Common Raven
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The common raven (''Corvus corax'') is a large all-black
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
bird. It is the most widely distributed of all
corvids Corvidae is a cosmopolitan family of oscine passerine birds that contains the crows, ravens, rooks, jackdaws, jays, magpies, treepies, choughs, and nutcrackers. In colloquial English, they are known as the crow family or corvids. Currently, 13 ...
, found across the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the Equator. For other planets in the Solar System, north is defined as being in the same celestial hemisphere relative to the invariable plane of the solar system as Earth's Nort ...
. It is a raven known by many names at the subspecies level; there are at least eight
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
with little variation in appearance, although recent research has demonstrated significant genetic differences among populations from various regions. It is one of the two largest corvids, alongside the
thick-billed raven The thick-billed raven (''Corvus crassirostris''), a corvid from the Horn of Africa, shares with the common raven the distinction of being the largest bird in the corvid family, and indeed the largest of the passerines. The thick-billed raven a ...
, and is possibly the heaviest passerine bird; at maturity, the common raven averages in length and in mass. Although their typical lifespan is considerably shorter, common ravens can live more than 23 years in the wild. Young birds may travel in
flocks Flocking is the behaviour exhibited when a group of birds, called a flock, are foraging or in flight. Computer simulations and mathematical models that have been developed to emulate the flocking behaviours of birds can also generally be applie ...
but later
mate for life In biology, a pair bond is the strong affinity that develops in some species between a mating pair, often leading to the production and rearing of offspring and potentially a lifelong bond. Pair-bonding is a term coined in the 1940s that is freque ...
, with each
mated pair Breeding pair is a pair of animals which cooperate over time to produce offspring with some form of a bond between the individuals.Gaston, A. J.The evolution of group territorial behavior and cooperative breeding" The American Naturalist 112.988 ...
defending a
territory A territory is an area of land, sea, or space, particularly belonging or connected to a country, person, or animal. In international politics, a territory is usually either the total area from which a state may extract power resources or a ...
. Common ravens have coexisted with humans for thousands of years and in some areas have been so numerous that people have regarded them as
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
. Part of their success as a species is due to their omnivorous diet; they are extremely versatile and
opportunistic Opportunism is the practice of taking advantage of circumstances – with little regard for principles or with what the consequences are for others. Opportunist actions are expedient actions guided primarily by self-interested motives. The term ...
in finding sources of nutrition, feeding on
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
, insects, cereal grains, berries, fruit, small animals, nesting birds, and food waste. Some notable feats of problem-solving provide evidence that the common raven is unusually intelligent. Over the centuries, the raven has been the subject of mythology, folklore, art, and literature. In many cultures, including the indigenous cultures of
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion#Europe, subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, ...
, ancient Ireland and Wales,
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainous ...
, the
northwest coast of North America The Pacific Northwest (sometimes Cascadia, or simply abbreviated as PNW) is a geographic region in western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though ...
, and
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
and northeast Asia, the common raven has been revered as a spiritual figure or godlike creature.


Taxonomy

The common raven was one of the many species originally described, with its type locality given as Europe, by
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
in his landmark 1758 10th edition of ''Systema Naturae'', and it still bears its original name of ''Corvus corax''. It is the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
of the genus ''
Corvus ''Corvus'' is a widely distributed genus of medium-sized to large birds in the family Corvidae. It includes species commonly known as crows, ravens and Rook (bird), rooks. The species commonly encountered in Europe are the carrion crow, the hoo ...
'', derived from the
Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...
word for 'raven'. The specific epithet ''corax'' is the Latinized form of the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
word , meaning 'raven' or 'crow'.. The modern English word ''raven'' has cognates in many other
Germanic languages The Germanic languages are a branch of the Indo-European language family spoken natively by a population of about 515 million people mainly in Europe, North America, Oceania and Southern Africa. The most widely spoken Germanic language, Engli ...
, including
Old Norse Old Norse, Old Nordic, or Old Scandinavian, is a stage of development of North Germanic languages, North Germanic dialects before their final divergence into separate Nordic languages. Old Norse was spoken by inhabitants of Scandinavia and t ...
(and subsequently
modern Icelandic Icelandic (; is, íslenska, link=no ) is a North Germanic language spoken by about 314,000 people, the vast majority of whom live in Iceland, where it is the national language. Due to being a West Scandinavian language, it is most closely re ...
) and
Old High German Old High German (OHG; german: Althochdeutsch (Ahd.)) is the earliest stage of the German language, conventionally covering the period from around 750 to 1050. There is no standardised or supra-regional form of German at this period, and Old High ...
, all which descend from
Proto-Germanic Proto-Germanic (abbreviated PGmc; also called Common Germanic) is the reconstructed proto-language of the Germanic branch of the Indo-European languages. Proto-Germanic eventually developed from pre-Proto-Germanic into three Germanic branc ...
. An old Scottish word or , akin to the French , has been used for both this bird and the
carrion crow The carrion crow (''Corvus corone'') is a passerine bird of the family Corvidae and the genus ''Corvus'' which is native to western Europe and the eastern Palearctic. Taxonomy and systematics The carrion crow was one of the many species origi ...
.
Collective noun In linguistics, a collective noun is a word referring to a collection of things taken as a whole. Most collective nouns in everyday speech are not specific to one kind of thing. For example, the collective noun "group" can be applied to people (" ...
s for a group of ravens (or at least the common raven) include "unkindness" and "conspiracy".


Classification

The closest relatives of the common raven are the
brown-necked raven The brown-necked raven (''Corvus ruficollis'') is a larger bird (52–56 cm in length) than the carrion crow though not as large as the common raven. It has similar proportions to the common raven but the bill is not so large or deep and th ...
(''C. ruficollis''), the
pied crow The pied crow (''Corvus albus'') is a widely distributed African bird species in the crow genus. Structurally, the pied crow is better thought of as a small crow-sized raven, especially as it can hybridise with the Somali crow (dwarf raven) wher ...
(''C. albus'') of Africa, and the
Chihuahuan raven The Chihuahuan raven (''Corvus cryptoleucus'') is a species of crow in the family Corvidae that is native to the United States and Mexico. Description The proportions resemble the common raven with a heavy bill, but is about the same size as a c ...
(''C. cryptoleucus'') of the North American Southwest. While some authorities have recognized as many as 11
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
,Marzluff, J. M. (2009). "Common Raven (''Corvus corax'')". pp. 638–639 in ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...
.'' Bush-shrikes to Old World Sparrows. del Hoyo, J., Elliott, A. and Christie, D. A. (eds.). Lynx Edicions, Barcelona.
others recognize only eight:Clements, J. F. (2007). ''The Clements Checklist of the Birds of the World.'' 6th edition. Christopher Helm.


Evolutionary history

The common raven evolved in the
Old World The "Old World" is a term for Afro-Eurasia that originated in Europe , after Europeans became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia, which were previously thought of by the ...
and crossed the
Bering land bridge Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip of ...
into North America. Recent genetic studies, which examined the DNA of common ravens from across the world, have determined that the birds fall into at least two
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
s: a
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
clade, found only in the southwestern United States, and a
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
clade, found across the rest of the Northern Hemisphere. Birds from both clades look alike, but the groups are genetically distinct and began to diverge about two million years ago. The findings indicate that based on
mitochondrial DNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial D ...
, common ravens from the rest of the United States are more closely related to those in Europe and Asia than to those in the California clade, and that common ravens in the California clade are more closely related to the
Chihuahuan raven The Chihuahuan raven (''Corvus cryptoleucus'') is a species of crow in the family Corvidae that is native to the United States and Mexico. Description The proportions resemble the common raven with a heavy bill, but is about the same size as a c ...
(''C. cryptoleucus'') than to those in the Holarctic clade. Ravens in the Holarctic clade are more closely related to the
pied crow The pied crow (''Corvus albus'') is a widely distributed African bird species in the crow genus. Structurally, the pied crow is better thought of as a small crow-sized raven, especially as it can hybridise with the Somali crow (dwarf raven) wher ...
(''C. albus'') than they are to the California clade. Thus, the common raven species as traditionally delimited is considered to be
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
. One explanation for these genetic findings is that common ravens settled in California at least two million years ago and became separated from their relatives in Europe and Asia during a
glacial period A glacial period (alternatively glacial or glaciation) is an interval of time (thousands of years) within an ice age that is marked by colder temperatures and glacier advances. Interglacials, on the other hand, are periods of warmer climate betw ...
. One million years ago, a group from the California clade evolved into a new species, the Chihuahuan raven. Other members of the Holarctic clade arrived later in a separate migration from Asia, perhaps at the same time as humans. A 2011 study suggested that there are no restrictions on
gene flow In population genetics, gene flow (also known as gene migration or geneflow and allele flow) is the transfer of genetic material from one population to another. If the rate of gene flow is high enough, then two populations will have equivalent a ...
between the Californian and Holarctic common raven groups, and that the lineages can remerge, effectively reversing a potential speciation. A recent study of raven mitochondrial DNA showed that the isolated population from the
Canary Islands The Canary Islands (; es, Canarias, ), also known informally as the Canaries, are a Spanish autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to the African mainland, they are west of Morocc ...
is distinct from other populations. The study did not include any individuals from the North African population, and its position is therefore unclear, though its
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines * Morphology (archaeology), study of the shapes or forms of artifacts * Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies ...
is very close to the population of the Canaries (to the extent that the two are often considered part of a single subspecies).


Description

A mature common raven ranges between 54 and 67 cm (21" and 26") long, with a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan of ...
of 115 to 150 cm (45–51"). Recorded weights range from 0.69 to 2 kg (1.5 to 4.4 lb), thus making the common raven one of the heaviest
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by t ...
s. Birds from colder regions such as the
Himalayas The Himalayas, or Himalaya (; ; ), is a mountain range in Asia, separating the plains of the Indian subcontinent from the Tibetan Plateau. The range has some of the planet's highest peaks, including the very highest, Mount Everest. Over 100 ...
and Greenland are generally larger with slightly larger bills, while those from warmer regions are smaller with proportionally smaller bills. Representative of the size variation in the species, ravens from
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
weighed an average of , those from
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
weighed an average of and those from
Nova Scotia Nova Scotia ( ; ; ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. Nova Scotia is Latin for "New Scotland". Most of the population are native Eng ...
weighed an average of . The
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Plac ...
is large and slightly curved, with a culmen length of , easily one of the largest bills amongst passerines (perhaps only the
thick-billed raven The thick-billed raven (''Corvus crassirostris''), a corvid from the Horn of Africa, shares with the common raven the distinction of being the largest bird in the corvid family, and indeed the largest of the passerines. The thick-billed raven a ...
has a noticeably larger bill). It has a longish, strongly graduated tail, at , and mostly black
iridescent Iridescence (also known as goniochromism) is the phenomenon of certain surfaces that appear to gradually change color as the angle of view or the angle of illumination changes. Examples of iridescence include soap bubbles, feathers, butterfl ...
plumage, and a dark brown
iris Iris most often refers to: *Iris (anatomy), part of the eye *Iris (mythology), a Greek goddess * ''Iris'' (plant), a genus of flowering plants * Iris (color), an ambiguous color term Iris or IRIS may also refer to: Arts and media Fictional ent ...
. The throat feathers are elongated and pointed and the bases of the neck feathers are pale brownish-grey. The legs and feet are good-sized, with a tarsus length of . Juvenile plumage is similar but duller with a blue-grey iris. Apart from its greater size, the common raven differs from its cousins, the
crow A crow is a bird of the genus '' Corvus'', or more broadly a synonym for all of ''Corvus''. Crows are generally black in colour. The word "crow" is used as part of the common name of many species. The related term "raven" is not pinned scientifica ...
s, by having a larger and heavier black beak, shaggy feathers around the throat and above the beak, and a wedge-shaped tail. Flying ravens are distinguished from crows by their tail shape, larger wing area, and more stable soaring style, which generally involves less wing flapping. Despite their bulk, ravens are easily as agile in flight as their smaller cousins. In flight the feathers produce a creaking sound that has been likened to the rustle of silk. The voice of ravens is also quite distinct, its usual call being a deep croak of a much more sonorous quality than a crow's call. In North America, the
Chihuahuan raven The Chihuahuan raven (''Corvus cryptoleucus'') is a species of crow in the family Corvidae that is native to the United States and Mexico. Description The proportions resemble the common raven with a heavy bill, but is about the same size as a c ...
(''C. cryptoleucus'') is fairly similar to the relatively small common ravens of the American southwest and is best distinguished by the still relatively smaller size of its bill, beard and body and relatively longer tail. All-black
carrion crow The carrion crow (''Corvus corone'') is a passerine bird of the family Corvidae and the genus ''Corvus'' which is native to western Europe and the eastern Palearctic. Taxonomy and systematics The carrion crow was one of the many species origi ...
(''C. corone'') in Europe may suggest a raven due to their largish bill but are still distinctly smaller and have the wing and tail shapes typical of crows. In the
Faroe Islands The Faroe Islands ( ), or simply the Faroes ( fo, Føroyar ; da, Færøerne ), are a North Atlantic island group and an autonomous territory of the Kingdom of Denmark. They are located north-northwest of Scotland, and about halfway bet ...
, a now-extinct white-and-black colour morph of this species existed, known as the
pied raven The pied raven (''Corvus corax varius'' morpha ''leucophaeus'') is an extinct colour morph of the North Atlantic subspecies of the common raven which was only found on the Faroe Islands and was last seen in 1902. It had large areas of white feath ...
. White ravens are occasionally found in the wild. Birds in
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, ...
lack the pink eyes of an albino, and are instead
leucistic Leucism () is a wide variety of conditions that result in the partial loss of pigmentation in an animal—causing white, pale, or patchy coloration of the skin, hair, feathers, scales, or cuticles, but not the eyes. It is occasionally spelled ' ...
, a condition where an animal lacks any of several different types of pigment, not simply
melanin Melanin (; from el, μέλας, melas, black, dark) is a broad term for a group of natural pigments found in most organisms. Eumelanin is produced through a multistage chemical process known as melanogenesis, where the oxidation of the amino ...
. Common ravens have a wide range of vocalizations which are of interest to
ornithologist Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
s. Gwinner carried out important studies in the early 1960s, recording and photographing his findings in great detail. Fifteen to 30 categories of vocalization have been recorded for this species, most of which are used for social interaction. Calls recorded include alarm calls, chase calls, and flight calls. The species has a distinctive, deep, resonant ''prruk-prruk-prruk'' call, which to experienced listeners is unlike that of any other corvid. Its very wide and complex vocabulary includes a high, knocking ''toc-toc-toc'', a dry, grating ''kraa'', a low guttural rattle and some calls of an almost musical nature. Like other corvids, the common raven can mimic sounds from their environment, including human speech. Non-vocal sounds produced by the common raven include wing whistles and bill snapping. Clapping or clicking has been observed more often in females than in males. If a member of a pair is lost, its mate reproduces the calls of its lost partner to encourage its return.


Distribution and habitat

The common raven can thrive in varied climates; indeed this species has the largest range of any member of the genus, and one of the largest of any passerine. They range throughout the
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
from Arctic and
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
habitats in North America and Eurasia to the deserts of North Africa, and to islands in the Pacific Ocean. In the British Isles, they are more common in Scotland, Wales, northern England and the west of Ireland. In
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
, they have been recorded at altitudes up to 5,000 m (16,400 ft), and as high as 6,350 m (20,600 ft) on
Mount Everest Mount Everest (; Tibetan: ''Chomolungma'' ; ) is Earth's highest mountain above sea level, located in the Mahalangur Himal sub-range of the Himalayas. The China–Nepal border runs across its summit point. Its elevation (snow heig ...
. The population sometimes known as the Punjab raven—described as ''Corvus corax laurencei'' (also spelt ''lawrencii'' or ''laurencii'') by
Allan Octavian Hume Allan Octavian Hume, CB ICS (4 June 1829 – 31 July 1912) was a British civil servant, political reformer, ornithologist and botanist who worked in British India. He was the founder of the Indian National Congress. A notable ornithologist, Hum ...
but more often considered synonymous with ''subcorax''—is restricted to the
Sindh Sindh (; ; ur, , ; historically romanized as Sind) is one of the four provinces of Pakistan. Located in the southeastern region of the country, Sindh is the third-largest province of Pakistan by land area and the second-largest province ...
district of Pakistan and adjoining regions of northwestern India. They are generally resident within their range for the whole year. In his 1950 work, ''Grønlands Fugle'' 'Birds of Greenland'' noted ornithologist
Finn Salomonsen Finn Salomonsen (31 January 1909 – 23 April 1983) was a Danish ornithologist. He is best known for his work on the birds of Greenland. His interest in Greenland began at the age of 16 when he made a trip with Lehn Schioler to the Upernavik Dis ...
indicated that common ravens did not overwinter in the Arctic. However, in Arctic Canada and Alaska, they are found year-round. Young birds may disperse locally. Goodwin, p. 139 In the United Kingdom, the common raven's range has been increasing, though it favours mountainous or coastal terrain, but can also be found in parks with tall trees suitable for use as habitation. Its population is at its most dense in the north and west of the country, though the species is expanding its population southwards. Most common ravens prefer wooded areas with large expanses of open land nearby, or coastal regions for their nesting sites and feeding grounds. In some areas of dense human population, such as
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
in the United States, they take advantage of a plentiful food supply and have seen a surge in their numbers. On coasts, individuals of this species are often evenly distributed and prefer to build their nest sites along sea cliffs. Common ravens are often located in coastal regions because these areas provide easy access to water and a variety of food sources. Also, coastal regions have stable weather patterns without extreme cold or hot temperatures. In general, common ravens live in a wide array of environments but prefer heavily contoured landscapes. When the environment changes in vast degrees, these birds will respond with a stress response. The hormone known as
corticosterone Corticosterone, also known as 17-deoxycortisol and 11β,21-dihydroxyprogesterone, is a 21-carbon steroid hormone of the corticosteroid type produced in the cortex of the adrenal glands. It is of minor importance in humans, except in the very rare ...
is activated by the
hypothalamic–pituitary–adrenal axis The hypothalamic–pituitary–adrenal axis (HPA axis or HTPA axis) is a complex set of direct influences and feedback interactions among three components: the hypothalamus (a part of the brain located below the thalamus), the pituitary gland (a ...
. Corticosterone is activated when the bird is exposed to stress, such as migrating great distances.


Behaviour

Common ravens usually travel in mated pairs, although young birds may form
flocks Flocking is the behaviour exhibited when a group of birds, called a flock, are foraging or in flight. Computer simulations and mathematical models that have been developed to emulate the flocking behaviours of birds can also generally be applie ...
. Relationships between common ravens are often quarrelsome, yet they demonstrate considerable devotion to their families.


Predation

Owing to its size, gregariousness and its defensive abilities, the common raven has few natural predators. Predators of its eggs include
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s,
marten A marten is a weasel-like mammal in the genus ''Martes'' within the subfamily Guloninae, in the family Mustelidae. They have bushy tails and large paws with partially retractile claws. The fur varies from yellowish to dark brown, depending on t ...
s, and sometimes
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, just ...
s. Ravens are quite vigorous at defending their young and are usually successful at driving off perceived threats. They attack potential predators by flying at them and lunging with their large bills. Humans are occasionally attacked if they get close to a raven nest, though serious injuries are unlikely. There are a few records of predation by large birds of prey. Their attackers in America have reportedly included
great horned owl The great horned owl (''Bubo virginianus''), also known as the tiger owl (originally derived from early naturalists' description as the "winged tiger" or "tiger of the air"), or the hoot owl, is a large owl native to the Americas. It is an extrem ...
s,
northern goshawk The northern goshawk (; ''Accipiter gentilis'') is a species of medium-large bird of prey, raptor in the Family (biology), family Accipitridae, a family which also includes other extant diurnal raptors, such as eagles, buzzards and harrier (bird) ...
s,
bald eagle The bald eagle (''Haliaeetus leucocephalus'') is a bird of prey found in North America. A sea eagle, it has two known subspecies and forms a species pair with the white-tailed eagle (''Haliaeetus albicilla''), which occupies the same niche as ...
s,
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of p ...
s and
red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members with ...
s. It is possible that the two hawk species only attack young ravens; in one instance a
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a Cosmopolitan distribution, cosmopolitan bird of prey (Bird of prey, raptor) in the family (biology), family Falco ...
swooped at a newly fledged raven but was chased off by the parent ravens. In
Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...
, their reported predators include, in addition to golden eagles,
Eurasian eagle-owl The Eurasian eagle-owl (''Bubo bubo'') is a species of eagle-owl that resides in much of Palearctic, Eurasia. It is also called the Uhu and it is occasionally abbreviated to just the eagle-owl in Europe. It is one of the largest species of owl, ...
s,
white-tailed eagle The white-tailed eagle (''Haliaeetus albicilla'') is a very large species of sea eagle widely distributed across temperate Eurasia. Like all eagles, it is a member of the family Accipitridae (or accipitrids) which includes other diurnal raptors ...
s, Steller's sea-eagles,
eastern imperial eagle The eastern imperial eagle (''Aquila heliaca'') is a large bird of prey that breeds in southeastern Europe and extensively through West and Central Asia. Most populations are migratory and winter in northeastern Africa, the Middle East and South ...
s and
gyrfalcon The gyrfalcon ( or ) (), the largest of the falcon species, is a bird of prey. The abbreviation gyr is also used. It breeds on Arctic coasts and tundra, and the islands of northern North America and the Eurosiberian region. It is mainly a reside ...
s. Because they are potentially hazardous prey for raptorial birds, raptors must usually take them by surprise and most attacks are on fledgling ravens. More rarely still, large mammalian predators such as
lynx A lynx is a type of wild cat. Lynx may also refer to: Astronomy * Lynx (constellation) * Lynx (Chinese astronomy) * Lynx X-ray Observatory, a NASA-funded mission concept for a next-generation X-ray space observatory Places Canada * Lynx, Ontar ...
es,
coyote The coyote (''Canis latrans'') is a species of canis, canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecologica ...
s and
cougar The cougar (''Puma concolor'') is a large Felidae, cat native to the Americas. Its Species distribution, range spans from the Canadian Yukon to the southern Andes in South America and is the most widespread of any large wild terrestrial mamm ...
s have also attacked ravens. This principally occurs at a nest site and when other prey for the carnivores are scarce. Ravens are highly wary around novel carrion sites and, in North America, have been recorded waiting for the presence of
American crow The American crow (''Corvus brachyrhynchos'') is a large passerine bird species of the family Corvidae. It is a common bird found throughout much of North America. American crows are the New World counterpart to the carrion crow of Europe an ...
s and
blue jay The blue jay (''Cyanocitta cristata'') is a passerine bird in the family Corvidae, native to eastern North America. It lives in most of the eastern and central United States; some eastern populations may be migratory. Resident populations are a ...
s before approaching to eat.


Breeding

Juveniles begin to court at a very early age, but may not bond for another two or three years. Aerial acrobatics, demonstrations of intelligence, and ability to provide food are key behaviours of courting. Once paired, they tend to nest together for life, usually in the same location. Instances of non-monogamy have been observed in common ravens, by males visiting a female's nest when her mate is away. Breeding pairs must have a territory of their own before they begin nest-building and reproduction, and thus they aggressively defend a territory and its food resources. Nesting territories vary in size according to the density of food resources in the area. The nest is a deep bowl made of large sticks and twigs, bound with an inner layer of roots, mud, and bark and lined with a softer material, such as
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
fur. The nest is usually placed in a large tree or on a cliff ledge, or less frequently in old buildings or utility poles. Females lay between three and seven pale bluish-green, brown-blotched eggs. Incubation is about 18 to 21 days, by the female only. The male may stand or crouch over the young, sheltering but not actually brooding them. Young
fledge Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
at 35 to 42 days, and are fed by both parents. They stay with their parents for another six months after fledging. In most of their range, egg-laying begins in late February. In colder climates, it is later, e.g. April in
Greenland Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland is t ...
and
Tibet Tibet (; ''Böd''; ) is a region in East Asia, covering much of the Tibetan Plateau and spanning about . It is the traditional homeland of the Tibetan people. Also resident on the plateau are some other ethnic groups such as Monpa people, ...
. In Pakistan, egg-laying takes place in December. Eggs and hatchlings are preyed on, rarely, by large
hawk Hawks are bird of prey, birds of prey of the family Accipitridae. They are widely distributed and are found on all continents except Antarctica. * The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks and others. Th ...
s and
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, just ...
s, large
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s,
marten A marten is a weasel-like mammal in the genus ''Martes'' within the subfamily Guloninae, in the family Mustelidae. They have bushy tails and large paws with partially retractile claws. The fur varies from yellowish to dark brown, depending on t ...
s and
canid Canidae (; from Latin, '' canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found withi ...
s. The adults, which are very rarely preyed upon, are often successful in defending their young from these predators, due to their numbers, large size and cunning. They have been observed dropping stones on potential predators that venture close to their nests. Common ravens can be very long-lived, especially in captive or protected conditions; individuals at the Tower of London have lived for more than 40 years. Lifespans in the wild are considerably shorter at typically 10 to 15 years. The longest known lifespan of a banded wild common raven was 23 years, 3 months, which among passerines only is surpassed by a few Australian species such as the
satin bowerbird The satin bowerbird (''Ptilonorhynchus violaceus'') is a bowerbird endemic to eastern Australia. A rare natural intergeneric hybrid between the satin bowerbird and the regent bowerbird is known as Rawnsley's bowerbird. Description Mature mal ...
.


Feeding

Common ravens are
omnivorous An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutri ...
and highly
opportunistic Opportunism is the practice of taking advantage of circumstances – with little regard for principles or with what the consequences are for others. Opportunist actions are expedient actions guided primarily by self-interested motives. The term ...
: their diet may vary widely with location, season and
serendipity Serendipity is an unplanned fortunate discovery. Serendipity is a common occurrence throughout the history of product invention and scientific discovery. Etymology The first noted use of "serendipity" was by Horace Walpole on 28 January 1754. I ...
. For example, those foraging on
tundra In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless moun ...
on the Arctic North Slope of
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
obtained about half their energy needs from
predation Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
, mainly of microtine rodents, and half by scavenging, mainly of
caribou Reindeer (in North American English, known as caribou if wild and ''reindeer'' if domesticated) are deer in the genus ''Rangifer''. For the last few decades, reindeer were assigned to one species, ''Rangifer tarandus'', with about 10 subspe ...
and
ptarmigan ''Lagopus'' is a small genus of birds in the grouse subfamily commonly known as ptarmigans (). The genus contains three living species with numerous described subspecies, all living in tundra or cold upland areas. Taxonomy and etymology The ge ...
carcasses. In some places they are mainly
scavenger Scavengers are animals that consume dead organisms that have died from causes other than predation or have been killed by other predators. While scavenging generally refers to carnivores feeding on carrion, it is also a herbivorous feeding b ...
s, feeding on
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
as well as the associated
maggot A maggot is the larva of a fly (order Diptera); it is applied in particular to the larvae of Brachycera flies, such as houseflies, cheese flies, and blowflies, rather than larvae of the Nematocera, such as mosquitoes and crane flies. ...
s and
carrion beetle Silphidae is a family of beetles that are known commonly as large carrion beetles, carrion beetles or burying beetles. There are two subfamilies: Silphinae and Nicrophorinae. Nicrophorines are sometimes known as sexton beetles. The number of sp ...
s. With large-bodied carrion, which they are not equipped to tear through as well as birds such as hook-billed
vulture A vulture is a bird of prey that scavenges on carrion. There are 23 extant species of vulture (including Condors). Old World vultures include 16 living species native to Europe, Africa, and Asia; New World vultures are restricted to North and ...
s, they must wait for the prey to be torn open by another predator or flayed by other means. They are also known to eat the afterbirth of ewes and other large mammals. Plant food includes cereal grains, acorns, buds, berries and fruit. They prey on small
invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...
s, amphibians, reptiles, small mammals and birds. Ravens may also consume the undigested portions of animal feces, and human food waste. They store surplus food items, especially those containing fat, and will learn to hide such food out of the sight of other common ravens. Ravens also raid the food caches of other species, such as the
Arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small fox native to the Arctic regions of the Northern Hemisphere and common throughout the Arctic tundra biome. It is well adapted to living in co ...
. They sometimes associate with another canine, the
grey wolf The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly ...
, as a
kleptoparasite Kleptoparasitism (etymologically, parasitism by theft) is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct feeding, which can mean when fo ...
, following to scavenge wolf-kills in winter. Ravens are regular predators at bird nests, brazenly picking off eggs, nestlings and sometimes adult birds when they spot an opportunity. They are considered perhaps the primary natural threat to the nesting success of the critically endangered
California condor The California condor (''Gymnogyps californianus'') is a New World vulture and the largest North American land bird. It became extinct in the wild in 1987 when all remaining wild individuals were captured, but has since been reintroduced to nor ...
, since they readily take condor eggs and are very common in the areas where the species is being re-introduced. On the other hand, when they defend their own adjacent nests, they may incidentally benefit condors since they chase
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of p ...
s out of the area that may otherwise prey upon larger nestling and fledging condors. Condors, despite their large size, do not seem to have well developed nest defenses. Common ravens nesting near sources of human garbage included a higher percentage of food waste in their diet, birds nesting near roads consumed more road-killed
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...
s, and those nesting far from these sources of food ate more
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...
s and plant material.
Fledging Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
success was higher for those using human garbage as a food source. In contrast, a 1984–1986 study of common raven diet in an agricultural region of southwestern
Idaho Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and Wyom ...
found that cereal grains were the principal constituent of pellets, though small mammals, grasshoppers, cattle carrion and birds were also eaten. One behaviour is recruitment, where juvenile ravens call other ravens to a food bonanza, usually a carcass, with a series of loud yells. In '' Ravens in Winter'',
Bernd Heinrich Bernd Heinrich (born April 19, 1940 in Bad Polzin, Germany), is a professor emeritus in the biology department at the University of Vermont and is the author of a number of books about nature writing and biology. Heinrich has made major contribut ...
posited that this behaviour evolved to allow the juveniles to outnumber the resident adults, thus allowing them to feed on the carcass without being chased away. A more mundane explanation is that individuals co-operate in sharing information about carcasses of large mammals because they are too big for just a few birds to exploit.Heinrich, B. (1999). ''Mind of the Raven: Investigations and Adventures with Wolf-Birds'' pp. 119–120. New York: Cliff Street Books. Experiments with baits however show that such recruitment behaviour is independent of the size of the bait. Furthermore, there has been research suggesting that the common raven is involved in seed dispersal. In the wild, the common raven chooses the best habitat and disperses seeds in locations best suited for its survival.


Intelligence

The brain of the common raven is among the largest of any bird species. Specifically, their hyperpallium is large for a bird. They display ability in problem-solving, as well as other
cognitive Cognition refers to "the mental action or process of acquiring knowledge and understanding through thought, experience, and the senses". It encompasses all aspects of intellectual functions and processes such as: perception, attention, thought, ...
processes such as
imitation Imitation (from Latin ''imitatio'', "a copying, imitation") is a behavior whereby an individual observes and replicates another's behavior. Imitation is also a form of that leads to the "development of traditions, and ultimately our culture. ...
and
insight Insight is the understanding of a specific cause and effect within a particular context. The term insight can have several related meanings: *a piece of information *the act or result of understanding the inner nature of things or of seeing intu ...
. Linguist
Derek Bickerton Derek Bickerton (March 25, 1926 – March 5, 2018) was an English-born linguist and professor at the University of Hawaii in Manoa. Based on his work in creole languages in Guyana and Hawaii, he has proposed that the features of creole languages ...
, building on the work of biologist
Bernd Heinrich Bernd Heinrich (born April 19, 1940 in Bad Polzin, Germany), is a professor emeritus in the biology department at the University of Vermont and is the author of a number of books about nature writing and biology. Heinrich has made major contribut ...
, has argued that ravens are one of only four known animals (the others being bees,
ant Ants are eusocial insects of the family Formicidae and, along with the related wasps and bees, belong to the order Hymenoptera. Ants evolved from vespoid wasp ancestors in the Cretaceous period. More than 13,800 of an estimated total of 22 ...
s, and humans) who have demonstrated
displacement Displacement may refer to: Physical sciences Mathematics and Physics *Displacement (geometry), is the difference between the final and initial position of a point trajectory (for instance, the center of mass of a moving object). The actual path ...
, the capacity to communicate about objects or events that are distant in space or time. Subadult ravens roost together at night, but usually forage alone during the day. However, when one discovers a large carcass guarded by a pair of adult ravens, the unmated raven will return to the roost and communicate the find. The following day, a flock of unmated ravens will fly to the carcass and chase off the adults. Bickerton argues that the advent of linguistic displacement was perhaps the most important event in the evolution of human language, and that ravens are the only other
vertebrate Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...
to share this with humans. One experiment designed to evaluate insight and
problem-solving Problem solving is the process of achieving a goal by overcoming obstacles, a frequent part of most activities. Problems in need of solutions range from simple personal tasks (e.g. how to turn on an appliance) to complex issues in business an ...
ability involved a piece of meat attached to a string hanging from a perch. To reach the food, the bird needed to stand on the perch, pull the string up a little at a time, and step on the loops to gradually shorten the string. Four of five common ravens eventually succeeded, and "the transition from no success (ignoring the food or merely yanking at the string) to constant reliable access (pulling up the meat) occurred with no demonstrable
trial-and-error Trial and error is a fundamental method of problem-solving characterized by repeated, varied attempts which are continued until success, or until the practicer stops trying. According to W.H. Thorpe, the term was devised by C. Lloyd Morgan (1 ...
learning." This supports the hypothesis that common ravens are 'inventors', implying that they can solve problems. Many of the feats of common ravens were formerly argued to be stereotyped innate behaviour, but it now has been established that their aptitudes for solving problems individually and learning from each other reflect a flexible capacity for intelligent insight unusual among non-human animals. Another experiment showed that some common ravens could intentionally deceive their conspecifics. A study published in 2011 found that ravens can recognise when they are given an unfair trade during reciprocal interactions with conspecifics or humans, retaining memory of the interaction for a prolonged period of time. Birds that were given a fair trade by experimenters were found to prefer interacting with these experimenters compared to those that did not. Furthermore, ravens in the wild have also been observed to stop cooperating with other ravens if they observe them cheating during group tasks. Common ravens have been observed calling
wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
to the site of dead animals. The
wolves The wolf (''Canis lupus''; plural, : wolves), also known as the gray wolf or grey wolf, is a large Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been reco ...
open the carcass, leaving the scraps more accessible to the birds. They watch where other common ravens bury their food and remember the locations of each other's food caches, so they can steal from them. This type of theft occurs so regularly that common ravens will fly extra distances from a food source to find better hiding places for food. They have also been observed pretending to make a cache without actually depositing the food, presumably to confuse onlookers. Common ravens are known to steal and cache shiny objects such as pebbles, pieces of metal, and golf balls. One theory is that they hoard shiny objects to impress other ravens. Other research indicates that juveniles are deeply curious about all new things, and that common ravens retain an attraction to bright, round objects based on their similarity to bird eggs. Mature birds lose their intense interest in the unusual, and become highly neophobic. The first large-scale assessment of ravens' cognitive abilities suggests that, by four months of age, ravens do about as well as adult chimps and
orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genus ...
s on tests of
causal reasoning Causal reasoning is the process of identifying causality: the relationship between a cause and its effect. The study of causality extends from ancient philosophy to contemporary neuropsychology; assumptions about the nature of causality may be ...
, social learning,
theory of mind In psychology, theory of mind refers to the capacity to understand other people by ascribing mental states to them (that is, surmising what is happening in their mind). This includes the knowledge that others' mental states may be different fro ...
, etc.


Play

There has been increasing recognition of the extent to which birds engage in
play Play most commonly refers to: * Play (activity), an activity done for enjoyment * Play (theatre), a work of drama Play may refer also to: Computers and technology * Google Play, a digital content service * Play Framework, a Java framework * Pla ...
. Juvenile common ravens are among the most playful of bird species. They have been observed to slide down snowbanks, apparently purely for fun. They even engage in games with other species, such as playing catch-me-if-you-can with wolves, otters and dogs. Common ravens are known for spectacular aerobatic displays, such as flying in loops or interlocking talons with each other in flight. They are also one of only a few wild animals who make their own toys. They have been observed breaking off twigs to play with socially.


Relationship with humans


Conservation and management

Compared to many smaller ''
Corvus ''Corvus'' is a widely distributed genus of medium-sized to large birds in the family Corvidae. It includes species commonly known as crows, ravens and Rook (bird), rooks. The species commonly encountered in Europe are the carrion crow, the hoo ...
'' species (such as
American crow The American crow (''Corvus brachyrhynchos'') is a large passerine bird species of the family Corvidae. It is a common bird found throughout much of North America. American crows are the New World counterpart to the carrion crow of Europe an ...
), ravens prefer undisturbed mountain or forest habitat or rural areas over urban areas. In other areas, their numbers have increased dramatically and they have become agricultural
pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...
. Common ravens can cause damage to crops, such as nuts and grain, or can harm livestock, particularly by killing young goat kids, lambs and calves. Ravens generally attack the faces of young livestock, but the more common raven behaviour of scavenging may be misidentified as predation by ranchers. In the western
Mojave Desert The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily in ...
, human settlement and land development have led to an estimated 16-fold increase in the common raven population over 25 years. Towns, landfills, sewage treatment plants and artificial ponds create sources of food and water for scavenging birds. Ravens also find nesting sites in utility poles and ornamental trees, and are attracted to roadkill on highways. The explosion in the common raven population in the Mojave has raised concerns for the
desert tortoise The desert tortoise (''Gopherus agassizii''), is a species of tortoise in the Family (biology), family Testudinidae. The species is native to the Mojave Desert, Mojave and Sonoran Deserts of the southwestern United States and northwestern Mexic ...
, a
threatened species Threatened species are any species (including animals, plants and fungi) which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensa ...
. Common ravens prey upon juvenile tortoises, which have soft shells and move slowly. Plans to control the population have included shooting and trapping birds, as well as contacting landfill operators to ask that they reduce the amount of exposed garbage. A hunting bounty as a method of control was historically used in Finland from the mid-18th century until 1923. Culling has taken place to a limited extent in
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U.S., ...
, where the population increase in common ravens is threatening the vulnerable
Steller's eider Steller's eider (''Polysticta stelleri'') is a migrating Arctic diving duck that breeds along the coastlines of eastern Russia and Alaska. It is the rarest, smallest, and fastest flying of the eider species. Amongst the Inupiat, Steller's eider is ...
(''Polysticta stelleri''). Ravens, like other corvids, are definitive hosts of West Nile Virus (WNV). The transmission can be from infected birds to humans, and ravens are susceptible to WNV. However, in a 2010 study, it was shown that the California Common Ravens did not have a high positivity rate of WNV.


Cultural depictions

Across its range in the Northern Hemisphere, and throughout human history, the common raven has been a powerful symbol and a popular subject of mythology and folklore. In some Western traditions, ravens have long been considered to be birds of ill omen, death and evil in general, in part because of the negative symbolism of their all-
black Black is a color which results from the absence or complete absorption of visible light. It is an achromatic color, without hue, like white and grey. It is often used symbolically or figuratively to represent darkness. Black and white have o ...
plumage and the eating of
carrion Carrion () is the decaying flesh of dead animals, including human flesh. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures, c ...
. In Sweden, ravens are known as the ghosts of murdered people, and in Germany as the souls of the damned. In
Danish folklore Danish folklore consists of folk tales, legends, songs, music, dancing, popular beliefs, myths and traditions communicated by the inhabitants of towns and villages across the country, often passed on from generation to generation by word of mout ...
,
valravn In Danish folklore, a valravn ( Danish "raven of the slain") is a supernatural raven. Those ravens appear in traditional Danish folksongs, where they are described as originating from ravens who consume the bodies of the dead on the battlefield, a ...
e that ate a king's heart gained human knowledge, could perform great malicious acts, could lead people astray, had superhuman powers, and were "terrible animals".Kristensen, Evald Tang. (1980) ''Danske Sagn: Som De Har Lyd I Folkemunde'', Nyt Nordisk Forlag Arnold Busck, Copenhagen. . p. 132. As in traditional mythology and folklore, the common raven features frequently in more modern writings such as the works of
William Shakespeare William Shakespeare ( 26 April 1564 – 23 April 1616) was an English playwright, poet and actor. He is widely regarded as the greatest writer in the English language and the world's pre-eminent dramatist. He is often called England's nation ...
, and, perhaps most famously, in the poem "
The Raven "The Raven" is a narrative poem by American writer Edgar Allan Poe. First published in January 1845, the poem is often noted for its musicality, stylized language, and supernatural atmosphere. It tells of a distraught lover who is paid a myste ...
" by
Edgar Allan Poe Edgar Allan Poe (; Edgar Poe; January 19, 1809 – October 7, 1849) was an American writer, poet, editor, and literary critic. Poe is best known for his poetry and short stories, particularly his tales of mystery and the macabre. He is wide ...
. Ravens have appeared in the works of
Charles Dickens Charles John Huffam Dickens (; 7 February 1812 – 9 June 1870) was an English writer and social critic. He created some of the world's best-known fictional characters and is regarded by many as the greatest novelist of the Victorian e ...
,
J. R. R. Tolkien John Ronald Reuel Tolkien (, ; 3 January 1892 – 2 September 1973) was an English writer and philologist. He was the author of the high fantasy works ''The Hobbit'' and ''The Lord of the Rings''. From 1925 to 1945, Tolkien was the Rawlins ...
,
Stephen King Stephen Edwin King (born September 21, 1947) is an American author of horror, supernatural fiction, suspense, crime, science-fiction, and fantasy novels. Described as the "King of Horror", a play on his surname and a reference to his high s ...
,
George R. R. Martin George Raymond Richard Martin (born George Raymond Martin; September 20, 1948), also known as GRRM, is an American novelist, screenwriter, television producer and short story writer. He is the author of the series of epic fantasy novels ''A Song ...
and
Joan Aiken Joan Delano Aiken (4 September 1924 – 4 January 2004) was an English writer specialising in supernatural fiction and children's alternative history novels. In 1999 she was awarded an MBE for her services to children's literature. For ''The ...
among others. It continues to be used as a symbol in areas where it once had mythological status: as the
national bird This is a list of national birds, including official birds of overseas territories and other states described as nations. Most species in the list are officially designated. Some species hold only an "unofficial" status. National birds See al ...
of
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainous ...
( Kings of Bhutan wear the
Raven Crown The Raven Crown (Dzongkha: དབུ་ཞྭ་བྱ་རོག་ཅན་; Wylie: ''dbu-zhva bya-rog-can'') is worn by the Kings of Bhutan. It is a hat surmounted by the head of a raven. History The hereditary monarchy of the Wangchuck dy ...
), official bird of the
Yukon Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...
territory, and on the coat of arms of the
Isle of Man ) , anthem = "O Land of Our Birth" , image = Isle of Man by Sentinel-2.jpg , image_map = Europe-Isle_of_Man.svg , mapsize = , map_alt = Location of the Isle of Man in Europe , map_caption = Location of the Isle of Man (green) in Europe ...
(once a Viking colony). In Persia and Arabia the raven was held as a bird of bad omen but a 14th-century Arabic work reports use of the raven in falconry. The modern unisex given name '' Raven'' is derived from the English word "raven". As a masculine name, ''Raven'' parallels the Old Norse ''
Hrafn Hrafn (; ) is both a masculine byname, and personal name in Old Norse. The name translates into English as " raven". The Old English form of the name is ''*Hræfn''. The name is paralleled by the English masculine given name '' Raven'', which is ...
'', and Old English ''*Hræfn'', which were both
byname An epithet (, ), also byname, is a descriptive term (word or phrase) known for accompanying or occurring in place of a name and having entered common usage. It has various shades of meaning when applied to seemingly real or fictitious people, di ...
s and
personal name A personal name, or full name, in onomastic terminology also known as prosoponym (from Ancient Greek πρόσωπον / ''prósōpon'' - person, and ὄνομα / ''onoma'' - name), is the set of names by which an individual person is known ...
s.


Mythology

In
Tlingit The Tlingit ( or ; also spelled Tlinkit) are indigenous peoples of the Pacific Northwest Coast of North America. Their language is the Tlingit language (natively , pronounced ),
and Haida cultures, Raven was both a
trickster In mythology and the study of folklore and religion, a trickster is a character in a story (god, goddess, spirit, human or anthropomorphisation) who exhibits a great degree of intellect or secret knowledge and uses it to play tricks or otherwi ...
and
creator god A creator deity or creator god (often called the Creator) is a deity responsible for the creation of the Earth, world, and universe in human religion and mythology. In monotheism, the single God is often also the creator. A number of monolatris ...
. Related beliefs are widespread among the peoples of
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
and northeastern Asia. The
Kamchatka Peninsula The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and we ...
, for example, was supposed to have been created by the raven god
Kutkh Kutkh (also ''Kutkha'', ''Kootkha'', ''Kutq,'' ''Kutcha'' and other variants, russian: Кутх) is a Raven spirit traditionally revered in various forms by various indigenous peoples of the Russian Far East. Kutkh appears in many legends: as a k ...
. There are several references to common ravens in the
Old Testament The Old Testament (often abbreviated OT) is the first division of the Christian biblical canon, which is based primarily upon the 24 books of the Hebrew Bible or Tanakh, a collection of ancient religious Hebrew writings by the Israelites. The ...
of the
Bible The Bible (from Koine Greek , , 'the books') is a collection of religious texts or scriptures that are held to be sacred in Christianity, Judaism, Samaritanism, and many other religions. The Bible is an anthologya compilation of texts of a ...
and it is an aspect of
Mahakala Mahākāla is a deity common to Hinduism and Tantric Buddhism. In Buddhism, Mahākāla is regarded as the sacred '' Dharmapāla'' ("Protector of the Dharma"), while in Hinduism, Mahākāla is a fierce manifestation of the Hindu god Shiva and th ...
in
Bhutan Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountainous ...
ese mythology. In
Norse mythology Norse, Nordic, or Scandinavian mythology is the body of myths belonging to the North Germanic peoples, stemming from Old Norse religion and continuing after the Christianization of Scandinavia, and into the Nordic folklore of the modern period ...
,
Huginn In Norse mythology, Huginn (Old Norse: "thought"Orchard (1997:92).) and Muninn (Old Norse "memory"Orchard (1997:115). or "mind"Lindow (2001:186).) are a pair of common raven, ravens that fly all over the world, Midgard, and bring information to t ...
(from the Old Norse for "thought") and
Muninn In Norse mythology, Huginn (Old Norse: "thought"Orchard (1997:92).) and Muninn (Old Norse "memory"Orchard (1997:115). or "mind"Lindow (2001:186).) are a pair of ravens that fly all over the world, Midgard, and bring information to the god Odin. ...
(from the Old Norse for "memory" or "mind") are a pair of ravens that fly all over the world of humans,
Midgard In Germanic cosmology, Midgard (an anglicised form of Old Norse ; Old English , Old Saxon , Old High German , and Gothic ''Midjun-gards''; "middle yard", "middle enclosure") is the name for Earth (equivalent in meaning to the Greek term , "inhab ...
, and bring the god
Odin Odin (; from non, Óðinn, ) is a widely revered Æsir, god in Germanic paganism. Norse mythology, the source of most surviving information about him, associates him with wisdom, healing, death, royalty, the gallows, knowledge, war, battle, v ...
information. Additionally among the Norse,
raven banner The raven banner ( non, hrafnsmerki ; enm, hravenlandeye) was a flag, possibly totemic in nature, flown by various Viking chieftains and other Scandinavian rulers during the 9th, 10th and 11th centuries. The flag, as depicted in Norse artwork, ...
standards were carried by such figures as the Jarls of Orkney, King
Cnut the Great Cnut (; ang, Cnut cyning; non, Knútr inn ríki ; or , no, Knut den mektige, sv, Knut den Store. died 12 November 1035), also known as Cnut the Great and Canute, was King of England from 1016, King of Denmark from 1018, and King of Norwa ...
of England, Norway and Denmark, and
Harald Hardrada Harald Sigurdsson (; – 25 September 1066), also known as Harald III of Norway and given the epithet ''Hardrada'' (; modern no, Hardråde, roughly translated as "stern counsel" or "hard ruler") in the sagas, was King of Norway from 1046 t ...
. In the British Isles, ravens also were symbolic to the
Celts The Celts (, see pronunciation for different usages) or Celtic peoples () are. "CELTS location: Greater Europe time period: Second millennium B.C.E. to present ancestry: Celtic a collection of Indo-European peoples. "The Celts, an ancien ...
. In
Irish mythology Irish mythology is the body of myths native to the island of Ireland. It was originally passed down orally in the prehistoric era, being part of ancient Celtic religion. Many myths were later written down in the early medieval era by C ...
, the goddess Morrígan alighted on the hero
Cú Chulainn Cú Chulainn ( ), called the Hound of Ulster (Irish: ''Cú Uladh''), is a warrior hero and demigod in the Ulster Cycle of Irish mythology, as well as in Scottish and Manx folklore. He is believed to be an incarnation of the Irish god Lugh, ...
's shoulder in the form of a raven after his death. In
Welsh mythology Welsh mythology (Welsh: ''Mytholeg Cymru'') consists of both folk traditions developed in Wales, and traditions developed by the Celtic Britons elsewhere before the end of the first millennium. As in most of the predominantly oral societies Cel ...
they were associated with the Welsh god
Bran the Blessed Bran, also known as miller's bran, is the hard outer layers of cereal grain. It consists of the combined aleurone and pericarp. Corn (maize) bran also includes the pedicel (tip cap). Along with germ, it is an integral part of whole grains, a ...
, whose name translates to "raven." According to the ''
Mabinogion The ''Mabinogion'' () are the earliest Welsh prose stories, and belong to the Matter of Britain. The stories were compiled in Middle Welsh in the 12th–13th centuries from earlier oral traditions. There are two main source manuscripts, create ...
'', Bran's head was buried in the White Hill of London as a
talisman A talisman is any object ascribed with religious or magical powers intended to protect, heal, or harm individuals for whom they are made. Talismans are often portable objects carried on someone in a variety of ways, but can also be installed perm ...
against invasion. A legend developed that England would not fall to a foreign invader as long as there were ravens at the Tower of London; although this is often thought to be an ancient belief, the official Tower of London historian, Geoff Parnell, believes that this is actually a romantic Victorian invention. In the
Jewish Jews ( he, יְהוּדִים, , ) or Jewish people are an ethnoreligious group and nation originating from the Israelites Israelite origins and kingdom: "The first act in the long drama of Jewish history is the age of the Israelites""The ...
,
Christian Christians () are people who follow or adhere to Christianity, a monotheistic Abrahamic religion based on the life and teachings of Jesus Christ. The words ''Christ'' and ''Christian'' derive from the Koine Greek title ''Christós'' (Χρι ...
and
Islamic Islam (; ar, ۘالِإسلَام, , ) is an Abrahamic monotheistic religion centred primarily around the Quran, a religious text considered by Muslims to be the direct word of God (or '' Allah'') as it was revealed to Muhammad, the mai ...
traditions, the raven was the first animal to be released from
Noah's Ark Noah's Ark ( he, תיבת נח; Biblical Hebrew: ''Tevat Noaḥ'')The word "ark" in modern English comes from Old English ''aerca'', meaning a chest or box. (See Cresswell 2010, p.22) The Hebrew word for the vessel, ''teva'', occurs twice in t ...
. "So it came to pass, at the end of forty days, that Noah opened the window of the ark which he had made. Then he sent out a raven, which kept going to and fro until the waters had dried up from the earth. He also sent out from himself a dove, to see if the waters had receded from the face of the ground." The raven is mentioned 12 times in the Bible. In the
New Testament The New Testament grc, Ἡ Καινὴ Διαθήκη, transl. ; la, Novum Testamentum. (NT) is the second division of the Christian biblical canon. It discusses the teachings and person of Jesus, as well as events in first-century Christ ...
Jesus Jesus, likely from he, יֵשׁוּעַ, translit=Yēšūaʿ, label=Hebrew/Aramaic ( AD 30 or 33), also referred to as Jesus Christ or Jesus of Nazareth (among other names and titles), was a first-century Jewish preacher and religious ...
tells a parable using the raven to show how people should rely on God for their needs and not riches ( Luke 12:24).Luke 12:24 (New Living Translation)
Biblegateway.com. Retrieved on 2012-12-19.
The raven is also mentioned in the
Quran The Quran (, ; Standard Arabic: , Classical Arabic, Quranic Arabic: , , 'the recitation'), also romanized Qur'an or Koran, is the central religious text of Islam, believed by Muslims to be a revelation in Islam, revelation from God in Islam, ...
at the story of
Cain and Abel In the biblical Book of Genesis, Cain ''Qayīn'', in pausa ''Qāyīn''; gr, Κάϊν ''Káïn''; ar, قابيل/قايين, Qābīl / Qāyīn and Abel ''Heḇel'', in pausa ''Hāḇel''; gr, Ἅβελ ''Hábel''; ar, هابيل, Hāb ...
.
Adam Adam; el, Ἀδάμ, Adám; la, Adam is the name given in Genesis 1-5 to the first human. Beyond its use as the name of the first man, ''adam'' is also used in the Bible as a pronoun, individually as "a human" and in a collective sense as " ...
's firstborn son Cain kills his brother Abel, but he does not know what to do with the corpse: "Then Allah sent a raven scratching up the ground, to show him how to hide his brother's naked corpse. He said: Woe unto me! Am I not able to be as this raven and so hide my brother's naked corpse? And he became repentant."(بَعَثَ اللّهُ غُرَابًا يَبْحَثُ فِي الأَرْضِ لِيُرِيَهُ كَيْفَ يُوَارِي سَوْءةَ أَخِيهِ قَالَ يَا وَيْلَتَا أَعَجَزْتُ أَنْ أَكُونَ مِثْلَ هَذَا الْغُرَابِ فَأُوَارِيَ سَوْءةَ أَخِي فَأَصْبَحَ مِنَ النَّادِمِينَ) Qur'an 5:31, translation: Pickthall, Marmaduke ''The Meaning of the Glorious Koran: An Explanatory Translation'', Amana Publications, 1st edition (1996)


References


Cited texts

* * *


Further reading

* Heinrich, B. (1999). ''Mind of the Raven: Investigations and Adventures with Wolf-Birds.'' New York: Cliff Street Books.


External links


RSPB: Raven
– Cornell Lab of Ornithology *
Ageing and sexing (PDF; 3.1 MB) by Javier Blasco-Zumeta & Gerd-Michael HeinzeCommon raven images at ARKive''Corvus corax''
at Encyclopedia of Life

– USGS Patuxent Bird Identification InfoCenter (includes CBC/BBS range maps) *

at naturesongs.com {{Featured article Corvus Holarctic birds Ravens Talking birds Territorial symbols of Yukon Birds described in 1758 Articles containing video clips Fauna of the San Francisco Bay Area Taxa named by Carl Linnaeus