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Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily
Coccinelloidea Coccinelloidea is a superfamily of beetles in the order Coleoptera, formerly included in the superfamily Cucujoidea. There are more than 10,000 species in Coccinelloidea, including more than 6000 in the lady beetle family Coccinellidae. Morphol ...
. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark beetles ( Scolytinae), which are
weevil Weevils are beetles belonging to the Taxonomic rank, superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and Herbivore, herbivorous. Approximately 97,000 species of weevils are known. They b ...
s (superfamily
Curculionoidea Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, ...
).


Morphology

The morphology of Cucujoidea is varied and there are no features uniting all members of the superfamily. Adults can be recognised by the procoxal cavities being internally open in most taxa, females having tarsal formula 5-5-5 and males 5-5-5 or 5-5-4 (rarely 4-4-4), females with tergite VIII concealed dorsally by tergite VII, and males with tergite X completely membraneous. Larvae have frontal arms usually lyriform, the mandible mesal surface usually with well-developed mola, a maxillary articulating area usually present, a hypopharyngeal sclerome usually present, and two pretarsal setae.


Taxonomy

According to a 2015 revision, the following 25 families make up superfamily Cucujoidea: * Agapythidae Sen Gupta and Crowson, 1969 *
Boganiidae Boganiidae is a family of beetles, in the superfamily Cucujoidea. Members of the family are found in southern Africa, Australia and New Caledonia. Adults and larvae are pollenivorous, feeding on the pollen of cycads and flowering plants of the f ...
Sen Gupta & Crowson, 1966 * Cavognathidae Sen Gupta & Crowson, 1966 *
Cryptophagidae Cryptophagidae is a family of beetles with representatives found in all biogeographic realms. Members of this family are commonly called silken fungus beetles and both adults and larvae appear to feed exclusively on fungi although in a wide vari ...
Kirby, 1826
silken fungus beetle Cryptophagidae is a family of beetles with representatives found in all biogeographic realms. Members of this family are commonly called silken fungus beetles and both adults and larvae appear to feed exclusively on fungi although in a wide vari ...
s *
Cucujidae The Cucujidae, "flat bark beetles," are a family of distinctively flat beetles found worldwide (except Africa and Antarctica) under the bark of dead trees. The family has received considerable taxonomic attention in recent years and now consists ...
Latreille, 1802flat bark beetles * Cybocephalidae Jacquelin du Val, 1858 *
Cyclaxyridae Cyclaxyridae are a family of beetles in the superfamily Cucujoidea. The only living genus is '' Cyclaxyra'', with two species endemic to New Zealand. Other species have been named from fossils. They are also known as sooty mould beetles due to the ...
Gimmel, Leschen & Ślipiński, 2009 – sooty mould beetles *
Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present circumscription, it contains 6 tribes ( Tritomini, Dacnini, Megalodacnini, Er ...
Latreille, 1802
pleasing fungus beetle file:Ischyrus quadripunctatus P1390172a.jpg, ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family (biology), family of beetles belonging to Cucujoidea containing over 100 genus, genera. In the present circumscrip ...
s * Helotidae Reitter, 1876 * Hobartiidae Sen Gupta & Crowson, 1966 *
Kateretidae Kateretidae also known as short-winged flower beetles are a family of beetles in the superfamily Cucujoidea. There are 10 extant and 4 extinct genera, and at least 40 described species. They are found worldwide except in New Zealand. Adults are a ...
Erichson in Agassiz, 1846
short-winged flower beetle Kateretidae also known as short-winged flower beetles are a family of beetles in the superfamily Cucujoidea. There are 10 extant and 4 extinct genera, and at least 40 described species. They are found worldwide except in New Zealand. Adults are ...
s (= Brachypteridae) *
Laemophloeidae Laemophloeidae, "lined flat bark beetles," is a family in the superfamily Cucujoidea characterized by predominantly dorso-ventrally compressed bodies, head and pronotal discs bordered by ridges or grooves, and inverted male genitalia. Size range ...
Ganglbauer 1899lined flat bark beetles *
Lamingtoniidae ''Lamingtonium'' is the only genus in the family Lamingtoniidae, of the beetle superfamily Cucujoidea. It contains three species endemic to Australia.Lawrence, J. F., and Leschen, R. A. B. (2003). Review of Lamingtoniidae (Coleoptera: Cucujoidea ...
Sen Gupta & Crowson, 1966 *
Monotomidae Monotomidae is a family of beetles in the superfamily Cucujoidea. The family is found worldwide, with approximately 240 species in 33 genera. The ecological habits of the family are diverse, with different members of the group being found under t ...
Laporte, 1840root-eating beetles *
Myraboliidae ''Myrabolia'' is the only genus in the beetle family Myraboliidae in the superfamily Cucujoidea. It has about 13 species, found in Australia. Adults and possibly larvae live under the bark of ''Eucalyptus ''Eucalyptus'' () is a genus of over s ...
Lawrence and Britton, 1991 *
Nitidulidae The sap beetles, also known as Nitidulidae, are a family (biology), family of beetles. They are small (2–6 mm) ovoid, usually dull-coloured beetles, with knobbed antenna (biology), antennae. Some have red or yellow spots or bands. They fe ...
Latreille, 1802
sap beetle The sap beetles, also known as Nitidulidae, are a family of beetles. They are small (2–6 mm) ovoid, usually dull-coloured beetles, with knobbed antennae. Some have red or yellow spots or bands. They feed mainly on decaying vegetable ma ...
s * Passandridae Erichson, 1845parasitic flat bark beetles *
Phalacridae The Phalacridae are a family of beetles commonly called the shining flower beetles, They are often found in composite flowers. They are oval-shaped, usually tan, and aboout 2 mm in length. Most species feed on fungus, although a number feed ...
Leach, 1815shining flower beetles * Phloeostichidae Reitter, 1911 * Priasilphidae Crowson, 1973 * Protocucujidae Crowson, 1954 *
Silvanidae Silvanidae, "silvan flat bark beetles", is a family of beetles in the superfamily Cucujoidea,Thomas, M. C., and R.A. B. Leschen. 2010. Silvanidae Kirby, 1837. p. 346-350. In: Leschen, R.A.B., R.G. Beutel, and J.F. Lawrence. Coleoptera, Beetles. Vo ...
Kirby, 1937
silvanid flat bark beetle Silvanidae, "silvan flat bark beetles", is a family of beetles in the superfamily Cucujoidea,Thomas, M. C., and R.A. B. Leschen. 2010. Silvanidae Kirby, 1837. p. 346-350. In: Leschen, R.A.B., R.G. Beutel, and J.F. Lawrence. Coleoptera, Beetles. Vo ...
s * Smicripidae Horn, 1879palmetto beetles * Sphindidae Jacquelin du Val, 1860dry-fungus beetles * Tasmosalpingidae Lawrence and Britton, 1991 * Parandrexidae Kirejtshuk, 1994 (Middle Jurassic-Early Cretaceous)


Extinct genera

* '' Alloterocucus'' Li et al., 2022
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The ...
, Myanmar, mid-
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of th ...
(latest
Albian The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 M ...
- earliest
Cenomanian The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
)


References


External links

* * Beetle superfamilies Taxa named by Pierre André Latreille {{Cucujoidea-stub