''Cyphornis'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of the

prehistoric Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of ...

pseudotooth bird The Pelagornithidae, commonly called pelagornithids, pseudodontorns, bony-toothed birds, false-toothed birds or pseudotooth birds, are a prehistoric family (biology), family of large seabirds. Their fossil remains have been found all over the wor ...

s. These were probably rather close relatives of either

pelican Pelicans (genus ''Pelecanus'') are a genus of large water birds that make up the family Pelecanidae. They are characterized by a long beak and a large throat pouch used for catching prey and draining water from the scooped-up contents before s ...

s and

stork Storks are large, long-legged, long-necked wading birds with long, stout bills. They belong to the family called Ciconiidae, and make up the order Ciconiiformes . Ciconiiformes previously included a number of other families, such as herons an ...

s, or of

waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which in ...

, and are here placed in the

order Order, ORDER or Orders may refer to: * Categorization, the process in which ideas and objects are recognized, differentiated, and understood * Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of d ...

Odontopterygiformes The Pelagornithidae, commonly called pelagornithids, pseudodontorns, bony-toothed birds, false-toothed birds or pseudotooth birds, are a prehistoric family of large seabirds. Their fossil remains have been found all over the world in rocks dating ...

to account for this uncertainty.

Description

Only a single

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

, ''Cyphornis magnus'', is known to date. It is only known with certainty from a single specimen, the rather abraded

proximal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

part of a left

tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) and meta ...

which was found at Carmanah Point on

Vancouver Island Vancouver Island is an island in the northeastern Pacific Ocean and part of the Canadian Provinces and territories of Canada, province of British Columbia. The island is in length, in width at its widest point, and in total area, while are o ...

(

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tot ...

), where the

Juan de Fuca Strait The Strait of Juan de Fuca (officially named Juan de Fuca Strait in Canada) is a body of water about long that is the Salish Sea's outlet to the Pacific Ocean. The international boundary between Canada and the United States runs down the centre ...

opens into the

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

. The deposits from which it originated were initially dated to the

Eocene The Eocene ( ) Epoch is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene' ...

; subsequent authors have usually assigned them to the

Early Miocene The Early Miocene (also known as Lower Miocene) is a sub-epoch of the Miocene Epoch made up of two stages: the Aquitanian and Burdigalian stages. The sub-epoch lasted from 23.03 ± 0.05 Ma to 15.97 ± 0.05 Ma (million years ago). It was prece ...

though certainly rocks from around the Eo-

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...

boundary also occur in the region where it was found. At the time of its discovery, it "probably represent dthe largest known bird of flight." Even today it is one of the largest (though not heaviest) flying birds known. Some huge pseudotooth wing bone fossils have been found in

Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...

. Specimen

LACM The Natural History Museum of Los Angeles County is the largest natural and historical museum in the western United States. Its collections include nearly 35 million specimens and artifacts and cover 4.5 billion years of history. This large coll ...

128462, a mostly complete

end of a left

ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...

, originates from the Keasey Formation of Washington County. LACM 127875 are fragments of the proximal

humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...

ends, the proximal right ulna and the right radius of a single individual presumed to be of the same species; they were found in the Pittsburg Bluff Formation near

Mist Mist is a phenomenon caused by small droplets of water suspended in the cold air, usually by condensation. Physically, it is an example of a dispersion. It is most commonly seen where water vapor in warm, moist air meets sudden cooling, such a ...

. These remains all date from the Eo-Oligocene boundary, and considering their size they may well be of ''C. magnus'' if it is in fact that old, or of its ancestor or older relative.

Systematics

Due to its fragmentary nature – the bones of pseudotooth bords are very thin-walled and light and notoriously easily broken and crushed when

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

izing – it was often allied with the enigmatic ''

Cladornis The Pelecaniformes are an order of medium-sized and large waterbirds found worldwide. As traditionally—but erroneously—defined, they encompass all birds that have feet with all four toes webbed. Hence, they were formerly also known by such n ...

'', and though placed in the

Pelecaniformes The Pelecaniformes are an order of medium-sized and large waterbirds found worldwide. As traditionally—but erroneously—defined, they encompass all birds that have feet with all four toes webbed. Hence, they were formerly also known by such ...

(as pseudotooth birds often were) separated in a

suborder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...

Cladornithes. But the slightly older (

Late Oligocene The Chattian is, in the geologic timescale, the younger of two ages or upper of two stages of the Oligocene Epoch/Series. It spans the time between . The Chattian is preceded by the Rupelian and is followed by the Aquitanian (the lowest stage ...

) ''Cladornis'' from the

Argentinian Argentines (mistakenly translated Argentineans in the past; in Spanish (masculine) or ( feminine)) are people identified with the country of Argentina. This connection may be residential, legal, historical or cultural. For most Argentines, ...

part of

Patagonia Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and gl ...

is known from a

distal Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...

right tarsometatarsus only, and thus not directly comparable to ''Cyphornis''. The two genera were allied simply because of their size and because they both vaguely reminded of the tarsometatarsus of pelicans. Today however, ''Cladornis'' is more generally held to be a

terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...

bird rather than a

seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same enviro ...

. Other authors had been more conservative all along, and considered ''Cyphornis'' quite close to

s, uniting these as a

superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...

Pelecanides in suborder Pelecanae, or later on (after the endings of

taxonomic rank In biological classification, taxonomic rank is the relative level of a group of organisms (a taxon) in an ancestral or hereditary hierarchy. A common system consists of species, genus, family (biology), family, order (biology), order, class (b ...

s were fixed to today's standard) Pelecanoidea in suborder Pelecani. To set it apart from its alleged relatives, ''Cyphornis'' was early on separated in a

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Cyphornithidae together with '' Palaeochenoides mioceanus'' and eventually also '' Tympanonesiotes wetmorei'' which are also little-known pseudotooth birds but inhabited the

Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe an ...

. But these are probably all closely related to the better-known ''

Pelagornis ''Pelagornis'' is a widespread genus of prehistoric pseudotooth birds. These were probably rather close relatives of either pelicans and storks, or of waterfowl, and are here placed in the order Odontopterygiformes to account for this uncertain ...

'',

type genus In biological taxonomy, the type genus is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearing type of a nominal f ...

of the family Pelagornithidae. And even if ''Cyphornis'' is the

senior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linnae ...

of all the later-described genera (which is not very likely), according to the rules of

zoological nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the In ...

the family name Pelagornithidae would not change. Thus Cyphornithidae would almost certainly be a

junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...

of Pelagornithidae even if the pseudotooth birds are (as some have proposed) divided into several families – rather than being all placed in the Pelagornithidae as is usual nowadays –, as ''Cyphornis'', ''Osteodontornis'', ''Palaeochenoides'', ''Pelagornis'' and perhaps the smaller ''Tympanoneisiotes'' appear to be very closely related and are probably all part of a

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic gro ...

lineage of (usually) giant pseudotooth birds. Only if the

lineage is sufficiently distinct, the Cyphornithidae would remain valid, but in this case they would presumably not include the Atlantic forms.Stone (1928), Brodkorb (1963: pp. 264–265), Hopson (1964), Olson (1985: pp. 195–198), Mlíkovský (2002: p. 81), Mayr (2009: pp. 58–59)

Footnotes

References

Electronic supplement
* * * * * * * * * * {{Taxonbar, from=Q5200258 Pelagornithidae Miocene birds Miocene birds of North America Fossil taxa described in 1894