Coptotermes Acinaciformis
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''Coptotermes acinaciformis'' is a species of subterranean
termite Termites are small insects that live in colonies and have distinct castes (eusocial) and feed on wood or other dead plant matter. Termites comprise the infraorder Isoptera, or alternatively the epifamily Termitoidae, within the order Blattode ...
in the
family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...
Rhinotermitidae Rhinotermitidae is a family of termites (Isoptera). They feed on wood and can cause extensive damage to buildings or other wooden structures. About 345 species are recognized, among these are severe pests such as '' Coptotermes formosanus'', ''Co ...
native to Australia. Termites are
social insects Eusociality (from Greek εὖ ''eu'' "good" and social), the highest level of organization of sociality, is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping genera ...
and build a communal nest. In the case of ''C. acinaciformis'', this is either in the
root crown A root crown, also known as the root collar or root neck, is that part of a root system from which a stem arises. Since roots and stems have quite different vascular The blood vessels are the components of the circulatory system that transport ...
of a tree or underground. From this, a network of galleries extends through the nearby soil, enabling the workers to forage in the surrounding area without emerging on the surface of the ground. This termite can cause substantial damage to trees and the wooden parts of buildings.


The colony

''C. acinaciformis'' builds its nest at the base of a tree trunk or underground, and may form a mound. Subterranean passages extend from the nest up to and allow the workers to access other sources of wood on which to feed. Subsidiary nests are often built in such places as the cavities in the walls of buildings, but only in places where some moisture exists, perhaps from a leaking pipe. A large colony may contain more than a million termites. Where the nest is in a living tree, a central "pipe" of material is eaten out and replaced with
frass Frass refers loosely to the more or less solid excreta of insects, and to certain other related matter. Definition and etymology ''Frass'' is an informal term and accordingly it is variously used and variously defined. It is derived from the ...
. A number of other trees in the immediate vicinity can be similarly tunnelled, invasion taking place through the living bark. This is in contrast to the initial invasion of a tree by a newly mated pair of reproductives. In the latter case, the invasion is via pre-existing damage to the trunk. Tree stumps and fallen logs in the area are also affected, and mound nests have been shown to persist even when the host trees have been eaten out. The galleries from the colony may extend over an area of . Another favoured nesting site is where soil has been disturbed in a building site and pieces of timber have been accidentally buried in moist soil. This sometimes happens under patios or around new buildings.


The termites

Several species of termite are found in Australia and they are difficult to distinguish from one another. ''C. acinaciformis'' is most likely to be confused with '' C. frenchi'' or '' C. lacteus''. At a length of , the soldiers of ''C. acinaciformis'' are slightly larger than the other two species. When viewed from above, the soldiers of ''C. frenchi'' and ''C. lacteus'' have pear-shaped heads, while the heads of ''C. acinaciformis'' soldiers are more rectangular. Their lifecycle starts with the emergence of winged reproductives from a temporary hole in an established nest. A
nuptial flight Nuptial flight is an important phase in the reproduction of most ant, termite, and some bee species. It is also observed in some fly species, such as ''Rhamphomyia longicauda''. During the flight, virgin queens mate with males and then land to ...
takes place, and after they have returned to the ground, they shed their wings and search for a suitable place to found their colony. This is usually a piece of rotting wood buried in moist soil or a patch of decay at the base of a tree. The first eggs laid develop into workers and only when these are mature enough to take over the care of the young can the queen concentrate on egg-laying, the thing she will do for the rest of her life. Usually, three to five years are needed before the colony is big enough to cause damage to nearby buildings and structures.


Distribution and habitat

''C. acinaciformis'' is the most widely distributed termite in Australia and occurs throughout the country with the exception of Tasmania, parts of coastal Victoria, and a few other areas with high rainfall. It is found in both rural forested areas and in urban areas where trees are near buildings.


Research

Researchers have shown that when individual termites have been separated from a colony for some time and then returned to the colony, they are recognised and accepted by other members of the colony, whereas individuals from a different nest are rejected.


References

{{Taxonbar, from=Q16752093 Termites Insects described in 1898 Insects of Australia