genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
frog
A frog is any member of a diverse and largely Carnivore, carnivorous group of short-bodied, tailless amphibians composing the order (biology), order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-f ...
s in the family
Hemiphractidae
The Hemiphractidae are a family (biology), family of frogs from South America, South and Central America. Previously, this group had been classified as a subfamily (Hemiphractinae) under family Hylidae. More recent research classifies these gen ...
. They are found in
Colombia
Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular regions in North America—near Nicaragua's Caribbean coast—as well as in the Pacific Ocean. The Colombian mainland is bordered by the Car ...
and
Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
. They are also known as backpack frogs, as the females have the habit of carrying their
egg clutch
__NOTOC__
A clutch of eggs is the group of eggs produced by birds, amphibians, or reptiles, often at a single time, particularly those laid in a nest.
In birds, destruction of a clutch by predators (or removal by humans, for example the Califor ...
on their backs until the young hatch; this behavior also occurs in the related hemiphractid genera ''
Hemiphractus
''Hemiphractus'' is a genus of frogs, the horned treefrogs, in the family Hemiphractidae. These overall brownish frogs have a pointed snout and a casque on the head. They are nocturnal, relatively rare and native to humid lowland and highland for ...
'' and ''
Stefania
''Stefania'' is a genus of frogs in the family Hemiphractidae. They are native to the highlands of the Guiana Shield in southern Venezuela, Guyana, and adjacent far northern Brazil. Most are restricted to the tepui highlands, but '' S. evansi'' ...
''.
Description
''Cryptobatrachus'' have a treefrog-like habitus. Males measure and females in
snout–vent length
Snout–vent length (SVL) is a morphometric measurement taken in herpetology from the tip of the snout to the most posterior opening of the cloacal slit (vent)."direct line distance from tip of snout to posterior margin of vent" It is the most c ...
. Fingers have no webbing whereas the toes are webbed. The finger and toe tips bear disks that are larger on the former. Also adhesive pads are present on the penultimate subarticular tubercles on the fingers. Males lack
vocal sac
The vocal sac is the flexible membrane of skin possessed by most male frogs and toads. The purpose of the vocal sac is usually as an amplification of their mating or advertisement call. The presence or development of the vocal sac is one way o ...
and these frogs appear not to vocalize.
Species
There are six species in the genus ''Cryptobatrachus'':
* ''
Cryptobatrachus boulengeri
''Cryptobatrachus boulengeri'', also known as Boulenger's backpack frog, is a species of frogs in the family Hemiphractidae. It is endemic to Sierra Nevada de Santa Marta in northern Colombia. The specific name honours George Albert Boulenger, ...
'' Ruthven, 1916
* ''
Cryptobatrachus conditus
''Cryptobatrachus'' is a genus of frogs in the family Hemiphractidae. They are found in Colombia and Venezuela. They are also known as backpack frogs, as the females have the habit of carrying their egg clutch on their backs until the young hatch ...
Cryptobatrachus remotus
''Cryptobatrachus'' is a genus of frogs in the family Hemiphractidae. They are found in Colombia and Venezuela. They are also known as backpack frogs, as the females have the habit of carrying their egg clutch on their backs until the young hatch ...
'' Infante-Rivero, Rojas-Runjaic, and Barrio-Amorós, 2009
* '' Cryptobatrachus ruthveni'' Lynch, 2008
The
AmphibiaWeb
AmphibiaWeb is an American non-profit website that provides information about amphibians. It is run by a group of universities working with the California Academy of Sciences: San Francisco State University, the University of California at Berkel ...
lists the six species above, but also ''Cryptobatrachus nicefori'', which the
Amphibian Species of the World
''Amphibian Species of the World 6.1: An Online Reference'' (ASW) is a herpetology database. It lists the names of frogs, salamanders and other amphibians, which scientists first described each species and what year, and the animal's known range. ...
''Cryptobatrachus'' occur on humid forested slopes in the mountain ranges of northern Colombia and northeast Venezuela at elevations of
above sea level
Height above mean sea level is a measure of the vertical distance (height, elevation or altitude) of a location in reference to a historic mean sea level taken as a vertical datum. In geodesy, it is formalized as ''orthometric heights''.
The comb ...
. They are
nocturnal
Nocturnality is an animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnal meaning the opposite.
Nocturnal creatures generally have highly developed sens ...
and perch on low bushes or cling to rocks and cliffs in spray zones of waterfalls.