander Brooke, 3rd Baronet (5 January 1843 – 23 November 1891), was an Anglo-Irish sportsman- naturalist and baronet. He was the father of Field Marshal The 1st Viscount Alanbrooke, and grandfather of The 1st Viscount Brookeboro ...
suggested in 1878 that deer could be bifurcated into two classes on the according to the features of the second and fifth
metacarpal bone In human anatomy, the metacarpal bones or metacarpus form the intermediate part of the skeletal hand located between the phalanges of the fingers and the carpal bones of the wrist, which forms the connection to the forearm. The metacarpal bones ar ...
s of their forelimbs: Plesiometacarpalia (most Old World deer) and Telemetacarpalia (most New World deer). He treated the
musk deer Musk deer can refer to any one, or all seven, of the species that make up ''Moschus'', the only extant genus of the family Moschidae. Despite being commonly called deer, they are not true deer belonging to the family Cervidae, but rather their fa ...
as a cervid, placing it under Telemetacarpalia. While the telemetacarpal deer showed only those elements located far from the joint, the plesiometacarpal deer retained the elements closer to the joint as well. Differentiation on the basis of
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Sets of chromosomes refer to the number of maternal and paternal chromosome copies, respectively ...
number of
chromosome A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins are ...
s in the late 20th century has been flawed by several inconsistencies. In 1987, the zoologists
Colin Groves Colin Peter Groves (24 June 1942 – 30 November 2017) was a British-Australian biologist and anthropologist. Groves was Professor of Biological Anthropology at the Australian National University in Canberra, Australia. Education Born in Englan ...
and Peter Grubb identified three subfamilies: Cervinae, Hydropotinae and Odocoileinae; they noted that the hydropotines lack antlers, and the other two subfamilies differ in their skeletal morphology. They reverted from this classification in 2000. Molecular phylogenetic analyses since the latter half of the 2000s all show that ''
hydropotes The water deer (''Hydropotes inermis'') is a small deer superficially more similar to a musk deer than a true deer. Native to China and Korea, there are two subspecies: the Chinese water deer (''Hydropotes inermis inermis'') and the Korean water ...
'' is a sister taxon of ''
Capreolus ''Capreolus'' is a genus of deer, the roe deer. Etymology English ''roe'' is from Old English ''ra'' or ''rá'', from ''raha'', from Proto-Germanic ''*raikhaz'', cognate to Old Norse ''ra'', Old Saxon ''reho'', Middle Dutch and Dutch ''ree'' ...
'', and “Hydropotinae” became outdated subfamily.Hassanin, A., Delsuc, F., Ropiquet, A., Hammer, C., van Vuuren, B. J., Matthee, C., Ruiz-Garcia, M., Catzeflis, F., Areskoug, V., Nguyen, T. T., & Couloux, A. (2012)
Pattern and timing of diversification of Cetartiodactyla (Mammalia, Laurasiatheria), as revealed by a comprehensive analysis of mitochondrial genomes.
Comptes Rendus Biologies, 335(1), 32–50.
Heckeberg, N. S., Erpenbeck, D., Wörheide, G., & Rössner, G. E. (2016)
Systematic relationships of five newly sequenced cervid species.
PeerJ, 4, e2307.
Heckeberg, N. S. (2020)
The systematics of the Cervidae: A total evidence approach.
PeerJ, 8, e8114.


External relationships

Until 2003, it was understood that the family
Moschidae Moschidae is a family of pecoran even-toed ungulates, containing the musk deer (''Moschus'') and its extinct relatives. They are characterized by long 'saber teeth' instead of horns, antlers or ossicones, modest size (''Moschus'' only reaches ; ...
(musk deer) was
sister A sister is a woman or a girl who shares one or more parents with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to refer to ...
to Cervidae. Then a
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
study by Alexandre Hassanin (of
National Museum of Natural History, France The French National Museum of Natural History, known in French as the ' (abbreviation MNHN), is the national natural history museum of France and a ' of higher education part of Sorbonne Universities. The main museum, with four galleries, is loc ...
) and colleagues, based on
mitochondria A mitochondrion (; ) is an organelle found in the Cell (biology), cells of most Eukaryotes, such as animals, plants and Fungus, fungi. Mitochondria have a double lipid bilayer, membrane structure and use aerobic respiration to generate adenosi ...
l and
nuclear Nuclear may refer to: Physics Relating to the nucleus of the atom: * Nuclear engineering *Nuclear physics *Nuclear power *Nuclear reactor *Nuclear weapon *Nuclear medicine *Radiation therapy *Nuclear warfare Mathematics *Nuclear space *Nuclear ...
analyses, revealed that Moschidae and
Bovidae The Bovidae comprise the biological family of cloven-hoofed, ruminant mammals that includes cattle, bison, buffalo, antelopes, and caprines. A member of this family is called a bovid. With 143 extant species and 300 known extinct species, ...
form a
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
sister to Cervidae. According to the study, Cervidae diverged from the Bovidae-Moschidae clade 27 to 28 million years ago. The following
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...
is based on the 2003 study.


Internal relationships

A 2006
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
study of the internal relationships in Cervidae by Clément Gilbert and colleagues divided the family into two major clades: Capreolinae (telemetacarpal or New World deer) and Cervinae (plesiometacarpal or Old World deer). Studies in the late 20th century suggested a similar bifurcation in the family. This as well as previous studies support
monophyly In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic grou ...
in Cervinae, while Capreolinae appears
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
. The 2006 study identified two lineages in Cervinae, Cervini (comprising the genera ''
Axis An axis (plural ''axes'') is an imaginary line around which an object rotates or is symmetrical. Axis may also refer to: Mathematics * Axis of rotation: see rotation around a fixed axis *Axis (mathematics), a designator for a Cartesian-coordinate ...
'', ''
Cervus ''Cervus'' is a genus of deer that primarily are native to Eurasia, although one species occurs in northern Africa and another in North America. In addition to the species presently placed in this genus, it has included a whole range of other sp ...
'', '' Dama'' and ''
Rucervus ''Rucervus'' is a genus of deer from India, Nepal, Indochina, and the Chinese island of Hainan. The only extant representatives, the barasingha (''R. duvaucelii)'' and Eld's deer (''R. eldii''), are threatened by habitat loss and hunting, and ano ...
'') and Muntiacini (''
Muntiacus Muntjacs ( ), also known as the barking deer or rib-faced deer, (URL is Google Books) are small deer of the genus ''Muntiacus'' native to South Asia and Southeast Asia. Muntjacs are thought to have begun appearing 15–35 million years ago, ...
'' and ''
Elaphodus The tufted deer (''Elaphodus cephalophus'') is a small species of deer characterized by a prominent tuft of black hair on its forehead and fang-like canines for the males. It is a close relative of the muntjac, living somewhat further north over ...
''). Capreolinae featured three lineages, Alceini (''
Alces The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult male ...
'' species), Capreolini (''
Capreolus ''Capreolus'' is a genus of deer, the roe deer. Etymology English ''roe'' is from Old English ''ra'' or ''rá'', from ''raha'', from Proto-Germanic ''*raikhaz'', cognate to Old Norse ''ra'', Old Saxon ''reho'', Middle Dutch and Dutch ''ree'' ...
'' and the subfamily Hydropotinae) and Rangiferini (''
Blastocerus The marsh deer (''Blastocerus dichotomus'') is the largest deer species from South America reaching a length of and a shoulder height of . It is found in Argentina, Bolivia, Peru, Brazil, Uruguay and Paraguay. Formerly found in much of tropical ...
'', ''
Hippocamelus ''Hippocamelus'' is a genus of Cervidae, the deer family. It comprises two extant Andean and two fossil species. The living members are commonly known as the huemul (from the Mapuche language), and the taruca. Both species have a stocky, thick, ...
'', '' Mazama'', ''
Odocoileus ''Odocoileus'' is a genus of medium-sized deer (family Cervidae) containing three species native to the Americas The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North and South Am ...
'', ''
Pudu The pudus (Mapudungun ''püdü'' or ''püdu'', es, pudú, ) are two species of South American deer from the genus ''Pudu'', and are the world's smallest deer. The chevrotains (mouse-deer; Tragulidae) are smaller, but they are not true deer. The ...
'' and '' Rangifer'' species). The following cladogram is based on the 2006 study.


Human interaction


Prehistoric

Deer were an important source of food for early hominids. In China, ''
Homo erectus ''Homo erectus'' (; meaning "upright man") is an extinct species of archaic human from the Pleistocene, with its earliest occurrence about 2 million years ago. Several human species, such as '' H. heidelbergensis'' and '' H. antecessor' ...
'' fed upon the
sika deer The sika deer (''Cervus nippon''), also known as the Northern spotted deer or the Japanese deer, is a species of deer native to much of East Asia and introduced to other parts of the world. Previously found from northern Vietnam in the south to ...
, while the red deer was hunted in Germany. In the
Upper Palaeolithic The Upper Paleolithic (or Upper Palaeolithic) is the third and last subdivision of the Paleolithic or Old Stone Age. Very broadly, it dates to between 50,000 and 12,000 years ago (the beginning of the Holocene), according to some theories coin ...
, the reindeer was the staple food for
Cro-Magnon Early European modern humans (EEMH), or Cro-Magnons, were the first early modern humans (''Homo sapiens'') to settle in Europe, migrating from Western Asia, continuously occupying the continent possibly from as early as 56,800 years ago. They ...
people, while the
cave paintings In archaeology, Cave paintings are a type of parietal art (which category also includes petroglyphs, or engravings), found on the wall or ceilings of caves. The term usually implies prehistoric art, prehistoric origin, and the oldest known are mor ...
at
Lascaux Lascaux ( , ; french: Grotte de Lascaux , "Lascaux Cave") is a network of caves near the village of Montignac, in the department of Dordogne in southwestern France. Over 600 parietal wall paintings cover the interior walls and ceilings of ...
in southwestern France include some 90 images of stags. In
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...
, deer continued to be a main source of food for millennia even after people began farming, and it is possible that sika and other deer benefited from the frequently abandoned field sites.


Historic

Deer had a central role in the ancient art, culture and mythology of the
Hittites The Hittites () were an Anatolian people who played an important role in establishing first a kingdom in Kussara (before 1750 BC), then the Kanesh or Nesha kingdom (c. 1750–1650 BC), and next an empire centered on Hattusa in north-centra ...
, the ancient Egyptians, the
Celts The Celts (, see pronunciation for different usages) or Celtic peoples () are. "CELTS location: Greater Europe time period: Second millennium B.C.E. to present ancestry: Celtic a collection of Indo-European peoples. "The Celts, an ancien ...
, the
ancient Greeks Ancient Greece ( el, Ἑλλάς, Hellás) was a northeastern Mediterranean civilization, existing from the Greek Dark Ages of the 12th–9th centuries BC to the end of classical antiquity ( AD 600), that comprised a loose collection of cultu ...
, the Asians and several others. For instance, the
Stag Hunt Mosaic The Stag Hunt mosaic ( BC) is a mosaic from a wealthy home of the late 4th century BC, the so-called "House of the Abduction of Helen" (or "House of the Rape of Helen"), in Pella, the capital of the Macedonian Kingdom. It bears the signature of th ...
of ancient
Pella Pella ( el, Πέλλα) is an ancient city located in Central Macedonia, Greece. It is best-known for serving as the capital city of the ancient Greek kingdom of Macedon, and was the birthplace of Alexander the Great. On site of the ancient cit ...
, under the
Kingdom of Macedonia Macedonia (; grc-gre, Μακεδονία), also called Macedon (), was an ancient kingdom on the periphery of Archaic and Classical Greece, and later the dominant state of Hellenistic Greece. The kingdom was founded and initially ruled by ...
(4th century BC), possibly depicts
Alexander the Great Alexander III of Macedon ( grc, wikt:Ἀλέξανδρος, Ἀλέξανδρος, Alexandros; 20/21 July 356 BC – 10/11 June 323 BC), commonly known as Alexander the Great, was a king of the Ancient Greece, ancient Greek kingdom of Maced ...
hunting a deer with
Hephaestion Hephaestion ( grc, Ἡφαιστίων ''Hephaistíon''; c. 356 BC  –  October 324 BC), son of Amyntor, was an ancient Macedonian nobleman and a general in the army of Alexander the Great. He was "by far the dearest of all the ...
. In Japanese
Shintoism Shinto () is a religion from Japan. Classified as an East Asian religion by scholars of religion, its practitioners often regard it as Japan's indigenous religion and as a nature religion. Scholars sometimes call its practitioners ''Shintoists ...
, the sika deer is believed to be a messenger to the gods.
In China IN, In or in may refer to: Places * India (country code IN) * Indiana, United States (postal code IN) * Ingolstadt, Germany (license plate code IN) * In, Russia, a town in the Jewish Autonomous Oblast Businesses and organizations * Independ ...
, deer are associated with great medicinal significance;
deer penis In traditional Chinese medicine, a deer penis (; Vietnamese: ''Lộc pín'') is said to have important therapeutic properties. Purported properties In Angang, Taiwan, women are reported to consume deer penis during pregnancy as it is said to hav ...
is thought by some in China to have
aphrodisiac An aphrodisiac is a substance that increases sexual desire, sexual attraction, sexual pleasure, or sexual behavior. Substances range from a variety of plants, spices, foods, and synthetic chemicals. Natural aphrodisiacs like cannabis or cocain ...
properties. Spotted deer are believed in China to accompany the god of longevity. Deer was the principal sacrificial animal for the Huichal Indians of Mexico. In medieval Europe, deer appeared in hunting scenes and coats-of-arms. Deer are depicted in many materials by various pre-Hispanic civilizations in the Andes.Berrin, Katherine & Larco Museum (1997) ''The Spirit of Ancient Peru:Treasures from the Museo Arqueológico Rafael Larco Herrera.'' New York:
Thames and Hudson Thames & Hudson (sometimes T&H for brevity) is a publisher of illustrated books in all visually creative categories: art, architecture, design, photography, fashion, film, and the performing arts. It also publishes books on archaeology, history, ...
, .
The common male first name ''
Oscar Oscar, OSCAR, or The Oscar may refer to: People * Oscar (given name), an Irish- and English-language name also used in other languages; the article includes the names Oskar, Oskari, Oszkár, Óscar, and other forms. * Oscar (Irish mythology), ...
'' is taken from the
Irish Language Irish ( Standard Irish: ), also known as Gaelic, is a Goidelic language of the Insular Celtic branch of the Celtic language family, which is a part of the Indo-European language family. Irish is indigenous to the island of Ireland and was ...
, where it is derived from two elements: the first, ''os'', means "deer"; the second element, ''cara'', means "friend". The name is borne by a famous hero of
Irish mythology Irish mythology is the body of myths native to the island of Ireland. It was originally passed down orally in the prehistoric era, being part of ancient Celtic religion. Many myths were later written down in the early medieval era by C ...
Oscar Oscar, OSCAR, or The Oscar may refer to: People * Oscar (given name), an Irish- and English-language name also used in other languages; the article includes the names Oskar, Oskari, Oszkár, Óscar, and other forms. * Oscar (Irish mythology), ...
, grandson of
Fionn Mac Cumhail Fionn mac Cumhaill ( ; Old and mga, Find or ''mac Cumail'' or ''mac Umaill''), often anglicized Finn McCool or MacCool, is a hero in Irish mythology, as well as in later Scottish and Manx folklore. He is leader of the ''Fianna'' bands of ...
. The name was popularised in the 18th century by
James Macpherson James Macpherson (Gaelic: ''Seumas MacMhuirich'' or ''Seumas Mac a' Phearsain''; 27 October 1736 – 17 February 1796) was a Scottish writer, poet, literary collector and politician, known as the "translator" of the Ossian cycle of epic poem ...
, creator of 'Ossianic poetry'.


Literary

Deer have been an integral part of fables and other literary works since the inception of writing. Stags were used as symbols in the latter Sumerian writings. For instance, the boat of Sumerian god Enki is named the ''Stag of Azbu''. There are several mentions of the animal in the
Rigveda The ''Rigveda'' or ''Rig Veda'' ( ', from ' "praise" and ' "knowledge") is an ancient Indian collection of Vedic Sanskrit hymns (''sūktas''). It is one of the four sacred canonical Hindu texts (''śruti'') known as the Vedas. Only one Sh ...
as well as the
Bible The Bible (from Koine Greek , , 'the books') is a collection of religious texts or scriptures that are held to be sacred in Christianity, Judaism, Samaritanism, and many other religions. The Bible is an anthologya compilation of texts of a ...
. In the Indian epic
Ramayana The ''Rāmāyana'' (; sa, रामायणम्, ) is a Sanskrit literature, Sanskrit Indian epic poetry, epic composed over a period of nearly a millennium, with scholars' estimates for the earliest stage of the text ranging from the 8th ...
,
Sita Sita (; ) also called as Janaki and Vaidehi is a Hindu goddess and the female protagonist of the Hindu epic, ''Ramayana''. She is the consort of Rama, the avatar of the god Vishnu, and is regarded as a form of Vishnu's consort, Lakshmi. She ...
is lured by a golden deer which
Rama Rama (; ), Ram, Raman or Ramar, also known as Ramachandra (; , ), is a major deity in Hinduism. He is the seventh and one of the most popular '' avatars'' of Vishnu. In Rama-centric traditions of Hinduism, he is considered the Supreme Bein ...
tries to catch. In the absence of both Rama and
Lakshman Lakshmana ( sa, लक्ष्मण, lit=the fortunate one, translit=Lakṣmaṇa), also spelled as Laxmana, is the younger brother of Rama and his loyalist in the Hindu epic ''Ramayana''. He bears the epithets of Saumitra () and Ramanuja (). ...
,
Ravana Ravana (; , , ) is a rakshasa king of the island of Lanka, and the chief antagonist of the Hindu epic ''Ramayana'' and its adaptations. In the ''Ramayana'', Ravana is described to be the eldest son of sage Vishrava and rakshasi Kaikesi. He a ...
kidnaps Sita. Many of the allegorical
Aesop's fables Aesop's Fables, or the Aesopica, is a collection of fables credited to Aesop, a slave and storyteller believed to have lived in ancient Greece between 620 and 564 BCE. Of diverse origins, the stories associated with his name have descended to ...
, such as "The Stag at the Pool", "The One-Eyed Doe" and "The Stag and a Lion", personify deer to give moral lessons. For instance, "The Sick Stag" gives the message that uncaring friends can do more harm than good. The
Yaqui The Yaqui, Hiaki, or Yoeme, are a Native American people of the southwest, who speak a Uto-Aztecan language. Their homelands include the Río Yaqui valley in Sonora, Mexico, and the area below the Gila River in Arizona, Southwestern United Stat ...
deer song accompanies the deer dance which is performed by a pascola
rom the Spanish 'pascua', Easter Rom, or ROM may refer to: Biomechanics and medicine * Risk of mortality, a medical classification to estimate the likelihood of death for a patient * Rupture of membranes, a term used during pregnancy to describe a rupture of the amniotic sac * R ...
dancer (also known as a deer dancer). Pascolas would perform at religious and social functions many times of the year, especially during Lent and Easter. In one of
Rudolf Erich Raspe Rudolf Erich Raspe (March 1736 – 16 November 1794) was a German librarian, writer, and scientist, called by his biographer John Patrick Carswell a "rogue". He is best known for his collection of tall tales '' The Surprising Adventures of Bar ...
's 1785 stories of ''
Baron Munchausen Baron Munchausen (; ) is a fictional German nobleman created by the German writer Rudolf Erich Raspe in his 1785 book '' Baron Munchausen's Narrative of his Marvellous Travels and Campaigns in Russia''. The character is loosely based on a real ...
's Narrative of his Marvellous Travels and Campaigns in Russia'', the baron encounters a stag while eating cherries and, without ammunition, fires the cherry-pits at the stag with his musket, but it escapes. The next year, the baron encounters a stag with a
cherry tree A cherry is the fruit of many plants of the genus ''Prunus'', and is a fleshy drupe (stone fruit). Commercial cherries are obtained from cultivars of several species, such as the sweet ''Prunus avium'' and the sour ''Prunus cerasus''. The nam ...
growing from its head; presumably this is the animal he had shot at the previous year. In
Christmas Christmas is an annual festival commemorating Nativity of Jesus, the birth of Jesus, Jesus Christ, observed primarily on December 25 as a religious and cultural celebration among billions of people Observance of Christmas by country, around t ...
lore (such as in the narrative poem "
A Visit from St. Nicholas ''A Visit from St. Nicholas'', more commonly known as ''The Night Before Christmas'' and ''Twas the Night Before Christmas'' from its first line, is a poem first published anonymously under the title ''Account of a Visit from St. Nicholas'' i ...
"),
reindeer Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subspe ...
are often depicted pulling the
sleigh A sled, skid, sledge, or sleigh is a land vehicle that slides across a surface, usually of ice or snow. It is built with either a smooth underside or a separate body supported by two or more smooth, relatively narrow, longitudinal runners s ...
of
Santa Claus Santa Claus, also known as Father Christmas, Saint Nicholas, Saint Nick, Kris Kringle, or simply Santa, is a Legend, legendary figure originating in Western Christianity, Western Christian culture who is said to Christmas gift-bringer, bring ...
.
Marjorie Kinnan Rawlings Marjorie Kinnan Rawlings (August 8, 1896 – December 14, 1953)
accessed December 8, 2014.
was an
's
Pulitzer Prize The Pulitzer Prize () is an award for achievements in newspaper, magazine, online journalism, literature, and musical composition within the United States. It was established in 1917 by provisions in the will of Joseph Pulitzer, who had made h ...
-winning 1938 novel ''
The Yearling ''The Yearling'' is a novel by American writer Marjorie Kinnan Rawlings, published in March 1938. It was the main selection of the Book of the Month Club in April 1938. It won the 1939 Pulitzer Prize for the Novel. It was the best-selling no ...
'' was about a boy's relationship with a baby deer. The fiction book ''
Fire Bringer ''Fire Bringer'' is a young adult fiction, young adult fantasy novel by David Clement-Davies published in 1999, in the United Kingdom and 2000, in the United States. It tells the story of Rannoch, a red deer whose life is the subject of an old p ...
'' is about a young fawn who goes on a quest to save the Herla, the deer kind. In the 1942
Walt Disney Pictures Walt Disney Pictures is an American film production company and subsidiary of Walt Disney Studios, which is owned by The Walt Disney Company. The studio is the flagship producer of live-action feature films within the Walt Disney Studios uni ...
film, ''
Bambi ''Bambi'' is a 1942 American animated drama film directed by David Hand (supervising a team of sequence directors), produced by Walt Disney and based on the 1923 book ''Bambi, a Life in the Woods'' by Austrian author and hunter Felix Salten. ...
'' is a
white-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known as the whitetail or Virginia deer, is a medium-sized deer native to North America, Central America, and South America as far south as Peru and Bolivia. It has also been introduced t ...
, while in
Felix Salten Felix Salten (; 6 September 1869 – 8 October 1945) was an Austro-Hungarian author and literary critic in Vienna. Life and death Salten was born Siegmund Salzmann on 6 September 1869 in Pest, Austria-Hungary. His father was Fülöp Salzmann, ...
's original 1923 book ''
Bambi, a Life in the Woods ''Bambi, a Life in the Woods'' (German title: ''Bambi: Eine Lebensgeschichte aus dem Walde'') is a 1923 Austrian literature, Austrian coming-of-age novel written by Felix Salten and originally published in Berlin by Ullstein Verlag. The novel tra ...
'', he is a
roe deer The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
. In
C. S. Lewis Clive Staples Lewis (29 November 1898 – 22 November 1963) was a British writer and Anglican lay theologian. He held academic positions in English literature at both Oxford University (Magdalen College, 1925–1954) and Cambridge Univers ...
's 1950 fantasy novel ''
The Lion, the Witch and the Wardrobe ''The Lion, the Witch and the Wardrobe'' is a fantasy novel for children by C. S. Lewis, published by Geoffrey Bles in 1950. It is the first published and best known of seven novels in ''The Chronicles of Narnia'' (1950–1956). Among all the ...
'' the adult Pevensies, now kings and queens of
Narnia ''The Chronicles of Narnia'' is a series of seven high fantasy novels by British author C. S. Lewis. Illustrated by Pauline Baynes and originally published between 1950 and 1956, ''The Chronicles of Narnia'' has been adapted for radio, telev ...
, chase the White Stag on a hunt, as the Stag is said to grant its captor a wish. The hunt is key in returning the Pevensies to their home in England. In the 1979 book ''
The Animals of Farthing Wood The Animals of Farthing Wood is a series of books about a group of woodland animals. It originated with the 1979 book, ''The Animals of Farthing Wood'', by Colin Dann, and was followed by six sequels and a prequel by Dann. An animated ''Anim ...
'', The Great White Stag is the leader of all the animals.


Heraldic

Deer of various types appear frequently in European
heraldry Heraldry is a discipline relating to the design, display and study of armorial bearings (known as armory), as well as related disciplines, such as vexillology, together with the study of ceremony, rank and pedigree. Armory, the best-known branch ...
. In the British armory, the term "stag" is typically used to refer to antlered male red deer, while "buck" indicates an antlered male fallow deer. Stags and bucks appear in a number of attitudes, referred to as "lodged" when the deer is lying down, "trippant" when it has one leg raised, "courant" when it is running, "springing" when in the act of leaping, "statant" when it is standing with all hooves on the ground and looking ahead, and "at gaze" when otherwise statant but looking at the viewer. Stags' heads are also frequently used; these are typically portrayed without an attached neck and as facing the viewer, in which case they are termed "caboshed".
Arthur Fox-Davies Arthur Charles Fox-Davies (28 February 1871 – 19 May 1928) was a British expert on heraldry. His ''Complete Guide to Heraldry'', published in 1909, has become a standard work on heraldry in England. A barrister by profession, Fox-Davies worke ...

''A Complete Guide to Heraldry''
T.C. and E.C. Jack, London, 1909, 208–210,
Examples of deer in
coats of arms A coat of arms is a heraldic visual design on an escutcheon (i.e., shield), surcoat, or tabard (the latter two being outer garments). The coat of arms on an escutcheon forms the central element of the full heraldic achievement, which in its wh ...
can be found in the arms of
Hertfordshire Hertfordshire ( or ; often abbreviated Herts) is one of the home counties in southern England. It borders Bedfordshire and Cambridgeshire to the north, Essex to the east, Greater London to the south, and Buckinghamshire to the west. For govern ...
, England, and its county town of
Hertford Hertford ( ) is the county town of Hertfordshire, England, and is also a civil parish in the East Hertfordshire district of the county. The parish had a population of 26,783 at the 2011 census. The town grew around a ford on the River Lea, ne ...
; both are examples of
canting arms Canting arms are heraldic bearings that represent the bearer's name (or, less often, some attribute or function) in a visual pun or rebus. French heralds used the term (), as they would sound out the name of the armiger. Many armorial allus ...
. A deer appears on the arms of the Israeli Postal Authority. Coats of arms featuring deer include those of
Dotternhausen Dotternhausen is a municipality in the Zollernalbkreis district, in Baden-Württemberg, Germany. History Dotternhausen became a possession of the Kingdom of Württemberg in 1805 and was assigned to . In 1810, it was reassigned to and then to in ...
,
Thierachern Thierachern is a municipality in the administrative district of Thun in the canton of Bern in Switzerland. History Thierachern is first mentioned in 1250 as ''Tierachern''. The area around Thierachern was occupied during the neolithic era, the ...
,
Friolzheim Friolzheim is a municipality of the Enz district in Baden-Württemberg, Germany. The Friolzheimer Riese telecommunications tower is located here. History The village of Friolzheim gradually became a possession of the in the 15th century, but ...
,
Bauen Bauen is a former municipality on Lake Lucerne in the Swiss canton of Uri. On 1 January 2021 the former municipality of Bauen merged into the municipality of Seedorf. History Bauen is first mentioned in 1150 as ''Bawen''. The municipalities of ...
,
Albstadt Albstadt () is the largest city in the district of Zollernalbkreis in Baden-Württemberg, Germany. It is located on the Swabian Jura mountains, about halfway between Stuttgart and Lake Constance. Geography Albstadt is spread across a variety of ...
, and
Dassel Dassel is a town in southern Lower Saxony, Germany, located in the district Northeim. It is located near the hills of the Solling mountains. Geography The city covers an area of . Buildings and streets make up about 10% of this area while 26% a ...
in Germany; of the
Earls Bathurst Earl Bathurst, of Bathurst in the County of Sussex, is a title in the Peerage of Great Britain. The medieval English word was Botehurst, thought to date at least from the 13th century. Bote is the origination of Battle, although the family m ...
in England; of
Balakhna Balakhna (russian: Балахна́) is a town and the administrative center of Balakhninsky District in Nizhny Novgorod Oblast, Russia, located on the right bank of the Volga River, north of Nizhny Novgorod, the administrative center of the obl ...
, Russia; of
Åland Åland ( fi, Ahvenanmaa: ; ; ) is an Federacy, autonomous and Demilitarized zone, demilitarised region of Finland since 1920 by a decision of the League of Nations. It is the smallest region of Finland by area and population, with a size of 1 ...
, Finland; of
Gjemnes Gjemnes is a municipality in Møre og Romsdal county, Norway on the Romsdal peninsula. It is part of the Nordmøre region. The administrative centre is the village of Batnfjordsøra, which lies along the Batnfjorden and it is a former steamship ...
,
Hitra Hitra is a municipality in Trøndelag county, Norway. The municipality covers the island of Hitra and hundreds smaller islands, islets, and skerries. It is part of the Fosen region. The administrative centre of the municipality is the village ...
,
Hjartdal Hjartdal is a municipality in Telemark in the county of Vestfold og Telemark in Norway. It is part of the traditional regions of Upper Telemark and Øst-Telemark. The administrative centre of the municipality is the village of Sauland. The munic ...
,
Rendalen Rendalen is a municipality in Innlandet county, Norway. It is located in the traditional district of Østerdalen. The administrative centre of the municipality is the village of Bergset. Other villages in the municipality include Hanestad, Ot ...
and
Voss Voss () is a municipality and a traditional district in Vestland county, Norway. The administrative center of the municipality is the village of Vossevangen. Other villages include Bolstadøyri, Borstrondi, Evanger, Kvitheim, Mjølfjell, Opphe ...
in Norway; of
Jelenia Góra Jelenia Góra (pron. ; Polish: ; german: Hirschberg im Riesengebirge; Exonym: ''Deer Mountain''; szl, Jelyniŏ Gōra) is a historic city in southwestern Poland, within the historical region of Lower Silesia. Jelenia Góra is situated in the Low ...
, Poland; of
Umeå Umeå ( , , , locally ; South Westrobothnian: ;). fi, Uumaja; sju, Ubmeje; sma, Upmeje; se, Ubmi) is a city in northeast Sweden. It is the seat of Umeå Municipality and the capital of Västerbotten County. Situated on the Ume River, Ume ...
, Sweden; of
Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...
, Australia; of
Cervera Cervera () is the capital of the ''comarca'' of Segarra, in the province of Lleida, Autonomous Community of Catalonia, Spain. The title Comte de Cervera is a courtesy title, formerly part of the Crown of Aragon, that has been revived for Leonor, ...
, Catalonia; of Northern Ireland; and of Chile. Other types of deer used in heraldry include the hind, portrayed much like the stag or buck but without antlers, as well as the reindeer and winged stags. Winged stags are used as
supporter In heraldry, supporters, sometimes referred to as ''attendants'', are figures or objects usually placed on either side of the shield and depicted holding it up. Early forms of supporters are found in medieval seals. However, unlike the coro ...
s in the arms of the
de Carteret family The de Carteret family was perhaps the greatest of the patrician families of the Channel Islands. Their influence on the Island would last from the 10th century until the present time. Origins The family originated from Normandy where their ance ...
. The sea-stag, possessing the antlers, head, forelegs and upper body of a stag and the tail of a
mermaid In folklore, a mermaid is an aquatic creature with the head and upper body of a female human and the tail of a fish. Mermaids appear in the folklore of many cultures worldwide, including Europe, Asia, and Africa. Mermaids are sometimes asso ...
, is often found in German heraldry.


Economic

Deer have long had economic significance to humans. Deer meat, known as
venison Venison originally meant the meat of a game animal but now refers primarily to the meat of antlered ungulates such as elk or deer (or antelope in South Africa). Venison can be used to refer to any part of the animal, so long as it is edible, in ...
, is highly nutritious. Due to the inherently wild nature and diet of deer, venison is most often obtained through deer hunting. In the United States, it is produced in small amounts compared to
beef Beef is the culinary name for meat from cattle (''Bos taurus''). In prehistoric times, humankind hunted aurochs and later domesticated them. Since that time, numerous breeds of cattle have been bred specifically for the quality or quantity ...
, but still represents a significant trade. Deer hunting is a popular activity in the U.S. that can provide the hunter's family with high quality meat and generates revenue for states and the federal government from the sales of licenses, permits and tags. The 2006 survey by the
U.S. Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with oth ...
estimates that license sales generate approximately $700 million annually. This revenue generally goes to support conservation efforts in the states where the licenses are purchased. Overall, the U.S. Fish and Wildlife Service estimates that big game hunting for deer and elk generates approximately $11.8 billion annually in hunting-related travel, equipment and related expenditures. Conservation laws prevent the sale of unlicensed wild game meat, although it may be donated. Deer have often been bred in captivity as ornaments for parks, but only in the case of reindeer has thorough domestication succeeded. By 2012, some 25,000 tons of red deer were raised on farms in North America. The
Sami Acronyms * SAMI, ''Synchronized Accessible Media Interchange'', a closed-captioning format developed by Microsoft * Saudi Arabian Military Industries, a government-owned defence company * South African Malaria Initiative, a virtual expertise net ...
of Scandinavia and the
Kola Peninsula sjd, Куэлнэгк нёа̄ррк , image_name= Kola peninsula.png , image_caption= Kola Peninsula as a part of Murmansk Oblast , image_size= 300px , image_alt= , map_image= Murmansk in Russia.svg , map_caption = Location of Murmansk Oblas ...
of Russia and other nomadic peoples of northern Asia use reindeer for food, clothing, and transport. Others are bred for hunting are selected based on the size of the antlers. The major deer-producing countries are New Zealand, the market leader, with Ireland, Great Britain and Germany. The trade earns over $100 million annually for these countries. Automobile collisions with deer can impose a significant cost on the economy. In the U.S., about 1.5 million deer-vehicle collisions occur each year, according to the
National Highway Traffic Safety Administration The National Highway Traffic Safety Administration (NHTSA ) is an agency of the U.S. federal government, part of the Department of Transportation. It describes its mission as "Save lives, prevent injuries, reduce vehicle-related crashes" rela ...
. Those accidents cause about 150 human deaths and $1.1 billion in property damage annually. In Scotland, several roads including the A82, the A87 and the
A835 The A835 is a road in the Scottish Highlands linking Inverness to Ullapool and the Far North of Scotland. Route The A835 starts at Tore on the Black Isle, seven miles north-west of Inverness at a junction with the A9. The A835 crosses the Bla ...
have had significant enough problems with ''deer vehicle collisions'' (DVCs) that sets of vehicle activated automatic warning signs have been installed along these roads. The skins make a peculiarly strong, soft leather, known as buckskin. There is nothing special about skins with the fur still on since the hair is brittle and soon falls off. The hooves and antlers are used for ornamental purposes, especially the antlers of the
roe deer The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
, which are utilized for making umbrella handles, and for similar purposes; elk antlers is often employed in making knife handles. Among the
Inuit Inuit (; iu, ᐃᓄᐃᑦ 'the people', singular: Inuk, , dual: Inuuk, ) are a group of culturally similar indigenous peoples inhabiting the Arctic and subarctic regions of Greenland, Labrador, Quebec, Nunavut, the Northwest Territories ...
, the traditional ''
ulu An ulu ( iu, ᐅᓗ, plural: ''uluit'', 'woman's knife') is an all-purpose knife traditionally used by Inuit, Iñupiat, Yupik, and Aleut women. It is utilized in applications as diverse as skinning and cleaning animals, cutting a child's hair, cu ...
'' women's knife was made with an antler or ivory handle. In China, a
traditional chinese medicine Traditional Chinese medicine (TCM) is an alternative medical practice drawn from traditional medicine in China. It has been described as "fraught with pseudoscience", with the majority of its treatments having no logical mechanism of action ...
is made from stag antler, and the antlers of certain species are eaten when "in the velvet". Antlers can also be boiled down to release the protein gelatin, which is used as a topical treatment for skin irritation and is also used in cooking. Since the early 20th century, deer have become commonly thought of as pests in New Zealand due to a lack of predators on the island causing population numbers to increase and begin encroaching on more populated areas. They compete with livestock for resources, as well as cause excess erosion and wreak havoc on wild plant species and agriculture alike. They can also have an effect on the conservation efforts of other plant and animal species, as they can critically offset the balance within an environment by drastically depleting diversity within forests.


See also

*
Australian Deer Association The Australian Deer Association Inc (ADA) is an Australian not-for-profit association incorporated in South Australia with branches in all states and territories. The Australian Deer Association was established on September 5, 1969 (first named as ...
*
Deer forest The deer forest (Gaelic: frìth) is a sporting estate which is kept and managed largely or solely for the purposes of maintaining a resident population of red deer for sporting (deer stalking) purposes. It is an institution and phenomenon peculi ...
*
Deer management Deer management is the practice and philosophy of Wildlife Management, wildlife management employed to regulate the population of deer in an area. The purpose of deer management is to regulate the population's size, reduce negative effects of popula ...
*
Largest cervids Cervids are one of the most common wild herbivores of the world. Of these moose can grow up to 2.33 m tall and weigh as much as 820 kg. The smallest of them all is the northern pudu. See also * List of cervids Cervidae is a family of hoofed ...
*
Reindeer hunting in Greenland Reindeer hunting in Greenland is of great importance to the Greenlandic Inuit and sports hunters, both residents and tourists.


References


Further reading

* ''Deerland: America's Hunt for Ecological Balance and the Essence of Wildness'' by Al Cambronne, Lyons Press (2013),


External links


Family Cervidae
at the
Animal Diversity Web Animal Diversity Web (ADW) is an online database that collects the natural history, classification, species characteristics, conservation biology, and distribution information on thousands of species of animals. The website includes thousands ...

Chronic Wasting Disease InformationWorld of Deer Museum


* {{Authority control Articles containing video clips Extant Rupelian first appearances Livestock Mammal common names Mammal families Taxa named by Georg August Goldfuss