''Coturnix'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

of five extant

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

and five to eight known extinct species of Old World

quail Quail is a collective name for several genera of mid-sized birds generally placed in the order Galliformes. The collective noun for a group of quail is a flock, covey, or bevy. Old World quail are placed in the family Phasianidae, and New Wor ...

. These species are distributed throughout

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

Eurasia Eurasia (, ) is the largest continental area on Earth, comprising all of Europe and Asia. Primarily in the Northern and Eastern Hemispheres, it spans from the British Isles and the Iberian Peninsula in the west to the Japanese archipelago a ...

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

, and formerly

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

. An extinct

radiation In physics, radiation is the emission or transmission of energy in the form of waves or particles through space or through a material medium. This includes: * ''electromagnetic radiation'', such as radio waves, microwaves, infrared, visi ...

flightless Flightless birds are birds that through evolution lost the ability to fly. There are over 60 extant species, including the well known ratites (ostriches, emu, cassowaries, rheas, and kiwi) and penguins. The smallest flightless bird is the ...

, insular species is known through fossil remains from

Macaronesia Macaronesia (Portuguese: ''Macaronésia,'' Spanish: ''Macaronesia'') is a collection of four volcanic archipelagos in the North Atlantic, off the coasts of Africa and Europe. Each archipelago is made up of a number of List of islands in the At ...

, which were likely wiped out by human arrival. The quails are related to the

African spurfowl ''Pternistis'' is a genus of galliform birds formerly classified in the spurfowl group of the partridge subfamily of the pheasant family. They are described as "partridge-francolins" in literature establishing their phylogenetic placement ou ...

jungle bush quail The jungle bush quail (''Perdicula asiatica'') is a species of quail in the family Phasianidae. It is native to the Indian subcontinent, where it is found in peninsular India and Sri Lanka. It has also been reported from Nepal but has not been se ...

snowcock The snowcocks or snowfowl are a group of bird species in the genus ''Tetraogallus'' of the pheasant family, Phasianidae. They are ground-nesting birds that breed in the mountain ranges of southern Eurasia from the Caucasus to the Himalayas and we ...

s and rock partridges, which together with the species of ''Coturnix'', ''

Synoicus ''Synoicus'' is a genus of 4 species of Old World quail. The species in the genus are distributed throughout Sub-Saharan Africa, tropical Asia, and Australasia. Two of the four species in the genus were originally classified in ''Excalfactoria'' ...

'', and a few others make up a

clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...

called

Coturnicini Coturnicini is a tribe of birds in the subfamily Phasianinae. It contains the Old World quail, snowcocks, and African spurfowl, among others. Members of this tribe have a wide range throughout Africa, Eurasia, and Australasia. This tribe conta ...

, a

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English language, English is in the discipline of anthropology. This definition is contested, in p ...

within the subfamily

Pavoninae The Phasianinae ( Horsfield, 1821) are a subfamily of the pheasant family (Phasianidae) of landfowl, the order Galliformes. The subfamily includes true pheasants, tragopans, grouse, turkey and similar birds. Although this subfamily was consid ...

. Quail of ''Coturnix'' live in pairs or small social groups and form larger groups during migration. Not all species migrate, but most are capable of extremely rapid, upward flight to escape from danger. Unlike related genera, Old World quail do not perch in trees. They devote much of their time to scratching and foraging for seeds and invertebrates on the ground. Typical habitats are dense vegetation such as grasslands, bushes alongside rivers and cereal fields. They are predated upon heavily by the diurnal hawks.

Taxonomy

The genus ''Coturnix'' was introduced in 1764 by the French naturalist

François Alexandre Pierre de Garsault François Alexandre Pierre de Garsault was a French botanist, zoologist and painter. de Garsault was born on 16 April 1691 in Aix-en-Provence, France and died 3 August 1778 in Paris, France. de Garsault was a member of the French Academy of Sci ...

. The

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...

is the common quail (''Coturnix coturnix''). The genus name is the

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

for the common quail. The genus contains six species, of which one, the New Zealand quail (''Coturnix novaezelandiae''), is now extinct but was described from a living specimen. The

brown quail The brown quail (''Synoicus ypsilophorus''), also known as the swamp quail, silver quail and Tasmanian quail, is an Australasian true quail of the family Phasianidae. It is a small, ground-dwelling bird and is native to mainland Australia, Tasman ...

(''S. ypsilophora''),

king quail The king quail (''Synoicus chinensis''), also known as the blue-breasted quail, Asian blue quail, Chinese painted quail, or Chung-Chi, is a species of Old World quail in the family Phasianidae. This species is the smallest "true quail", rangin ...

(''S. chinensis'') and blue quail (''S. adansonii''), were formerly classified in this genus, but were later reclassified into ''

''.

Species

Fragmentary remains representing three other ''Coturnix'' species were also recovered from Macaronesia: ''Coturnix'' sp. A from

Bugio Island Bugio Island ( pt, Ilha do Bugio) — is one of the three islands of the Portuguese Desertas Islands archipelago, a small chain of islands in the Madeira Islands Archipelago of Macaronesia. It is located in the Atlantic Ocean off the western co ...

Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...

, ''Coturnix'' sp. B from Santa Maria in the

Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...

(likely representing another extinct island endemic species) and ''Coturnix'' sp. C from

Graciosa Graciosa Island () (literally "graceful" or "enchanting" in Portuguese) is referred to as the ''White Island'', the northernmost of the Central Group of islands in the Azores. The ovular Portuguese island has an area of , a length of and a width ...

in the Azores. Due to their fragmentary nature, it is uncertain whether these represented their own species or were synonymous with one of the already-described extinct ''Coturnix'' species or the extant common quail (''Coturnix coturnix''), which also has fossil remains known from Macaronesia and is still present there. A

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

species from the Late

Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...

- Late

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

of SW and Central Europe was described as ''Coturnix gallica''. Another, ''C. donnezani'', was widespread in

Early Pliocene Early may refer to: History * The beginning or oldest part of a defined historical period, as opposed to middle or late periods, e.g.: ** Early Christianity ** Early modern Europe Places in the United States * Early, Iowa * Early, Texas * Early ...

to Early Pleistocene Europe.Mlíkovský (2002)

Footnotes

References

* Mlíkovský, Jirí (2002a): Early Pleistocene birds of Stránská skála, Czech Republic: 2. Absolon's cave. ''Sylvia'' 38: 19-28 nglish with Czech abstractbr>PDF fulltext
* The genetic link between the Chinese bamboo partridge (Bambusicola thoracica) and the chicken and junglefowls of the genus Gallus.A Fumihito, T Miyake, M Takada, S Ohno, and N KondoYamashina Institute for Ornithology, Chiba Prefecture, Japan. * Phylogenetic analysis of gallinaceous birds inferred from mitochondrial NADH dehydrogenase subunit 5 gene sequences Wee Hui Kit Publisher: 2002. * A Molecular Phylogeny of the Pheasants and Partridges Suggests That These Lineages Are Not Monophyletic R. T. Kimball,* E. L. Braun,*,† P. W. Zwartjes,* T. M. Crowe,‡,§ and J. D. Ligon*

External links

{{Taxonbar, from=Q1869243 Bird genera Taxa named by François Alexandre Pierre de Garsault