''Cryptogramma crispa'', the parsley fern, is an

Arctic–alpine An Arctic–alpine taxon is one whose natural distribution includes the Arctic and more southerly mountain ranges, particularly the Alps. The presence of identical or similar taxa in both the tundra of the far north, and high mountain ranges much f ...

species of

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...

. It produces separate sterile and fertile fronds, up to tall, and is a pioneer species on acidic screes.

Description

The fronds of ''C. crispa'' are long and appear in two distinct forms. Sterile leaves are 2–3-

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...

with the pinnules long by wide, while fertile leaves are 3–4-pinnate, and with narrower pinnules. The fertile leaves have sori scattered along the veins, each with a strongly enrolled false

indusium A sorus (pl. sori) is a cluster of sporangia (structures producing and containing spores) in ferns and fungi. A coenosorus (plural coenosori) is a compound sorus composed of multiple, fused sori. Etymology This New Latin word is from Ancient ...

. The

sporangia A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cy ...

are yellow and mature around midsummer.

Distribution and ecology

''Cryptogramma crispa'' grows among acidic rocks in areas where snow lies until late in the year. It is a pioneer species on stable scree slopes and also occurs on

cliff In geography and geology, a cliff is an area of rock which has a general angle defined by the vertical, or nearly vertical. Cliffs are formed by the processes of weathering and erosion, with the effect of gravity. Cliffs are common on co ...

s and dry stone walls. In Europe, ''C. crispa'' has an

distribution, growing in the mountains of

Central Central is an adjective usually referring to being in the center of some place or (mathematical) object. Central may also refer to: Directions and generalised locations * Central Africa, a region in the centre of Africa continent, also known as ...

and

Southern Europe Southern Europe is the southern regions of Europe, region of Europe. It is also known as Mediterranean Europe, as its geography is essentially marked by the Mediterranean Sea. Definitions of Southern Europe include some or all of these countrie ...

, as well as in the north of the continent, including

Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion#Europe, subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, ...

and higher ground in the

British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...

. In

Ireland Ireland ( ; ga, Éire ; Ulster Scots dialect, Ulster-Scots: ) is an island in the Atlantic Ocean, North Atlantic Ocean, in Northwestern Europe, north-western Europe. It is separated from Great Britain to its east by the North Channel (Grea ...

, it is rare and concentrated in the east of the country, leading Praeger to conjecture that the Irish examples are recent colonists from

Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It is ...

, arriving as airborne spores. Similar plants, which may belong to the same species occur in

East Asia East Asia is the eastern region of Asia, which is defined in both geographical and ethno-cultural terms. The modern states of East Asia include China, Japan, Mongolia, North Korea, South Korea, and Taiwan. China, North Korea, South Korea and ...

and

North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...

, although these are usually considered a separate taxon.

Spore In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, f ...

s attributable to ''C. crispa'' have been discovered in deposits in

Snowdonia Snowdonia or Eryri (), is a mountainous region in northwestern Wales and a national park of in area. It was the first to be designated of the three national parks in Wales, in 1951. Name and extent It was a commonly held belief that the nam ...

from the last glacial period, as well as from more low-lying sites in

Cheshire Cheshire ( ) is a ceremonial and historic county in North West England, bordered by Wales to the west, Merseyside and Greater Manchester to the north, Derbyshire to the east, and Staffordshire and Shropshire to the south. Cheshire's county t ...

Taxonomic history

The parsley fern appeared in

Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...

' 1753 work ''

Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...

'', the starting point for

botanical nomenclature Botanical nomenclature is the formal, scientific naming of plants. It is related to, but distinct from Alpha taxonomy, taxonomy. Plant taxonomy is concerned with grouping and classifying plants; botanical nomenclature then provides names for the ...

, under the name ''Osmunda crispa''. The

specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...

''crispa'' means finely waved or curled. It is placed in the family Pteridaceae, part of the order Polypodiales.

References

External links

* * {{Taxonbar, from=Q1574713 crispa Alpine flora Flora of Europe Flora of temperate Asia Plants described in 1753 Taxa named by Carl Linnaeus