''Danielsraptor'' (meaning "Michael Daniels's thief") is an extinct genus of

masillaraptorid Masillaraptoridae is an extinct family of stem-group falconiform birds from the Eocene of Europe. They are noted for their relatively long legs. Two genera have been named: '' Danielsraptor'', from the London Clay of England, and ''Masillaraptor' ...

bird from the

Early Eocene In the geologic timescale the Ypresian is the oldest age or lowest stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by the Eocene Lutetian Age. The Ypresian i ...

(

Ypresian In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...

) Walton Member of the London Clay Formation in

Essex Essex () is a county in the East of England. One of the home counties, it borders Suffolk and Cambridgeshire to the north, the North Sea to the east, Hertfordshire to the west, Kent across the estuary of the River Thames to the south, and G ...

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...

. The genus contains a single species, ''D. phorusrhacoides'', known from a partial skeleton.

Discovery and naming

The ''Danielsraptor'' fossil material was discovered in layers of the London Clay Formation (Walton Member), dated to the early Ypresian. This formation is located near

Walton-on-the-Naze Walton-on-the-Naze is a seaside town on the North Sea coast and (as Walton le Soken) a former civil parish, now in the parish of Frinton and Walton, in the Tendring district in Essex, England. It is north of Clacton and south of the port of Ha ...

England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...

. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

, NMS.Z.2021.40.12, was collected in 1991 by Michael Daniels (who the taxon is named after). This specimen consists of a partial skeleton including the

skull The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...

vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic i ...

, fragmentary wing bones, elements of the

pectoral girdle The shoulder girdle or pectoral girdle is the set of bones in the appendicular skeleton which connects to the arm on each side. In humans it consists of the clavicle and scapula; in those species with three bones in the shoulder, it consists of t ...

pelvis The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton). The ...

, the majority of the legs, and various bone fragments still partially enclosed in

matrix Matrix most commonly refers to: * ''The Matrix'' (franchise), an American media franchise ** ''The Matrix'', a 1999 science-fiction action film ** "The Matrix", a fictional setting, a virtual reality environment, within ''The Matrix'' (franchis ...

. An additional referred specimen, NMS.Z.2021.40.13, was found by Daniels in 1986. This specimen includes a partial

sternum The sternum or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major blood vessels from injury. Sh ...

, right

coracoid A coracoid (from Greek κόραξ, ''koraks'', raven) is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is prese ...

scapula The scapula (plural scapulae or scapulas), also known as the shoulder blade, is the bone that connects the humerus (upper arm bone) with the clavicle (collar bone). Like their connected bones, the scapulae are paired, with each scapula on eithe ...

furcula The (Latin for "little fork") or wishbone is a forked bone found in most birds and some species of non-avian dinosaurs, and is formed by the fusion of the two pink clavicles. In birds, its primary function is in the strengthening of the thoracic ...

, and

pygostyle Pygostyle describes a skeletal condition in which the final few caudal vertebrae are fused into a single ossification, supporting the tail feathers and musculature. In modern birds, the rectrices attach to these. The pygostyle is the main compon ...

. In 2022,

Gerald Mayr Gerald Mayr is a German palaeontologist who is Curator of Ornithology at the Senckenberg Research Institute in Frankfurt am Main, Hesse. He has published extensively on fossil birds, especially the Paleogene avifauna of Europe. He is an expert on ...

and Andrew C. Kitchener described ''Danielsraptor phorusrhacoides'', a new genus and species of masillaraptorid, based on these fossil remains. The generic name, "''Danielsraptor''", honours Michael Daniels, who collected and studied bird fossils from the type locality. This is combined with the

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

"raptor", meaning "thief", in reference to a

vernacular A vernacular or vernacular language is in contrast with a "standard language". It refers to the language or dialect that is spoken by people that are inhabiting a particular country or region. The vernacular is typically the native language, n ...

term for

birds of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predators ...

. The specific name, "''phorusrhacoides''", references the extinct "terror bird" clade

Phorusrhacidae Phorusrhacids, colloquially known as terror birds, are an extinct clade of large carnivorous flightless birds that were one of the largest species of apex predators in South America during the Cenozoic era; their conventionally accepted temporal ...

, whose members had a similar bill shape.

Description

The describing authors concluded that ''Danielsraptor'' had a body shape very similar to the

middle Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn ...

Masillaraptor ''Masillaraptor'' is an extinct genus of masillaraptorid, a groups basal falconiforms, from the Middle Eocene Messel Pit, Germany. It is a long-legged relative of the living falcons. Known species Only one species of ''Masillaraptor'' is kno ...

'' of

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

. It had long legs, which may indicate that it had a

terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...

lifestyle, foraging on the ground similar to modern caracaras. It had a large pygostyle, indicating that it would have likely had long tail feathers. This, in addition to its long ulna, suggests that it was capable of well-developed flight. Its beak was similar to those of extinct phorusrhacids and extant caracaras.

Classification

''Danielsraptor'' represents one of two members of Masillaraptoridae within the Falconiformes. The

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ...

below displays the results of the

phylogenetic analysis In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

by Mayr & Kitchener (2022):

References

{{reflist Falconiformes Eocene birds Extinct birds of Europe Prehistoric bird genera Birds described in 2022 Fossil taxa described in 2022

Discovery and naming

Description

Classification

See also

References