''Crocodylus palaeindicus'' is an extinct

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of crocodile from southern

Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...

. ''C. palaeindicus'' lived from the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

to the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Mugger crocodile The mugger crocodile (''Crocodylus palustris'') is a medium-sized broad- snouted crocodile, also known as mugger and marsh crocodile. It is native to freshwater habitats from southern Iran to the Indian subcontinent, where it inhabits marshes ...

History

''C. palaeindicus'' was first named by Scottish paleontologist

Hugh Falconer Hugh Falconer MD FRS (29 February 1808 – 31 January 1865) was a Scottish geologist, botanist, palaeontologist, and paleoanthropologist. He studied the flora, fauna, and geology of India, Assam,Burma,and most of the Mediterranean islands a ...

in 1859. Falconer found fossils of the species in the

Siwalik Hills The Sivalik Hills, also known as the Shivalik Hills and Churia Hills, are a mountain range of the outer Himalayas that stretches over about from the Indus River eastwards close to the Brahmaputra River, spanning the northern parts of the Indian ...

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

along with the remains of many other animals like

turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked t ...

ostrich Ostriches are large flightless birds of the genus ''Struthio'' in the order Struthioniformes, part of the infra-class Palaeognathae, a diverse group of flightless birds also known as ratites that includes the emus, rheas, and kiwis. There ...

es, camels,

saber-toothed cat Machairodontinae is an extinct subfamily of carnivoran mammals of the family Felidae (true cats). They were found in Asia, Africa, North America, South America, and Europe from the Miocene to the Pleistocene, living from about 16 million ...

mastodon A mastodon ( 'breast' + 'tooth') is any proboscidean belonging to the extinct genus ''Mammut'' (family Mammutidae). Mastodons inhabited North and Central America during the late Miocene or late Pliocene up to their extinction at the end of th ...

Richard Lydekker Richard Lydekker (; 25 July 1849 – 16 April 1915) was an English naturalist, geologist and writer of numerous books on natural history. Biography Richard Lydekker was born at Tavistock Square in London. His father was Gerard Wolfe Lydekker, ...

later named another crocodile from the Siwalik Hills which he called ''C. sivalensis''. Although the two crocodiles are very similar, ''C. sivalensis'' was distinguished from ''C. palaeindicus'' because the margin of its skull was less convex. ''C. sivalensis'' has recently been synonymized with ''C. palaeindicus'', as the slight differences in shape are thought to be from natural variation or from fossilization. In later years, fossils were also found from

Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's fifth-most populous country, with a population of almost 243 million people, and has the world's second-lar ...

and

Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...

Classification

Historically, ''C. palaeindicus'' was considered a direct ancestor of the Mugger crocodile ''C. palustris''. The two species are similar in appearance, and some fossils of ''C. palaeindicus'' were at first mistaken for ''C. palustris''. Most modern

phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...

analyses of crocodiles place ''C. palaeindicus'' in a basal position among members of the genus ''Crocodylus''. Below is a

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

modified from Brochu ''et al.'' (2010) showing the relation of ''C. palaeindicus'' with other crocodiles: A 2018

tip dating Tip dating is a technique used in molecular dating that allows the inference of time-calibrated phylogenetic trees. Its defining feature is that it uses the ages of the samples to provide time information for the analysis, in contrast with traditio ...

study by Lee & Yates simultaneously using morphological, molecular ( DNA sequencing), and

stratigraphic Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithostra ...

(

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

age) data established the inter-relationships within Crocodylidae. In 2021, Hekkala ''et al.'' were able to use

paleogenomics Paleogenomics is a field of science based on the reconstruction and analysis of genomic information in extinct species. Improved methods for the extraction of ancient DNA (aDNA) from museum artifacts, ice cores, archeological or paleontological site ...

, extracting DNA from the extinct ''

Voay ''Voay'' is an extinct genus of crocodile from Madagascar that lived during the Late Pleistocene to Holocene, containing only one species, ''V. robustus''. Numerous subfossils have been found, including complete skulls, noted for their distinctiv ...

'', to better establish the relationships within Crocodylidae, including the subfamilies Crocodylinae and Osteolaeminae. The below

shows the results of the latest study:

References

{{Taxonbar, from=Q5187414 Crocodylidae Miocene crocodylomorphs Miocene reptiles of Asia Pliocene reptiles of Asia Pliocene crocodylomorphs