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The domestic donkey is a hoofed mammal in the family
Equidae Equidae (sometimes known as the horse family) is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. All extant species are in the genus '' Equus'', ...
, the same family as the
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million y ...
. It derives from the
African wild ass The African wild ass (''Equus africanus'') or African wild donkey is a wild member of the horse family, Equidae. This species is thought to be the ancestor of the domestic donkey (''Equus asinus''), which is sometimes placed within the same spec ...
, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a separate
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, ''Equus asinus''. It was domesticated in Africa some years ago, and has been used mainly as a
working animal A working animal is an animal, usually domesticated, that is kept by humans and trained to perform tasks instead of being slaughtered to harvest animal products. Some are used for their physical strength (e.g. oxen and draft horses) or for ...
since that time. There are more than 40 million donkeys in the world, mostly in
underdeveloped Underdevelopment, in the context of international development, reflects a broad condition or phenomena defined and critiqued by theorists in fields such as economics, development studies, and postcolonial studies. Used primarily to distinguish st ...
countries, where they are used principally as draught or
pack animals A pack animal, also known as a sumpter animal or beast of burden, is an individual or type of working animal used by humans as means of transporting materials by attaching them so their weight bears on the animal's back, in contrast to draft ani ...
. While working donkeys are often associated with those living at or below subsistence, small numbers of donkeys or asses are kept for breeding or as pets in developed countries. A male donkey is known as a ''jack'' or ''jackass'', a female is a ''jenny'' or ''jennet'', and an immature donkey of either sex is a '' foal''. Jacks are often mated with female horses (mares) to produce '' mules''; the less common hybrid of a male horse (stallion) and jenny is a ''
hinny A hinny is a domestic equine hybrid, the offspring of a male horse (a stallion) and a female donkey (a jenny). It is the reciprocal cross to the more common mule, which is the product of a male donkey (a jack) and a female horse (a mare). The ...
''.


Nomenclature

Traditionally, the scientific name for the donkey is ''Equus asinus asinus'' based on the principle of priority used for scientific names of animals. However, the International Commission on Zoological Nomenclature ruled in 2003 that if the domestic species and the wild species are considered subspecies of one another, the scientific name of the wild species has priority, even when that subspecies was described after the domestic subspecies. This means that the proper scientific name for the donkey is ''Equus africanus asinus'' when it is considered a subspecies and ''Equus asinus'' when it is considered a species. At one time, the synonym ''ass'' was the more common term for the donkey. The first recorded use of ''donkey'' was in either 1784 or 1785. While the word ''ass'' has cognates in most other
Indo-European languages The Indo-European languages are a language family native to the overwhelming majority of Europe, the Iranian plateau, and the northern Indian subcontinent. Some European languages of this family, English, French, Portuguese, Russian, Dutc ...
, ''donkey'' is an
etymologically Etymology ()The New Oxford Dictionary of English (1998) – p. 633 "Etymology /ˌɛtɪˈmɒlədʒi/ the study of the class in words and the way their meanings have changed throughout time". is the study of the history of the form of words an ...
obscure word for which no credible cognate has been identified. Hypotheses on its derivation include the following: * perhaps from Spanish for its
don Don, don or DON and variants may refer to: Places *County Donegal, Ireland, Chapman code DON *Don (river), a river in European Russia *Don River (disambiguation), several other rivers with the name *Don, Benin, a town in Benin *Don, Dang, a vill ...
-like gravity; the donkey was also known as "the King of Spain's trumpeter". * perhaps a diminutive of ''
dun A dun is an ancient or medieval fort. In Ireland and Britain it is mainly a kind of hillfort and also a kind of Atlantic roundhouse. Etymology The term comes from Irish ''dún'' or Scottish Gaelic ''dùn'' (meaning "fort"), and is cognat ...
'' (dull grayish-brown), a typical donkey colour. * perhaps from the name ''Duncan''. * perhaps of imitative origin. From the 18th century, ''donkey'' gradually replaced ''ass'' and ''jenny'' replaced ''she-ass'', which is now considered archaic. The change may have come about through a tendency to avoid pejorative terms in speech and may be comparable to the substitution in North American English of ''
rooster The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adult m ...
'' for ''cock'', or that of ''rabbit'' for ''coney'', which was formerly
homophonic In music, homophony (;, Greek: ὁμόφωνος, ''homóphōnos'', from ὁμός, ''homós'', "same" and φωνή, ''phōnē'', "sound, tone") is a texture in which a primary part is supported by one or more additional strands that flesh ...
with ''cunny'' (a variation of the word cunt). By the end of the 17th century, changes in pronunciation of both ''ass'' and ''
arse Arse or ARSE may refer to: * A Commonwealth English slang term for the buttocks * Arse, the name for the Iberian settlement of modern-day Sagunto * Arse (district), a district in the South Tapanuli Regency, North Sumatra province, Sumatra, Indone ...
'' had caused them to become homophones in some varieties of English. Other words used for the ass in English from this time include ''cuddy'' in Scotland, ''neddy'' in southwestern England and ''dicky'' in southeastern England; ''moke'' is documented in the 19th century and may be of Welsh or Romani origin. ''Burro'' is a word for donkey in both Spanish and
Portuguese Portuguese may refer to: * anything of, from, or related to the country and nation of Portugal ** Portuguese cuisine, traditional foods ** Portuguese language, a Romance language *** Portuguese dialects, variants of the Portuguese language ** Portu ...
. In the United States, it is commonly applied to the
feral A feral () animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some ...
donkeys that live west of the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico ...
; it may also refer to any small donkey.


History

The genus ''Equus'', which includes all extant equines, is believed to have evolved from ''
Dinohippus ''Dinohippus'' (Greek: "Terrible horse") is an extinct equid which was endemic to North America from the late Hemphillian stage of the Miocene through the Zanclean stage of the Pliocene (10.3—3.6 mya) and in existence for approximately . Foss ...
'', via the intermediate form ''
Plesippus ''Plesippus'' is a genus of extinct horse from the Pleistocene of North America. Although commonly seen as a subgenus of ''Equus'' recent cladistic analysis considers it a distinct genus. Species Two species are recognized by Barron et al. (201 ...
''. One of the oldest species is ''
Equus simplicidens The Hagerman horse (''Equus simplicidens''), also called the Hagerman zebra or the American zebra, was a North American species of equid from the Pliocene epoch and the Pleistocene epoch. It was one of the oldest horses of the genus ''Equus'' and ...
'', described as zebra-like with a donkey-shaped head. The oldest fossil to date is ~3.5 million years old from Idaho, USA. The genus appears to have spread quickly into the Old World, with the similarly aged ''Equus livenzovensis'' documented from western Europe and Russia. Molecular phylogenies indicate the most recent common ancestor of all modern equids (members of the genus ''Equus'') lived ~5.6 (3.9–7.8) mya. Direct paleogenomic sequencing of a 700,000-year-old middle Pleistocene horse metapodial bone from Canada implies a more recent 4.07 Myr before present date for the most recent common ancestor (MRCA) within the range of 4.0 to 4.5 Myr BP. The oldest divergencies are the Asian hemiones (subgenus ''E. (
Asinus ''Asinus'' is a subgenus of '' Equus'' (single-toed (hooved) grazing animal) that encompasses several subspecies of the Equidae commonly known as wild asses, characterized by long ears, a lean, straight-backed build, lack of a true withers, ...
)'', including the kulan,
onager The onager (; ''Equus hemionus'' ), A new species called the kiang (''E. kiang''), a Tibetan relative, was previously considered to be a subspecies of the onager as ''E. hemionus kiang'', but recent molecular studies indicate it to be a distinct ...
, and
kiang The kiang (''Equus kiang'') is the largest of the '' Asinus'' subgenus. It is native to the Tibetan Plateau, where it inhabits montane and alpine grasslands. Its current range is restricted to the plains of the Tibetan plateau; Ladakh; and nort ...
), followed by the African zebras (subgenera ''E. ( Dolichohippus)'', and ''E. (
Hippotigris Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: the Grévy's zebra (''Equus grevyi''), plains zebra (''E. quagga''), and the mountain zebra (''E. zeb ...
)''). All other modern forms including the domesticated horse (and many fossil
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
forms) belong to the subgenus ''E. ( Equus)'' which diverged ~4.8 (3.2–6.5) million years ago. The ancestors of the modern donkey are the
Nubia Nubia () (Nobiin: Nobīn, ) is a region along the Nile river encompassing the area between the first cataract of the Nile (just south of Aswan in southern Egypt) and the confluence of the Blue and White Niles (in Khartoum in central Sudan), or ...
n and
Somalia Somalia, , Osmanya script: 𐒈𐒝𐒑𐒛𐒐𐒘𐒕𐒖; ar, الصومال, aṣ-Ṣūmāl officially the Federal Republic of SomaliaThe ''Federal Republic of Somalia'' is the country's name per Article 1 of thProvisional Constituti ...
n subspecies of
African wild ass The African wild ass (''Equus africanus'') or African wild donkey is a wild member of the horse family, Equidae. This species is thought to be the ancestor of the domestic donkey (''Equus asinus''), which is sometimes placed within the same spec ...
. Remains of domestic donkeys dating to the fourth millennium BC have been found in
Ma'adi Maadi ( ar, المعادي / transliterated:   ) is a leafy suburban district south of Cairo, Egypt, on the east bank of the Nile about upriver from downtown Cairo. The Nile at Maadi is parallelled by the Corniche, a waterfront promenade a ...
in Lower Egypt, and it is believed that the domestication of the donkey was accomplished long after the domestication of cattle, sheep and goats in the seventh and eighth millennia BC. Donkeys were probably first domesticated by pastoral people in
Nubia Nubia () (Nobiin: Nobīn, ) is a region along the Nile river encompassing the area between the first cataract of the Nile (just south of Aswan in southern Egypt) and the confluence of the Blue and White Niles (in Khartoum in central Sudan), or ...
, and they supplanted the ox as the chief pack animal of that culture. The domestication of donkeys served to increase the mobility of pastoral cultures, having the advantage over ruminants of not needing time to chew their cud, and were vital in the development of long-distance trade across Egypt. In the
Dynasty IV The Fourth Dynasty of ancient Egypt (notated Dynasty IV) is characterized as a "golden age" of the Old Kingdom of Egypt. Dynasty IV lasted from to 2494 BC. It was a time of peace and prosperity as well as one during which trade with other ...
era of Egypt, between 2675 and 2565 BC, wealthy members of society were known to own over 1,000 donkeys, employed in agriculture, as dairy and meat animals and as pack animals. In 2003, the tomb of either King Narmer or King Hor-Aha (two of the first Egyptian pharaohs) was excavated and the skeletons of ten donkeys were found buried in a manner usually used with high ranking humans. These burials show the importance of donkeys to the early Egyptian state and its ruler. By the end of the fourth millennium BC, the donkey had spread to Southwest Asia, and the main breeding centre had shifted to
Mesopotamia Mesopotamia ''Mesopotamíā''; ar, بِلَاد ٱلرَّافِدَيْن or ; syc, ܐܪܡ ܢܗܪ̈ܝܢ, or , ) is a historical region of Western Asia situated within the Tigris–Euphrates river system, in the northern part of the F ...
by 1800 BC. The breeding of large, white riding asses made
Damascus )), is an adjective which means "spacious". , motto = , image_flag = Flag of Damascus.svg , image_seal = Emblem of Damascus.svg , seal_type = Seal , map_caption = , ...
famous, while Syrian breeders developed at least three other breeds, including one preferred by women for its easy
gait Gait is the pattern of movement of the limbs of animals, including humans, during locomotion over a solid substrate. Most animals use a variety of gaits, selecting gait based on speed, terrain, the need to maneuver, and energetic efficiency. Di ...
. The Muscat or Yemen ass was developed in
Arabia The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate. ...
. By the second millennium BC, the donkey was brought to Europe, possibly at the same time as
viticulture Viticulture (from the Latin word for ''vine'') or winegrowing (wine growing) is the cultivation and harvesting of grapes. It is a branch of the science of horticulture. While the native territory of ''Vitis vinifera'', the common grape vine, ran ...
was introduced, as the donkey is associated with the Syrian god of wine,
Dionysus In ancient Greek religion and myth, Dionysus (; grc, Διόνυσος ) is the god of the grape-harvest, winemaking, orchards and fruit, vegetation, fertility, insanity, ritual madness, religious ecstasy, festivity, and theatre. The Romans ...
. Greeks spread both of these to many of their colonies, including those in what are now Italy, France and Spain; Romans dispersed them throughout their empire. The first donkeys came to the Americas on ships of the Second Voyage of
Christopher Columbus Christopher Columbus * lij, Cristoffa C(or)ombo * es, link=no, Cristóbal Colón * pt, Cristóvão Colombo * ca, Cristòfor (or ) * la, Christophorus Columbus. (; born between 25 August and 31 October 1451, died 20 May 1506) was a ...
, and were landed at
Hispaniola Hispaniola (, also ; es, La Española; Latin and french: Hispaniola; ht, Ispayola; tnq, Ayiti or Quisqueya) is an island in the Caribbean that is part of the Greater Antilles. Hispaniola is the most populous island in the West Indies, and th ...
in 1495. The first to reach North America may have been two animals taken to Mexico by
Juan de Zumárraga Juan de Zumárraga, OFM (1468 – June 3, 1548) was a Spanish Basque Franciscan prelate and the first Bishop of Mexico. He was also the region's first inquisitor. He wrote ''Doctrina breve'', the first book published in the Western Hemispher ...
, the first bishop of Mexico, who arrived there on 6 December 1528, while the first donkeys to reach what is now the United States may have crossed the
Rio Grande The Rio Grande ( and ), known in Mexico as the Río Bravo del Norte or simply the Río Bravo, is one of the principal rivers (along with the Colorado River) in the southwestern United States and in northern Mexico. The length of the Rio G ...
with
Juan de Oñate Juan de Oñate y Salazar (; 1550–1626) was a Spanish conquistador from New Spain, explorer, and colonial governor of the province of Santa Fe de Nuevo México in the viceroyalty of New Spain. He led early Spanish expeditions to the Great ...
in April 1598. From that time on they spread northward, finding use in missions and mines. Donkeys were documented as present in what today is Arizona in 1679. By the
Gold Rush A gold rush or gold fever is a discovery of gold—sometimes accompanied by other precious metals and rare-earth minerals—that brings an onrush of miners seeking their fortune. Major gold rushes took place in the 19th century in Australia, New Z ...
years of the 19th century, the burro was the
beast of burden A working animal is an animal, usually domesticated, that is kept by humans and trained to perform tasks instead of being slaughtered to harvest animal products. Some are used for their physical strength (e.g. oxen and draft horses) or for ...
of choice of early prospectors in the western United States. With the end of the
placer mining Placer mining () is the mining of stream bed (Alluvium, alluvial) deposits for minerals. This may be done by open-pit mining, open-pit (also called open-cast mining) or by various surface excavating equipment or tunneling equipment. Placer minin ...
boom, many of them escaped or were abandoned, and a
feral A feral () animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some ...
population established itself.


Conservation status

About 41 million donkeys were reported worldwide in 2006. China had the most with 11 million, followed by Pakistan,
Ethiopia Ethiopia, , om, Itiyoophiyaa, so, Itoobiya, ti, ኢትዮጵያ, Ítiyop'iya, aa, Itiyoppiya officially the Federal Democratic Republic of Ethiopia, is a landlocked country in the Horn of Africa. It shares borders with Eritrea to the ...
and Mexico. As of 2017, however, the Chinese population was reported to have dropped to 3 million, with African populations under pressure as well, due to increasing trade and demand for donkey products in China. Some researchers believe the actual number may be somewhat higher since many donkeys go uncounted. The number of breeds and percentage of world population for each of the FAO's world regions was in 2006: In 1997 the number of donkeys in the world was reported to be continuing to grow, as it had steadily done throughout most of history; factors cited as contributing to this were increasing human population, progress in economic development and social stability in some poorer nations, conversion of forests to farm and range land, rising prices of motor vehicles and fuel, and the popularity of donkeys as pets. Since then, the world population of donkeys is reported to be rapidly shrinking, falling from 43.7 million to 43.5 million between 1995 and 2000, and to only 41 million in 2006. The fall in population is pronounced in developed countries; in Europe, the total number of donkeys fell from 3 million in 1944 to just over 1 million in 1994. The Domestic Animal Diversity Information System (
DAD-IS DAD-IS is the acronym of the worldwide Domestic Animal Diversity Information System of the Food and Agriculture Organization of the United Nations, within the FAO's management of animal genetic resources programme.Domestic Animal Diversity Infor ...
) of the
FAO The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...
listed 189 breeds of ass in June 2011. In 2000 the number of breeds of donkey recorded worldwide was 97, and in 1995 it was 77. The rapid increase is attributed to attention paid to identification and recognition of donkey breeds by the FAO's Animal Genetic Resources project. The rate of recognition of new breeds has been particularly high in some developed countries. In France only one breed, the Baudet du Poitou, was recognised until the early 1990s; by 2005, a further six donkey breeds had official recognition. In developed countries, the welfare of donkeys both at home and abroad has become a concern, and a number of sanctuaries for retired and rescued donkeys have been set up. The largest is
The Donkey Sanctuary The Donkey Sanctuary is a British charitable organisation devoted to the welfare of donkeys. The charity, which is based near Sidmouth in Devon, England, was founded in 1969. It is one of the largest equine charities in the world with an ann ...
near
Sidmouth Sidmouth () is a town on the English Channel in Devon, South West England, southeast of Exeter. With a population of 12,569 in 2011, it is a tourist resort and a gateway to the Jurassic Coast World Heritage Site. A large part of the town has ...
, England, which also supports donkey welfare projects in Egypt, Ethiopia, India,
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
, and Mexico. In 2017, a drop in the number of Chinese donkeys, combined with the fact that they are slow to reproduce, meant that Chinese suppliers began to look to Africa. As a result of the increase in demand, and the price that could be charged,
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
opened three donkey abattoirs. Concerns for donkeys' well-being, however, have resulted in a number of African countries (including
Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...
,
Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...
,
Botswana Botswana (, ), officially the Republic of Botswana ( tn, Lefatshe la Botswana, label=Setswana, ), is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory being the Kalahar ...
,
Niger ) , official_languages = , languages_type = National languagesBurkina Faso Burkina Faso (, ; , ff, 𞤄𞤵𞤪𞤳𞤭𞤲𞤢 𞤊𞤢𞤧𞤮, italic=no) is a landlocked country in West Africa with an area of , bordered by Mali to the northwest, Niger to the northeast, Benin to the southeast, Togo and Ghana to the ...
,
Mali Mali (; ), officially the Republic of Mali,, , ff, 𞤈𞤫𞤲𞥆𞤣𞤢𞥄𞤲𞤣𞤭 𞤃𞤢𞥄𞤤𞤭, Renndaandi Maali, italics=no, ar, جمهورية مالي, Jumhūriyyāt Mālī is a landlocked country in West Africa. Mali ...
, and
Senegal Senegal,; Wolof: ''Senegaal''; Pulaar: 𞤅𞤫𞤲𞤫𞤺𞤢𞥄𞤤𞤭 (Senegaali); Arabic: السنغال ''As-Sinighal'') officially the Republic of Senegal,; Wolof: ''Réewum Senegaal''; Pulaar : 𞤈𞤫𞤲𞤣𞤢𞥄𞤲𞤣𞤭 ...
) banning China from buying their donkey products. In 2019, The Donkey Sanctuary warned that the global donkey population could be reduced by half over the next half decade as the demand for
ejiao Donkey-hide gelatin or ass-hide glue ( la, colla corii asini) is gelatin obtained from the skin of the donkey (''Equus asinus'') by soaking and stewing. It is used as an ingredient in the traditional medicine of China, where it is called ejiao ...
increases in China.


Characteristics

Donkeys vary considerably in size, depending on both breed and environmental conditions, and heights at the
withers The withers is the ridge between the shoulder blades of an animal, typically a quadruped. In many species, it is the tallest point of the body. In horses and dogs, it is the standard place to measure the animal's height. In contrast, cattle ar ...
range from less than to approximately . Working donkeys in the poorest countries have a life expectancy of 12 to 15 years; in more prosperous countries, they may have a lifespan of 30 to 50 years. Donkeys are adapted to marginal
desert A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
lands. Unlike
wild Wild, wild, wilds or wild may refer to: Common meanings * Wild animal * Wilderness, a wild natural environment * Wildness, the quality of being wild or untamed Art, media and entertainment Film and television * ''Wild'' (2014 film), a 2014 A ...
and
feral horse A feral horse is a free-roaming horse of domesticated stock. As such, a feral horse is not a wild animal in the sense of an animal without domesticated ancestors. However, some populations of feral horses are managed as wildlife, and these ...
s, wild donkeys in dry areas are solitary and do not form
harems Harem (Persian: حرمسرا ''haramsarā'', ar, حَرِيمٌ ''ḥarīm'', "a sacred inviolable place; harem; female members of the family") refers to domestic spaces that are reserved for the women of the house in a Muslim family. A har ...
. Each adult donkey establishes a home range; breeding over a large area may be dominated by one jack. The loud call or bray of the donkey, which typically lasts for twenty seconds and can be heard for over three kilometres, may help keep in contact with other donkeys over the wide spaces of the desert. Donkeys have large ears, which may pick up more distant sounds, and may help cool the donkey's blood. Donkeys can defend themselves by biting, striking with the front hooves or kicking with the hind legs. Their vocalization, called a bray, is often represented in English as "hee haw".


Cross on back

Most donkeys have dorsal and shoulder stripes,
primitive markings Primitive markings are a group of hair coat markings and qualities seen in several equine species, including horses, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such breeds. The markings are ...
which form a distinctive cross pattern on their backs.


Breeding

A jenny is normally pregnant for about 12 months, though the gestation period varies from 11 to 14 months, and usually gives birth to a single foal. Births of twins are rare, though less so than in horses. About 1.7 percent of donkey pregnancies result in twins; both foals survive in about 14 percent of those. In general jennies have a conception rate that is lower than that of horses (i.e., less than the 60–65% rate for mares). Although jennies come into heat within 9 or 10 days of giving birth, their fertility remains low, and it is likely the reproductive tract has not returned to normal. Thus it is usual to wait one or two further oestrous cycles before rebreeding, unlike the practice with mares. Jennies are usually very protective of their foals, and some will not come into estrus while they have a foal at side. The time lapse involved in rebreeding, and the length of a jenny's gestation, means that a jenny will have fewer than one foal per year. Because of this and the longer gestation period, donkey breeders do not expect to obtain a foal every year, as horse breeders often do, but may plan for three foals in four years. Donkeys can interbreed with other members of the family Equidae, and are commonly interbred with horses. The
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
between a jack and a
mare A mare is an adult female horse or other equine. In most cases, a mare is a female horse over the age of three, and a filly is a female horse three and younger. In Thoroughbred horse racing, a mare is defined as a female horse more than four ...
is a
mule The mule is a domestic equine hybrid between a donkey and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two pos ...
, valued as a working and riding animal in many countries. Some large donkey breeds such as the
Asino di Martina Franca The Martina Franca donkey, it, Asino di Martina Franca, italic=no, is a breed of donkey from Puglia in southern Italy. It is the largest Italian donkey breed, and was famous for its qualities in the production of mules. It is particularly ass ...
, the
Baudet du Poitou The Baudet du Poitou, also called the Poitevin or Poitou donkey, is a French breed of donkey. It is one of the largest breeds, and jacks (donkey stallions) were bred to mares of the Poitevin horse breed to produce Poitevin mules, which were f ...
and the Mammoth Jack are raised only for mule production. The hybrid between a
stallion A stallion is a male horse that has not been gelded (castrated). Stallions follow the conformation and phenotype of their breed, but within that standard, the presence of hormones such as testosterone may give stallions a thicker, "cresty" nec ...
and a jenny is a
hinny A hinny is a domestic equine hybrid, the offspring of a male horse (a stallion) and a female donkey (a jenny). It is the reciprocal cross to the more common mule, which is the product of a male donkey (a jack) and a female horse (a mare). The ...
, and is less common. Like other inter-species hybrids, mules and hinnies are usually sterile. Donkeys can also breed with
zebra Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: the Grévy's zebra (''Equus grevyi''), plains zebra (''E. quagga''), and the mountain zebra (''E. zeb ...
s, in which case the offspring is called a zonkey (among other names).


Behaviour

Donkeys have a notorious reputation for stubbornness, but this has been attributed to a much stronger sense of
self-preservation Self-preservation is a behavior or set of behaviors that ensures the survival Survival, or the act of surviving, is the propensity of something to continue existing, particularly when this is done despite conditions that might kill or destroy ...
than exhibited by horses. Likely based on a stronger prey instinct and a weaker connection with humans, it is considerably more difficult to force or frighten a donkey into doing something it perceives to be dangerous for whatever reason. Once a person has earned their confidence they can be willing and companionable partners and very dependable in work. Although formal studies of their
behaviour Behavior (American English) or behaviour (British English) is the range of actions and mannerisms made by individuals, organisms, systems or artificial entities in some environment. These systems can include other systems or organisms as wel ...
and
cognition Cognition refers to "the mental action or process of acquiring knowledge and understanding through thought, experience, and the senses". It encompasses all aspects of intellectual functions and processes such as: perception, attention, thought, ...
are rather limited, donkeys appear to be quite intelligent, cautious, friendly, playful, and eager to learn.


Use

File:Skegness4web.jpg, Classic British seaside donkeys in
Skegness Skegness ( ) is a seaside town and civil parish in the East Lindsey District of Lincolnshire, England. On the Lincolnshire coast of the North Sea, the town is east of Lincoln and north-east of Boston. With a population of 19,579 as of 2011, i ...
File:104 Donkeys in Tayrona Park Colombia.JPG, Pack donkeys in Tayrona National Natural Park in northern
Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
File:Esel auf Ydra.jpg, Donkeys for transport on the island of Hydra File:Uyghur man on his donkey cart. Kashgar.jpg, Uyghur man on his donkey cart.
Kashgar Kashgar ( ug, قەشقەر, Qeshqer) or Kashi ( zh, c=喀什) is an oasis city in the Tarim Basin region of Southern Xinjiang. It is one of the westernmost cities of China, near the border with Afghanistan, Kyrgyzstan, Tajikistan and Pakistan ...
The donkey has been used as a working animal for at least years. Of the more than 40 million donkeys in the world, about 96% are in
underdeveloped Underdevelopment, in the context of international development, reflects a broad condition or phenomena defined and critiqued by theorists in fields such as economics, development studies, and postcolonial studies. Used primarily to distinguish st ...
countries, where they are used principally as pack animals or for draught work in transport or agriculture. After human labour, the donkey is the cheapest form of agricultural power. They may also be ridden, or used for threshing, raising water, milling and other work. Some cultures that prohibit women from working with
oxen An ox ( : oxen, ), also known as a bullock (in BrE British English (BrE, en-GB, or BE) is, according to Oxford Dictionaries, "English as used in Great Britain, as distinct from that used elsewhere". More narrowly, it can refer spec ...
in agriculture do not extend this taboo to donkeys. In developed countries where their use as beasts of burden has disappeared, donkeys are used to sire mules, to guard
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus ''Ovis'', in everyday usage it almost always refers to domesticated s ...
, for
donkey rides The domestic donkey is a hoofed mammal in the family Equidae, the same family as the horse. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a ...
for children or tourists, and as pets. Donkeys may be pastured or stabled with horses and ponies, and are thought to have a calming effect on nervous horses. If a donkey is introduced to a
mare A mare is an adult female horse or other equine. In most cases, a mare is a female horse over the age of three, and a filly is a female horse three and younger. In Thoroughbred horse racing, a mare is defined as a female horse more than four ...
and foal, the foal may turn to the donkey for support after it has been weaned from its mother. A few donkeys are milked or raised for meat. Approximately 3.5 million donkeys and mules are slaughtered each year for meat worldwide. In Italy, which has the highest consumption of equine meat in Europe and where donkey meat is the main ingredient of several regional dishes, about 1,000 donkeys were slaughtered in 2010, yielding approximately of meat.
Asses' milk Donkey milk (or ass milk, or jenny milk) is the milk from the domesticated donkey ''(Equus asinus)''. It has been used since antiquity for cosmetic purposes as well as infant nutrition. History Donkey milk has been used by humans for alimenta ...
may command good prices: the average price in Italy in 2009 was €15 per litre, and a price of €6 per 100 ml was reported from Croatia in 2008; it is used for soaps and cosmetics as well as dietary purposes. The niche markets for both milk and meat are expanding. In the past, donkey skin was used in the production of
parchment Parchment is a writing material made from specially prepared untanned skins of animals—primarily sheep, calves, and goats. It has been used as a writing medium for over two millennia. Vellum is a finer quality parchment made from the skins of ...
. In 2017, the UK based charity
The Donkey Sanctuary The Donkey Sanctuary is a British charitable organisation devoted to the welfare of donkeys. The charity, which is based near Sidmouth in Devon, England, was founded in 1969. It is one of the largest equine charities in the world with an ann ...
estimated that 1.8 million skins were traded every year, but the demand could be as high as 10 million. In China, donkey meat is considered a delicacy with some restaurants specializing in such dishes, and Guo Li Zhuang restaurants offer the genitals of donkeys in dishes.
Donkey-hide gelatin Donkey-hide gelatin or ass-hide glue ( la, colla corii asini) is gelatin obtained from the skin of the donkey (''Equus asinus'') by soaking and stewing. It is used as an ingredient in the traditional medicine of China, where it is called ejia ...
is produced by soaking and stewing the hide to make a traditional Chinese medicine product.
Ejiao Donkey-hide gelatin or ass-hide glue ( la, colla corii asini) is gelatin obtained from the skin of the donkey (''Equus asinus'') by soaking and stewing. It is used as an ingredient in the traditional medicine of China, where it is called ejiao ...
, the gelatine produced by boiling donkey skins, can sell for up to $388 per kilo, at October 2017 prices.


In warfare

During World War I
John Simpson Kirkpatrick John Kirkpatrick (enlisted as John Simpson; 6 July 1892 – 19 May 1915) was a stretcher bearer with the 3rd Australian Field Ambulance brigade during the Gallipoli campaign – the Allied attempt to capture Constantinople, ca ...
, a British stretcher bearer serving with the
Australian and New Zealand Army Corps The Australian and New Zealand Army Corps (ANZAC) was a First World War army corps of the Mediterranean Expeditionary Force. It was formed in Egypt in December 1914, and operated during the Gallipoli campaign. General William Birdwood comma ...
, and Richard Alexander "Dick" Henderson of the
New Zealand Medical Corps New is an adjective referring to something recently made, discovered, or created. New or NEW may refer to: Music * New, singer of K-pop group The Boyz Albums and EPs * ''New'' (album), by Paul McCartney, 2013 * ''New'' (EP), by Regurgitator ...
used donkeys to rescue wounded soldiers from the battlefield at
Gallipoli The Gallipoli peninsula (; tr, Gelibolu Yarımadası; grc, Χερσόνησος της Καλλίπολης, ) is located in the southern part of East Thrace, the European part of Turkey, with the Aegean Sea to the west and the Dardanelles ...
. According to British food writer
Matthew Fort Matthew Fort (born 29 January 1947) is a British food writer and critic. Matthew Fort is the son of the Conservative MP Richard Fort, who died when he was 12. His brother is the writer Tom Fort. He attended Eton College, and later Lancaster Un ...
, donkeys were used in the
Italian Army "The safeguard of the republic shall be the supreme law" , colors = , colors_labels = , march = ''Parata d'Eroi'' ("Heroes's parade") by Francesco Pellegrino, ''4 Maggio'' (May 4) ...
. The Mountain Fusiliers each had a donkey to
carry Carry or carrying may refer to: People *Carry (name) Finance * Carried interest (or carry), the share of profits in an investment fund paid to the fund manager * Carry (investment), a financial term: the carry of an asset is the gain or cost of h ...
their gear, and in extreme circumstances the animal could be eaten. Donkeys have also been used to carry explosives in conflicts that include the
war in Afghanistan War in Afghanistan, Afghan war, or Afghan civil war may refer to: *Conquest of Afghanistan by Alexander the Great (330 BC – 327 BC) *Muslim conquests of Afghanistan (637–709) *Conquest of Afghanistan by the Mongol Empire (13th century), see als ...
and others.


Care


Shoeing

Donkey hooves are more elastic than those of horses, and do not naturally wear down as fast. Regular clipping may be required; neglect can lead to permanent damage. Working donkeys may need to be shod. Donkey shoes are similar to
horseshoe A horseshoe is a fabricated product designed to protect a horse hoof from wear. Shoes are attached on the palmar surface (ground side) of the hooves, usually nailed through the insensitive hoof wall that is anatomically akin to the human toen ...
s, but usually smaller and without toe-clips.


Nutrition

In their native arid and semi-arid climates, donkeys spend more than half of each day foraging and feeding, often on poor quality scrub. The donkey has a tough
digestive system The human digestive system consists of the gastrointestinal tract plus the accessory organs of digestion (the tongue, salivary glands, pancreas, liver, and gallbladder). Digestion involves the breakdown of food into smaller and smaller compone ...
in which roughage is efficiently broken down by hind gut fermentation, microbial action in the
caecum The cecum or caecum is a pouch within the peritoneum that is considered to be the beginning of the large intestine. It is typically located on the right side of the body (the same side of the body as the appendix, to which it is joined). The wo ...
and
large intestine The large intestine, also known as the large bowel, is the last part of the gastrointestinal tract and of the digestive system in tetrapods. Water is absorbed here and the remaining waste material is stored in the rectum as feces before being r ...
. While there is no marked structural difference between the gastro-intestinal tract of a donkey and that of a horse, the digestion of the donkey is more efficient. It needs less food than a horse or pony of comparable height and weight, approximately 1.5 percent of body weight per day in dry matter, compared to the 2–2.5 percent consumption rate possible for a horse.Hall, Marvin H. and Patricia M. Comerford. "Pasture and Hay for Horses – Argonomy facts 32," 1992 University of Pennsylvania, Cooperative Extension Service.
Web site accessed February 14, 2007.
Donkeys are also less prone to
colic Colic or cholic () is a form of pain that starts and stops abruptly. It occurs due to muscular contractions of a hollow tube ( small and large intestine, gall bladder, ureter, etc.) in an attempt to relieve an obstruction by forcing content out ...
. The reasons for this difference are not fully understood; the donkey may have different
intestinal flora Gut microbiota, gut microbiome, or gut flora, are the microorganisms, including bacteria, archaea, fungi, and viruses that live in the digestive tracts of animals. The gastrointestinal metagenome is the aggregate of all the genomes of the gut mi ...
to the horse, or a longer gut retention time. Donkeys obtain most of their energy from structural carbohydrates. Some suggest that a donkey needs to be fed only
straw Straw is an agricultural byproduct consisting of the dry stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the yield of cereal crops such as barley, oats, rice, rye and wheat. It has a number ...
(preferably barley straw), supplemented with controlled grazing in the summer or
hay Hay is grass, legumes, or other herbaceous plants that have been cut and dried to be stored for use as animal fodder, either for large grazing animals raised as livestock, such as cattle, horses, goats, and sheep, or for smaller domesticat ...
in the winter, to get all the energy, protein, fat and vitamins it requires; others recommend some grain to be fed, particularly to working animals, and others advise against feeding straw. They do best when allowed to consume small amounts of food over long periods. They can meet their nutritional needs on 6 to 7 hours of grazing per day on average dryland pasture that is not stressed by drought. If they are worked long hours or do not have access to pasture, they require hay or a similar dried forage, with no more than a 1:4 ratio of legumes to grass. They also require salt and mineral supplements, and access to clean, fresh water. In temperate climates the forage available is often too abundant and too rich; over-feeding may cause weight gain and obesity, and lead to metabolic disorders such as founder (
laminitis Laminitis is a disease that affects the feet of ungulates and is found mostly in horses and cattle. Clinical signs include foot tenderness progressing to inability to walk, increased digital pulses, and increased temperature in the hooves. Sever ...
) and
hyperlipaemia Hyperlipidemia is abnormally elevated levels of any or all lipids (fats, cholesterol, or triglycerides) or lipoproteins in the blood. citing: and The term ''hyperlipidemia'' refers to the laboratory finding itself and is also used as an umbre ...
, or to
gastric ulcers Peptic ulcer disease (PUD) is a break in the inner lining of the stomach, the first part of the small intestine, or sometimes the lower esophagus. An ulcer in the stomach is called a gastric ulcer, while one in the first part of the intestines i ...
. Throughout the world, working donkeys are associated with the very poor, with those living at or below subsistence level. Few receive adequate food, and in general donkeys throughout the
Third World The term "Third World" arose during the Cold War to define countries that remained non-aligned with either NATO or the Warsaw Pact. The United States, Canada, Japan, South Korea, Western European nations and their allies represented the " First ...
are under-nourished and over-worked.


Feral populations

In some areas domestic donkeys have returned to the wild and established
feral A feral () animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some ...
populations such as those of the burro of North America and the
Asinara donkey The Asinara, it, Asino dell'Asinara, italic=no (), is a breed of feral donkey indigenous to the island of Asinara, which lies off the north-west coast of Sardinia, Italy, in the province of Sassari. Most of the population is wholly or part ...
of
Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label=Algherese and Catalan) is the second-largest island in the Mediterranean Sea, after ...
, Italy, both of which have protected status. Feral donkeys can also cause problems, notably in environments that have evolved free of any form of
equid Equidae (sometimes known as the horse family) is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. All extant species are in the genus '' Equus'', w ...
, such as Hawaii. In Australia, where there may be 5 million feral donkeys, they are regarded as an invasive pest and have a serious impact on the environment. They may compete with livestock and native animals for resources, spread weeds and diseases, foul or damage watering holes and cause erosion.


Donkey hybrids

The earliest documented donkey hybrid was the kunga, which was used as a draft animal in the Syrian and Mesopotamian kingdoms of the second half of the 3rd millennium BCE. A cross between a captive male
Syrian wild ass The Syrian wild ass (''Equus hemionus hemippus''), less commonly known as a hemippe, an achdari, or a Mesopotamian or Syrian onager, is an extinct subspecies of onager native to the Arabian peninsula and surrounding areas. It ranged across present ...
and a female domesticated donkey (jenny), they represent the earliest known example of human-directed animal hybridization. They were produced at a breeding center at Nagar (modern
Tell Brak Tell Brak (Nagar, Nawar) was an ancient city in Syria; its remains constitute a tell located in the Upper Khabur region, near the modern village of Tell Brak, 50 kilometers north-east of Al-Hasaka city, Al-Hasakah Governorate. The city' ...
) and were sold or given as gifts throughout the region, where they became significant status symbols, pulling battle wagons and the chariots of kings, and also being sacrificed to bury with high-status people. They fell out of favor following the introduction of the domestic horse and its donkey hybrid, the
mule The mule is a domestic equine hybrid between a donkey and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two pos ...
, into the region at the end of the 3rd millennium BCE. A male donkey (jack) crossed with a female horse produces a
mule The mule is a domestic equine hybrid between a donkey and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two pos ...
, while a male horse crossed with a jenny produces a
hinny A hinny is a domestic equine hybrid, the offspring of a male horse (a stallion) and a female donkey (a jenny). It is the reciprocal cross to the more common mule, which is the product of a male donkey (a jack) and a female horse (a mare). The ...
. Horse-donkey hybrids are almost always sterile because of a failure of their developing gametes to complete meiosis. The lower progesterone production of the jenny may also lead to early embryonic loss. In addition, there are reasons not directly related to reproductive biology. Due to different
mating In biology, mating is the pairing of either opposite-sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. ''Copulation'' is the union of the sex organs of two sexually reproduc ...
behavior, jacks are often more willing to cover mares than stallions are to breed jennies. Further, mares are usually larger than jennies and thus have more room for the ensuing foal to grow in the womb, resulting in a larger animal at birth. It is commonly believed that mules are more easily handled and also physically stronger than hinnies, making them more desirable for breeders to produce. The offspring of a
zebra Zebras (, ) (subgenus ''Hippotigris'') are African equines with distinctive black-and-white striped coats. There are three living species: the Grévy's zebra (''Equus grevyi''), plains zebra (''E. quagga''), and the mountain zebra (''E. zeb ...
-donkey cross is called a zonkey,
zebroid A zebroid is the offspring of any cross between a zebra and any other equine to create a hybrid. In most cases, the sire is a zebra stallion. Offspring of a donkey sire and zebra dam are called a donkra and offspring of a horse sire and a z ...
, zebrass, or zedonk; ''zebra mule'' is an older term, but still used in some regions today. The foregoing terms generally refer to hybrids produced by breeding a male zebra to a female donkey. ''Zebra hinny, zebret'' and ''zebrinny'' all refer to the cross of a female zebra with a male donkey. Zebrinnies are rarer than zedonkies because female zebras in captivity are most valuable when used to produce full-blooded zebras. There are not enough female zebras breeding in captivity to spare them for hybridizing; there is no such limitation on the number of female donkeys breeding.


See also

*
Animal-borne bomb attacks Animal-borne bomb attacks are the use of animals as delivery systems for explosives. The explosives are strapped to a pack animal such as a horse, mule or donkey. The pack animal may be set off in a crowd. Projects of bat bombs, dog bombs, an ...
* Cultural references to donkeys *
Jennet A jennet or Spanish jennet was a small Spanish horse. It was noted for a smooth naturally ambling gait, compact and well-muscled build, and a good disposition. The jennet was an ideal light riding horse, and as such spread across Europe and prov ...
, a type of medieval horse


References

{{Authority control Feral animals Mammals as pets Mammals described in 1758 Taxa named by Carl Linnaeus African wild ass Pack animals Subspecies