Creeping Soft Grass
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''Holcus mollis'', known as creeping soft grass or creeping velvet grass, is a species of
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns a ...
, native to
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located entirel ...
and western
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
.


Description

''Holcus mollis'' is a
rhizomatous In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ho ...
perennial grass found in woods and hedgerows, growing to tall. It has rhizomes that occur around deep in soil or sometimes deeper. Rhizome growth occurs in the period May to November but is fastest from mid-June to mid-July. The rhizomes have many dormant buds that do not develop unless the rhizomes are disturbed and then fresh aerial shoots may arise from the broken fragments. It flowers from June to July. The main distinguishing characteristics from '' H. lanatus'' are the presence of rhizomes, and the bearded nodes or 'hairy knees' on the culm.


Habitat

''Holcus mollis'' is favoured by conditions in woodland clearings and at the early stages of
coppicing Coppicing is a traditional method of woodland management which exploits the capacity of many species of trees to put out new shoots from their stump or roots if cut down. In a coppiced wood, which is called a copse, young tree stems are repeate ...
. Growth and flowering are restricted as the tree canopy develops. It is often a relict of former woodland vegetation, surviving in open grassland and grassy heaths after woodland clearance despite being a shade lover. It is found mostly on moist, freely-drained acid soils, normally light to medium texture and high in organic matter; it is absent from areas of calcareous or base rich soil, and often grows with
bracken Bracken (''Pteridium'') is a genus of large, coarse ferns in the family Dennstaedtiaceae. Ferns (Pteridophyta) are vascular plants that have alternating generations, large plants that produce spores and small plants that produce sex cells (eggs ...
.


Insect foodplant

The caterpillars of some Lepidoptera use it as a food plant, e.g. the
Essex skipper __NOTOC__ ''Thymelicus lineola'', known in Europe as the Essex skipper and in North America as the European skipper, is a species of butterfly in the family Hesperiidae. With a wingspan of 2.5 to 2.9 cm, it is very similar in appearance to ...
(''Thymelicus lineola'').


Status as a weed

In a survey of weeds in conventional cereals in central southern
England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe b ...
in 1982, it was found in 1% of
winter Winter is the coldest season of the year in polar and temperate climates. It occurs after autumn and before spring. The tilt of Earth's axis causes seasons; winter occurs when a hemisphere is oriented away from the Sun. Different cultur ...
barley Barley (''Hordeum vulgare''), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains, particularly in Eurasia as early as 10,000 years ago. Globally 70% of barley p ...
but not at all in
winter wheat Winter wheat (usually '' Triticum aestivum'') are strains of wheat that are planted in the autumn to germinate and develop into young plants that remain in the vegetative phase during the winter and resume growth in early spring. Classificatio ...
or
spring Spring(s) may refer to: Common uses * Spring (season) Spring, also known as springtime, is one of the four temperate seasons, succeeding winter and preceding summer. There are various technical definitions of spring, but local usage of ...
barley. Each small piece of rhizome is capable of developing into a new plant. Research shows that within of the surface, of rhizome infested soil may contain up to of rhizome, the weight of roots and rhizomes being estimated at 7.5 tons per acre. Hubbard (1976), p. 263.


Varieties and hybridisation

A pentaploid variant of ''H. mollis'' is common in Britain; it is sterile but spreads vegetatively. ''H. mollis'' var. ''variegatus'' has striped green and white leaves; it is sometimes cultivated. A male sterile hybrid with ''
Holcus lanatus ''Holcus lanatus'' is a perennial grass. The specific epithet ' is Latin for 'woolly' which describes the plant's hairy texture. Common names include Yorkshire fog, tufted grass, and meadow soft grass. In North America, where it is an invasive ...
'' exists with 2''n'' = 21 chromosomes. Hybrids tend to resemble ''H. lanatus'' in their morphology.Weed Information
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References


Bibliography

* {{Taxonbar, from=Q158965 Pooideae Flora of Asia Flora of Europe Plants described in 1759 Taxa named by Carl Linnaeus