''Cyanothamnus quadrangulus'', commonly known as narrow-leaved boronia, is a plant in the citrus

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Rutaceae The Rutaceae is a family, commonly known as the rueRUTACEAE
in BoDD – Botanical Derm ...

and is

endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found elsew ...

to eastern

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

. It is an erect shrub with four-angled branches,

bipinnate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...

leaves and white, sometimes pale pink, four-

petal Petals are modified Leaf, leaves that surround the reproductive parts of flowers. They are often advertising coloration, brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''c ...

led flowers. Boronia anethifolia habit

Description

''Cyanothamnus quadrangulus'' is an erect shrub that grows to a height of with four-angled,

glabrous Glabrousness (from the Latin ''glaber'' meaning "bald", "hairless", "shaved", "smooth") is the technical term for a lack of hair, down, setae, trichomes or other such covering. A glabrous surface may be a natural characteristic of all or part of ...

stems with prominent leaf scars. The leaves are bipinnate, long and wide in outline and have a petiole long. The leaves have between five and eleven glabrous, linear to narrow elliptic leaflets. The end leaflet is long and wide, the others usually slightly longer. The flowers are white, sometimes pale pink and are arranged in leaf

axil A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...

s, mainly in groups of between three and fourteen or more. The groups are borne on a peduncle long. The four

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...

s are triangular to broadly egg-shaped, about long and wide. The four

s are long with their bases overlapping. The eight

stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filame ...

s have hairy edges. Flowering occurs from April to October and the fruit are glabrous, long and about wide.

Taxonomy and naming

This species was first formally described in 1863 by

Stephan Endlicher Stephan Ladislaus Endlicher also known as Endlicher István László (24 June 1804, Bratislava (Pozsony) – 28 March 1849, Vienna) was an Austrian botanist, numismatist and Sinologist. He was a director of the Botanical Garden of Vienna. Bio ...

from an unpublished manuscript by Allan Cunningham and was given the name ''Boronia anethifolia''. Endlicher's description was published in ''

Enumeratio plantarum quas in Novae Hollandiae ora austro-occidentali ad fluvium Cygnorum et in sinu Regis Georgii collegit Carolus Liber Baro de Hügel ''Enumeratio plantarum quas in Novae Hollandiæ ora austro-occidentali ad fluvium Cygnorum et in sinu Regis Georgii collegit Carolus Liber Baro de Hügel'' is a description of the plants collected at the Swan River colony and King George Sound in ...

''. In a 2013 paper in the journal ''

Taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

'',

Marco Duretto Marco Duretto (born 1964) is a manager and senior research scientist at the Royal Botanic Gardens Sydney in Australia. His primary research interests are systematics and conservation of Rutaceae, Rubiaceae, Orchidaceae, Stylidiaceae and evolut ...

and others moved this species to the genus ''Cyanothamnus'' on the basis of

cladistic analysis Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived chara ...

, but because the name ''Cyanothamnus anethifolia'' had already been used for a different taxon (''Cyanothamnus anethifolius'' Bartl.) the name ''Cyanothamnus quadrangulus'' was used. The

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

(''anethifolia'') referred to the deeply divided leaves with narrow leaflets which resemble those of

dill Dill (''Anethum graveolens'') is an annual herb in the celery family Apiaceae. It is the only species in the genus ''Anethum''. Dill is grown widely in Eurasia, where its leaves and seeds are used as a herb or spice for flavouring food. Growth ...

(''Anethum'') in the

Apiaceae Apiaceae or Umbelliferae is a family of mostly aromatic flowering plants named after the type genus ''Apium'' and commonly known as the celery, carrot or parsley family, or simply as umbellifers. It is the 16th-largest family of flowering plants ...

and the new epithet (''quadrangulus'') means "four-angled" and refers to the four-sided stems of this species.

Distribution and habitat

This boronia grows in heath and forest and in rocky slopes and ridges between the Border Ranges in

Queensland ) , nickname = Sunshine State , image_map = Queensland in Australia.svg , map_caption = Location of Queensland in Australia , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , established_ ...

and Wadbilliga Mountain in southern

New South Wales ) , nickname = , image_map = New South Wales in Australia.svg , map_caption = Location of New South Wales in AustraliaCoordinates: , subdivision_type = Country , subdivision_name = Australia , established_title = Before federation , es ...

References

{{Taxonbar, from=Q106902240 quadrangulus Flora of New South Wales Flora of Queensland Plants described in 1837 Taxa named by Stephan Endlicher